Chimarra (Curgia) amigo, Holzenthal & Blahnik & Ríos-Touma, 2022

Holzenthal, Ralph W., Blahnik, Roger J. & Rios-Touma, Blanca, 2022, A new genus and new species of Ecuadorian Philopotamidae (Trichoptera), ZooKeys 1117, pp. 95-122 : 95

Chimarra (Curgia) amigo sp. nov.

Fig. 11

Type material.

Holotype. Male (pinned). Ecuador: Carchi: small stream 1, road from Chilmá Bajo to Moldanado, 0.90574°N, 78.21870°W, 1669 m a.s.l., 15.ii.2017, Ríos-Touma, Holzenthal, Amigo, Huisman, UMSP000378196 (UMSP). Paratypes. Ecuador: Carchi: Río Blanco between El Goaltal and Las Juntas, 0.80433°N, 78.16975°W, 1258 m a.s.l., Holzenthal and Huisman, 1 male (pinned) (UMSP); Pichincha: Quebrada Amagusa, 0.15561°N, 78.85356°W, 1254 m a.s.l., 21.vii.2015, Rázuri, Morabowen, Hernández, 2 males (pinned) (MECN).

Diagnosis.

Chimarra amigo has a general similarity to other species of the Chimarra (Curgia) fernandezi group of Flint (1998). The group is characterized by the form of tergum X, with the dorsomesal part forming a projecting lobe, either entire or slightly notched apically, and with projecting ventrolateral lobes on either side, and particularly by having an enlarged and distinctly sclerotized phallotremal sclerite complex, varying in shape and complexity among the different species of the group. Chimarra amigo differs from other described species of the group by having the preanal appendages flattened and almost completely fused (Fig. 11A, B ), much as in C. oztucoensis Flint & Reyes, 1991, which Flint (1998) placed in its own species group, largely because of having a tergum X that is deeply divided mesally. Chimarra amigo further differs from other species of the fernadezi group in the form of its inferior appendages, which are nearly subquadrate in lateral view, with the apicomesal projection very short and acute, not visible in lateral view (Fig. 11A ). Also distinctive for this species is its elongate tubular endotheca, which is very sharply bent or elbowed (Fig. 11E ).

Description.

Adult. Forewing length male 5.8-6.5 mm (n = 4); female 6.8 mm (n = 1). Head setae brownish black, slightly darker than wings. Wings brownish black, chord hyaline, distinct. Appendages dark brown. Pretarsal claws of male foreleg unmodified.

Male. Tergum VIII longer than sternum; dorsomesal margin projecting, entire; sternum VIII short ventrally, widening dorsally. Segment IX short dorsally, long ventrally, subtriangular in lateral view; anterolateral margin slightly concave, ventral margin produced; posterolateral margin angularly produced at level of inferior appendage; ventral process long, narrow, projecting, subacute apically; anteroventral margin concave mesally; dorsomesal margin broadly concave. Preanal appendage completely fused to tergum X represented by elevated patch of setae. Tergum X moderately long; divided into dorsal and ventrolateral lobes, dorsal lobe strongly rounded apicodorsally, apex laterally compressed, crest-like, ventrolateral lobes ~ 1/2 length of dorsal lobe, rounded apically; in dorsal view with apex of dorsal lobe shallowly cleft, ventrolateral lobes very narrow; apicomesally with numerous sensilla. Inferior appendage moderately long in lateral view, subquadrate; apex in ventral view abruptly narrowed, strongly inturned, forming short subapicomesal tooth-like projection. Endotheca elongate, tubular, with dorsomesal membranous projection when everted and inflated; endotheca strongly bent ventrad at dorsomesal projection; phallotremal sclerite forming large, complex, lightly sclerotized structure with single ventral and paired lateral blade-like sclerites, but much shorter than in other Chimarra fernandezi group species.

Female. Unknown.

Etymology.

This species is named in honor of Xavier Amigo, one of the collectors of the new species. He has provided essential support as a member of our field expeditions in Ecuador and is the beloved husband of Blanca Ríos-Touma.

Figures

Figure 11. Chimarra (Curgia) amigo sp. nov. Male genitalia A segments VIII-X, lateral B segments VIII-X, dorsal C inferior appendage, ventral D inferior appendage, dorsal E phallus, lateral.