Basalys villumi, Brazidec & Vilhelmsen, 2022

Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, European Journal of Taxonomy 813, pp. 57-86 : 76-77

Basalys villumi sp. nov.

urn:lsid:zoobank.org:act:4204B17E-4659-432C-9266-1C723B434E35

Figs 1F , 5F–H , 6A , Table 1

Diagnosis

Body smooth; scape seven times as long as wide; pedicel conical; flagellomeres cylindrical and longer than wide; F1 longest; F2–F11 similar in length; F2 conspicuously emarginate ( Fig. 5D–E ); scutellum subquadrate without posterior scutellar pits ( Fig. 5F ); fore wing bordered with small hairs, r-rs short but distinct, M+Cu nebulous ( Fig. 1F ); hind wing narrow, two thirds of fore wing length, with long hairs along posterior margin; petiole three times as long as wide ( Figs 5D , 6A ).

Etymology

We dedicate this species to Villum Fonden, who generously contributed to funding for purchasing the amber pieces studied. The specific epithet is to be treated as a noun in the genitive case.

Type material

Holotype NHMD-608360 , a complete male; paratype NHMD-608369 , a complete male.

Locality and horizon

Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma.

Description

Male

BODY. BL = 1.82 mm. Body highly glabrous. Head globular, as long as high (HeL = 0.33 mm), with sparse short hairs; eye round; toruli situated at middle of eye height; antenna 14-segmented; scape seven times as long as wide; pedicel conical; flagellomeres cylindrical, elongate, distinctly longer than wide, with short hairs; F1 longest, F2–F11 subequal in length; F2 emarginate on anterior third; F12 as long as F1; tapering at apex (antennomeres length of holotype, in mm: Sc- 0.22 mm; P-0.09; F1-0.15; F2-0.09; F3-0.11; F4-0.10; F5-0.11; F6-0.10; F7-0.12; F8-0.10; F9-0.11; F10-0.12; F11-0.11; F12-0.14); mandibles weakly crossing at tip, not sickle or beak-shaped; occipital flange foveate, rather concave.

MESOSOMA. Shorter than metasoma (MsL = 0.63 mm); pronotum not elongate, anterior part setose, epomia absent; mesoscutum slightly convex, smooth, notauli absent; anterior scutellar pit present, round; scutellum subquadrate, without posterior scutellar pits. Fore wing extending beyond metasoma (FwL = 1.78 mm), bordered with small hairs; Sc+R distinctly separated from anterior margin; marginal vein thickened along wing margin, longer than wide; r-rs short; basal vein curved, perpendicular to Sc+R, almost reaching M+Cu; M+Cu nebulous. Hind wing narrow, two thirds of fore wing length; bordered with long setae along posterior margin. Legs slender, covered with scattered setae; femur clavate; tibia weakly broadened apically; tibial spur formula 1-2-2; tarsal claws simple.

METASOMA. Petiole cylindrical, narrow, three times as long as wide (PtL = 0.21 mm; PtW = 0.07 mm), longitudinally striated; gaster ellipsoidal (GL = 0.61 mm; GH = 0.30 mm), glabrous; T2 and S2 longest, covering two thirds of gaster, anterior margin T2 straight; segments 3 and 4 ring-like, segment 5 longer than 3 and 4 combined.

Female

Unknown.

Comments

Using Nixon’s (1980) and Masner & García’s (2002) keys, Basalys villumi sp. nov. keys out to Basalys Westwood, 1833 because of the following characters: antenna with 14 segments, notauli absent, scutellum with anterior scutellar pit, wings fully developed, Sc+R separated from fore margin of wing, fore wing with distinct basal vein, basal margin of large tergite straight. The description of the species is also consistent with the diagnosis of Hou & Xu (2016).

Basalys villumi sp. nov. differs from extant Basalys by having F1 longer than F2 ( Nixon 1980; Hou & Xu 2016).

Figures

Fig. 1. Wing venation of the species described here. A. Belyta knudhoejgaardi sp. nov. B. Cinetus breviscapus sp. nov.C. Cinetus elongatus sp. nov.D. Pantoclis globosasp. nov. E. Pantolyta augustinusii sp. nov. F. Basalys villumi sp. nov. G. Doliopria baltica sp. nov. H. Spilomicrus succinalis sp. nov. Scale bars: A–E = 1 mm; F, H = 0.5 mm; G = 0.25 mm.

Fig. 5. A–C. Pantolyta chemyrevae sp. nov., holotype (NHMD-608448). A. Habitus in dorsolateral view. B. Head and mesosoma in dorsal view. C. Petiole in lateral view. D–E. Pantolyta similis sp. nov., holotype (NHMD-608468). D. Habitus in lateral view. E. Mesosoma, petiole and anterior gaster in dorsal view. F–H. Basalys villumi sp. nov., holotype (NHMD-608360). F. Habitus in lateral view. G. Head in lateral view. H. Head and mesosoma in dorsal view. Abbreviations: asp = anterior scutellar pit; fe = flagellomere emargination; mpk = median propodeal keel. Scale bars: A, D, F = 1 mm; B, E, G–H = 0.5 mm; C = 0.25 mm.

Fig. 6. A. Basalys villumi sp. nov., paratype (NHMD-608369), habitus in lateral view. B–D. Doliopria baltica sp. nov., holotype (NHMD-608374). B. Habitus in lateral view. C. Head in lateral view. D. Mesosoma and metasoma in lateral view. E–H. Spilomicrus succinalis sp. nov. E–F. Holotype (NHMD-607131). E. Habitus in ventrolateral view. F. Head and mesosoma in dorsal view. G. Paratype (NHMD-608344), habitus in lateral view. H. Paratype (NHMD-608354), habitus in lateral view. Abbreviations: asp = anterior scutellar pit; te = tergite excision. Scale bars: A, E–H = 1 mm; B = 0.5 mm; C–D = 0.25 mm.