Ceroderma asperata Raffray, 1890

Yin, Zi-Wei, Nomura, Shûhei & Li, Li-Zhen, 2012, Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae), Zootaxa 3355, pp. 62-68 : 63-65

Ceroderma asperata Raffray, 1890

( Figs. 1 , 3–11 , 19–21 )

Ceroderma asperata Raffray, 1890b: 199 . Type locality: Penang Island, Malaysian Peninsula. Type material examined (MNHN). Holotype ( Figs. 19–20 ): MALAYSIA: 3, labeled ‘Penang // MUSÉUM PARIS / 1917 / COLL. A. RAFFRAY // TYPE // C. asperata / A. Raffray det. // Ceroderma / asperata Raff / det. Löbl, 1970’.

Other material examined ( NSMT). 2 3, 1 Ƥ, labeled ‘Ulu Gombak, Selangor / (termite nest) / [MALAY- SIA] / 27.vii.2003, M. Maruyama // Host: Coptotermes curvi - / gnathus Holmgren, 1913 / M. Maruyama det., 2003 // FAUNA MALAYSIA / Staphylinidae , Pselaphinae / Ceroderma asperata Raffray / S. Nomura det., 2003’.

Diagnosis. Head triangular; vertex largely concave; postocular margins roundly expanded laterally; triangular pronotum broadly incised at basal half; elytra evenly constricted at basal half, with three basal foveae; tergite IV slightly longer than V.

Redescription. Male. BL 2.66. Body ( Figs. 1 , 19 ) reddish brown, maxillary palpi and tarsi lighter; head, pronotum and elytra covered with dense setae, microtuberculate where setae arise. Head ( Figs. 3 , 19 ) much wider than long, HL 0.48, HW 0.81; vertex with tufted setae at anterior and posterior borders of large transverse median concavity, vertexal foveae and sulcus overlapped by concavity; postocular margins largely angularly expanded laterally, with dense long lateral setae; eyes well-developed, each composed of about 60 facets; antennomeres II–VIII ( Fig. 4 ) of similar width, clubs formed by enlarged antennomeres IX–XI. Pronotum much wider than long, PL 0.55, PW 0.81; with shallow longitudinal median and lateral sulci; short discal carinae at basal half; lateral spines slightly larger than antebasal spines. Elytra wider than long, EL 0.82, EW 1.03; evenly constricted at basal half. Metaventrite concave along midline. Legs with femora thickened; protibiae ( Fig. 5 ) angularly expanded mesally at apical half; mesotibiae ( Fig. 5 ) with tiny membranous apical projection; metatrochanters ( Fig. 7 ) with large elongate ventral projection. Abdomen wider than long, AL 0.81, AW 1.02; tergite IV with thick triangular lateral ridges; tergites V–VI with ridges much thinner; tergite VII ( Fig. 8 ) with small lateral tubercles sharp. Sternite IV twice length of V at midline, with short median carina; sternites V–VII with short triangular basolateral ridges. Aedeagus ( Figs. 9–11 ) well-sclerotized, length 0.37; broad basal bulb with large foramen, median lobe expanded apically.

Female. Similar to male in general. Measurements: BL 2.62, HL 0.47, HW 0.74, PL 0.55, PW 0.78, EL 0.78, EW 1.04, AL 0.82, AW 0.98. Vertex lacking setose concavity, widely separated foveae connected by shallow triangular sulcus in large impressed area; postocular margins less expanded than in male, lacking long lateral setae. Each eye composed of about 55 facets. Protibiae lacking mesal expansion at apical half; mesotibiae lacking apical tubercle.

Distribution. Malaysia: Penang, Selangor.

Host termite. Coptotermes curvignathus Holmgren (det. M. Maruyama).

Figures

FIGURE 1 – 2. Dorsal habitus of Ceroderma asperata (1) and Cerochusa cilioceps (2). Scales: 0.5 mm.

FIGURE 3 – 11. Ceroderma asperata. 3. Head and pronotum. 4. Antenna. 5. Protibia. 6. Apex of mesotibia. 7. Metatrochanter. 8. Right half of tergite VII. 9. Aedeagus, in dorsal view. 10. Same, in lateral view. 11. Same, in ventral view. Scales: a, b = 0.5 mm; c = 0.3 mm; d = 0.05 mm; e – i = 0.2 mm.

FIGURE 19 – 21. Ceroderma asperata. 19. Holotype. 20. Type label. 21. Raffray’s original habitus illustration. Scale: 1.0 mm.