Cephaloleia luctuosa Guerin-Meneville , 1844

Staines, Charles L. & Garcia-Robledo, Carlos, 2014, The genus Cephaloleia Chevrolat, 1836 (Coleoptera, Chrysomelidae, Cassidinae), ZooKeys 436, pp. 1-355 : 177-178

Taxon classification Animalia Coleoptera Chrysomelidae

Cephaloleia luctuosa Guerin-Meneville, 1844 Fig. 179

Cephaloleia luctuosa Guérin–Méneville 1844: 282. Baly 1885: 55; Blackwelder 1946: 719 (catalog); Papp 1953: 18 (catalog); Uhmann 1957a: 21 (catalog), 1966: 269 (noted); Staines 1996: 42 (Central America species), 1999: 241 (mimicry); McKenna and Farrell 2005: 119 (phylogeny), 2006: 10949 (phylogeny).

Cephalolia luctuosa Guérin-Méneville. Baly 1858: 55 (redescription); Gemminger and Harold 1876: 3601 (catalog); Donckier 1899: 550 (catalog); Weise 1910: 86 (noted), 1911a: 8 (catalog), 1911b: 10 (catalog), 1913: 101 (noted); Uhmann 1932b: 261 (museum list), 1936a: 111 (noted), 1936b: 484 (key); Bryant 1942: 205 (faunal list).

Description.

Elongate; subparallel; subdepressed; head (except reddish-yellow frons) and scutellum black; antennae entirely black or antennomeres 8-11 yellow; pronotum reddish-yellow with black quadrangular macula medially of anterior margin extending towards base; elytra black with yellow vitta from puncture rows 4-9, not reaching apex, lateral margin usually dark, sometimes pale; venter pro-, meso-, and metasterna yellow medially, black laterally, abdominal sternite 1 yellow medially, black laterally; 2-4 black except pale marginal vitta; 5 entirely black; leg femur yellow, tibia darker. Head: vertex punctate, not rugose; Y-shaped medial sulcus present; frons punctate, not projecting; not depressed between eyes. Antenna: reaches to humerus; slender; antennomere 1 elongate, with tuft of setae at apex; 2 ⅓ length of 1, with sharp inner angle; 3 2 × length of 2, triangular; 4-10 elongate, decreasing in length; 11 2 × length of 10, pointed at apex; 2 punctate with scattered setae; 1, 3-11 setose. Pronotum: subquadrate; lateral margin straight then rounding to anterior angle, margined; anterior angle angulate, slightly produced; posterior angle acute; anterior margin weakly emarginate behind head; disc subconvex; surface punctate basally; basal impression absent; pronotal length 1.0-1.3 mm; pronotal width 1.4-1.6 mm. Scutellum: broadly triangular; impunctate. Elytron: lateral margin straight, smooth, narrowly margined; apex rounded; sutural angle without tooth; humerus rounded, slightly produced; slightly constricted behind humerus; shallowly punctate-striate; declivity beginning just behind humerus at puncture row 7 not edged with faint carina; elytral length 4.6-5.3 mm; elytral width 1.9-2.1 mm. Venter: pro-, meso-, and metasterna impunctate medially, punctate laterally; abdominal sterna punctate, each puncture with pale seta; suture between sterna 1 and 2 complete; last sternite with apical margin sinuate medially in male, truncate in female. Leg: slender, sparsely punctate; tibia with tuft of setae at apex. Total length: 6.0-7.0 mm.

Diagnosis.

This species is similar to Cephaloleia bella , Cephaloleia championi , and Cephaloleia vicina . It can be distinguished by the vertex of the head not being depressed between the eyes, by the suture between abdominal sterna 1 and 2 being complete, and by antennomeres 2 and 3 being triangular.

Distribution.

Colombia, Panama.

Type material.

Type: Colombia, Santa Fe de Bogata (depository unknown, not seen).

Specimens examined.

Colombia: Meta- road to Bogota- Villavicencio, 1500 m, 29 June 1965 (AMNH). PANAMA: Bocas de Toro- Almirante, S 4 38 (USNM), 22 March 1959 (FMNH); 6 km N Punta Peña, 27 May 1993, 28 May 1993 (CDFA, AJGC); 2.3 mi. N Continental Divide, Reserva Fortuna, 26 May 1993 (AJGC), 28 May 1993 (CDFA). Canal Zone- 22 August 1970 (USNM). Chiriquí- 11.2 km E Chiriquí, 30 May 1993 (CDFA, AJGC); Repr. La Fortuna, 17-21 September 1976, 3200' (EGRC); Reserva Fortuna, Continental Divide Trail, 25 May 1993, 26 May 1993, 29 May 1993 (CDFA, EGRC). Coclé- Cerro Gaital, 10-12 June 1985 (EGRC). Colón- Achiote Road, 10 km SW Gatun, 12 June 1976 (USNM, EGRC); El Valle, alt 2400-2600 ft, 21 February 1959, 22 February 1959, 23 February 1959 (FMNH); vic. Fort Sherman, 15-16 February 1999 (USNM); Gamboa, 22 June 1976 (EGRC); 5 mi NW Gamboa, 27 April 1974 (EGRC); Pipeline road, 23 May 1993 (AJGC, CDFA); Pipeline rd. nr. Gamboa, 1 July 1976 (EGRC); Paraiso, 08 January 1911, 17 January 1911, 26 January 1911, 5 February 1911, 5 March 1911, 26 March 1911 (USNM); Madden Rd., 27 February 1959 (FSCA); Porto Bello, 23 February, 19 February 1911, 26 Febru ary, 27 February 1911, 2 March 1911 (USNM). Panamá- Alajuela, 5 April 1911, 18 April 1911 (USNM); Arraiján (USNM); Cerro Campana, 850 m, 17 February 1959 (FMNH), 11-15 May 1985 (EGRC), 17 May 1993 (CDFA); nr. Chepo, 3 April 1971 (EGRC); Corazal, 12 January 1911 (USNM); Cristobal, 9 February 1959 (FMNH); Fort Kobbe, 6 June 1976, 8 June 1976, 15 June 1976, 20 June 1976 (EGRC); Madden Forest, Mi 2.5, 2 August 1970 (CMNC); Ft. Howard, 24 October 1973 (FSCA); Madden Forest, 9 January 1971, 6 March 1971, 27 March 1971, 27 May 1971, 2 November 1973, 8 August 1977 (EGRC); 9 km N El Llano, 18 May 1993 (CDFA); Fort Kobbe, 22 May 1993 (AJGC, CDFA); Old Gamboa Road, 4 June 1993 (AJGC, CDFA); Las Cumbres, 8 January 1959 (FMNH); La Pita Signal Station rd., 8 June 1976, 16 June 1980 (EGRC); Panama Road leading to La Pita signal station, 2 May 1971 (EGRC); Reserva Sobrina, Powerline Road, 29 October 1972 (FSCA); Summit, September 1946 (USNM). San Blas- Salud, 30 December 1972 (EGRC). Total: 396.