Boreantrops oaxacensis, Kits, Joel H. & Marshall, Stephen A., 2015

Kits, Joel H. & Marshall, Stephen A., 2015, A revision of Boreantrops Kits & Marshall (Diptera: Sphaeroceridae: Archiborborinae), Zootaxa 3915 (3), pp. 301-355 : 328-329

Boreantrops oaxacensis sp. nov.

( Figs. 57, 58 , 74, 116)

Diagnosis. Separable from other members of the mexicanus subgroup by the following combination of characters: Prementum entirely orange. Leg joints broadly orange. Mid tibia mostly orange. Male sternite 5 without a notch in posterior margin.

Description. Head orange, occiput black medially, maxillary palp yellow. Mostly covered with microtomentum, frons with microtomentum medially reaching from postocellar bristles anteriorly through ocellar triangle to anterior margin of frons, microtomentum on orbital plates reaching bases of interfrontal setae, face with patch of microtomentum below lunule and antenna. Ocellar bristles just anterior to median ocellus. Subvibrissal bristle and anterior genal bristles about 0.3X length of vibrissa.

Thorax black, mostly covered with microtomentum. Proepisternum shiny, anepisternum with microtomentum along dorsal third and posterior third, katepisternum with a shiny spot behind fore coxa, meron shiny, metapleuron with shiny spot in posteroventral corner. Mesoscutum sculptured. Halter white, brownish below knob.

Coxae, basal three-fifths of fore femur, basal two-thirds of mid and hind femur, and distal half of fore and hind tibia dark reddish brown to black, extreme bases and distal portions of femora, mid tibia and basal half of fore and hind tibia orange, mid tibia sometimes with dorsal brown spot at 4/5, fore tarsus with 2 basal tarsomeres ivory white, mid and hind tarsi with 2 basal tarsomeres yellow, all tarsi with 3 distal tarsomeres brown. Mid tibia with 1 anteroventral, 6 subapical bristles. Hind tibia with 1 ventroapical bristle.

Wing brown, veins dark brown, crossveins r-m and dm-cu white, vein R4+5 with 2 spots distal to crossvein r-m.

Abdominal tergites 1–4 mostly heavily sclerotized, black, covered with microtomentum, syntergite 1+2 with a semicircular weakly sclerotized patch anteriorly. Tergite 5 weakly sclerotized. Sternites weakly sclerotized.

Male postabdomen: Sternite 5 longer than wide, lateral margins straight, widest posteriorly, posterior margin with a shallow notch and narrowly interrupted row of setae, posteromedial patch weakly sclerotized, anterior apodeme broad, about as long as external portion, without keel ( Fig. 58 ). Surstylus paddle-shaped. Hypandrial arms with ventral tab-like structures. Pregonite small, discrete. Postgonite narrowed above lobes, lobes widely separated, anterior lobe broad, rounded, posterior lobe pointed. Basiphallus with long epiphallus, long, transparent preepiphallus. Distiphallus with strongly curved spinose dorsal tube, distal portion short, pointed apically ( Fig. 57 ).

Female postabdomen: Tergites 6–7 with anterior and lateral margins heavily sclerotized, center and posterior margin not sclerotized. Tergite 8 covered with microtomentum, except anterior margin. Epiproct without anterior arms, mostly covered with microtomentum except anterior corners, with 2 setae. Cerci broad. Sternites 6–7 with margins heavily sclerotized, center not sclerotized. Sclerites of sternite 8 broadest posteriorly, posterior half covered with microtomentum. Hypoproct lozenge-shaped, covered with microtomentum. Spermathecae about 2X longer than wide, annulated, apex invaginated, sclerotized duct about 2X as long as bulb (Fig. 74).

Type material. Holotype ♂: MEXICO: Oaxaca: 1.7 mi W Jct Mex. 175-Yuvila Rd., 2865 m, mesic oak, fish trap, 9–19 Aug 1973, A. Newton, debu01040237 ( DEBU). Paratypes: same data as holotype (2 ♂, 2 ♀, DEBU); as above but mesic oak forest (3 ♂, 4 ♀, DEBU); Suchixtepec, 8 km S, 10 Aug 1986, H. & A. Howden (2 ♂, 8 ♀, DEBU); Valle Nacional, 29.7 mi S, 2073 m, cloud forest, dung, 11 Aug 1973, A. Newton (1 ♀, DEBU).

Distribution. Only known from Oaxaca ( Fig. 116 ). Elevations over 2000 m.

Etymology. The species name refers to the occurrence of this species in Oaxaca.

Figures

FIGURES 55 – 60. Boreantrops male terminalia. B. masneri: 55) phallus, postgonite, and phallapodeme, lateral view, 56) sternite 5; B. oaxacensis: 57) phallus, postgonite, and phallapodeme, lateral view, 58) sternite 5; B. peruvianus: 59) phallus, postgonite, and phallapodeme, lateral view, 60) sternite 5.

FIGURES 115 – 120. Boreantrops distribution maps. 115) B. masneri (diamonds), B. calceatus (circle); 116) B. oaxacensis; 117) B. peruvianus; 118) B. subfoveolatus; 119) B. suchixtepecensis; 120) B. zacapa. Elevation is indicated by grey shading at 500 m intervals.