Atractodes altoandinus, Bordera, Santiago, Mazón, Marina & Sääksjärvi, Ilari E., 2016

Bordera, Santiago, Mazón, Marina & Sääksjärvi, Ilari E., 2016, The Neotropical species of Atractodes (Hymenoptera, Ichneumonidae, Cryptinae), I: the A. propodeator and A. altoandinus species-groups, Zootaxa 4137 (1), pp. 108-120 : 118-119

Atractodes altoandinus sp. nov.

( Figs 3 C, 4E, 8A–C)

Diagnosis. Atractodes altoandinus can be distinguished from all other Neotropical species of Atractodes by the combination of the following characters: propodeum more or less convex and uniformly sloped downwards posteriorly, strongly rugose-granulate, almost mat. Area basalis absent. Mesopleuron mostly smooth and shiny. Distal abscissa of Cu 1 present only as a shallow trace. Flagellum thicker distally than in central part ( Figs 4 E, 8A), width of penultimate flagellomere about 1.4 × the width of first one. First flagellomere very short, about 3.0 × as long as broad ( Figs 4 E, 8B). Tergite I very short, 2.7 × as long as broad, in lateral view, dorsal part strongly curved ( Fig. 8 C). Sternaulus reaching hind rim of mesopleuron, posteriorly granulate ( Fig. 3 C).

Description. Female: Body length 3.3 mm. Fore wing 2.7 mm long.

Head.- Transverse, about 0.6 × as wide as long, very slightly narrowed behind compound eyes, rounded (viewed from above); gena about 0.7 × as long as eye, smooth and shiny with sparse shallow setiferous punctures, setae relatively short, relatively sparse at ventral part. Frons and vertex finely granulate with dense shallow setiferous punctures, frons smooth and shiny, setae relatively short. Occipital carina rounded in middle part. Posterior ocellus separated from eye by 2.1 × its own maximum diameter; distance between posterior ocelli 1.4 × its own maximum diameter. Face finely and very densely punctate on a granulate background. Clypeus weakly convex, 3.3 × as wide as long, densely punctate on a smooth and shiny background, apex slightly upturned medially, setae conspicuously longer than in face. Lower tooth of mandible 0.9 × the length of upper tooth. Eye with relatively long and more or less dense setae. Malar space about 1.4 × basal mandibular width, only slightly granulate just behind the mandible base. Antenna with 17 flagellomeres; first flagellomere, seventh and penultimate 3.0, 1.9 and 1.4 × as long as wide, respectively; antenna slightly widened towards apex, width of penultimate about 1.4 × the width of first flagellomere ( Figs 4 E, 8A).

Mesosoma ( Figs 3 C, 8B). Pronotum finely and evenly granulate; epomia shallow and short, only reaching the anterior submarginal depression. Mesoscutum very fine and densely punctate on a smooth and shiny background; notauli deep, reaching 0.5 × the length of mesoscutum; prescutellar groove deep, smooth, with longitudinal striae; scutellum with dense fine and shallow setiferous punctures, not delimited by a lateral carina. Mesopleuron shiny, with some fine and sparse setiferous punctures; central part smooth and glabrous; sternaulus deep, reaching the hind rim of mesopleuron, posteriorly granulate. Metapleuron strongly punctate and granulate, sparsely setose; juxtacoxal carina absent. Propodeum more or less convex and uniformly sloped downwards posteriorly; area basalis absent; strongly rugose-granulate, almost mat, setae relatively long and dense; median area clearly defined, granulate, narrowed towards base, about 2.5 × as long as its maximum width; spiracle very small, rounded, separated from pleural carina 3.7 × its diameter. Legs long and slender; length of hind femur about 4.9 × its width; hind tarsal claws thin and right-angled curved, as long as arolium.

Wings. Densely setose. Fore wing with pterostigma about 2.5 × wider than high, vein Rs +2 r about 1.2 × as long as its height; areolet pentagonal, open, about 0.7 × higher than wide; vein 2 m-cu with a single bullae; first abscissa of Cu 1a 1.4 × Cu 1b. Hind wing with vein cu-a + abscissa of Cu 1 between M and cu-a, angulate, intercepted at its lower 0.4, straight; distal abscissa of Cu 1 not pigmented, only present as a shallow trace.

Metasoma ( Fig. 8 C). Tergite I smooth, with few short lateral setae, 2.7 × longer than posteriorly broad, in lateral view dorsal part very strongly curved; cross section of petiole on the mid part more or less circular, upper side slightly convex; tergite II smooth and shiny, about 1.9 × longer than basally broad; lateral crease separating tergite II from epipleuron, absent; other tergites smooth and shiny, with some sparse setae.

Colouration ( Fig. 8 A). Body black to dark brown; flagellum dark brown, lighter towards the apex; mandibles light brown. Legs dark brown. Metasoma from tergite II to the apex, brown-reddish; tergite I black.

Male: unknown.

Etymology. The name of the species refers to the high Andean region of Chile where the holotype was collected.

Type material. Holotype, 1 ♀: Chile, La Raya, 4300 m, 21.II.1950, 1 ♀, leg. P. Guzemon ( AEIC).

Figures

FIGURE 3. Morphology of female meospleuron: A— Atractodes species-group A; B— A. propodeator species-group; C— A. altoandinus species-group; D— Atractodes species-group B.

FIGURE 4. Morphology of antennae. A—D, Atractodes propodeator species-group; A— A. verapacensis, ♀; B— A. reijoi, ♀; C— A. propodeator, ♀; D— A. verapacensis, ♂; E— Atractodes altoandinus species-group (A. altoandinus, ♀).

FIGURE 8. Morphology of A. altoandinus (holotype), lateral view: A—Habitus; B—Head and mesosoma; C—Metasoma.