Anthobium cornatipenis, Shavrin, 2022

Shavrin, Alexey V., 2022, A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records, Zootaxa 5104 (3), pp. 301-346 : 326-327

Anthobium cornatipenis sp. n.

( Figs. 3 , 65–66 , 93 )

Type material examined: Holotype ♂ [dissected]: ‘ CHINA: W-Sichuan 1999 | Ganzi Tibet. Aut. Pref., Batang Co. | Shalui Shan, Fichtenwald 55 km | NE Batang, 30°17N, 99°31E | Rinde, Pilze, Nadelstreu, 4300m | 1. VII., leg. M. Schülke’ <printed>, ‘Sammlung | M. Schülke | Berlin’ <printed>, ‘HOLOTYPE | Anthobium | cornatipenis sp. n. | Shavrin A.V. des. 2022’ <red, printed> (cSch) .

Paratypes (3 specimens): 3 ♀♀: same data as the holotype (1 ♀: cSh; 2 ♀♀: cSch).All paratypes with additional red printed label: ‘ PARATYPE | Anthobium | cornatipenis sp. n. | Shavrin A.V. des. 2022’.

Description. Measurements (n=4): HL: 0.38–0.43; HW: 0.62–0.68; AL (holotype): 1.17; OL: 0.15–0.17; PL: 0.55–0.58; PW: 1.00–1.06; ESL: 1.45–1.48; EW: 1.35–1.47; AW: 1.15–1.21; MTbL (holotype): 0.70; MTrL (holotype): 0.37 (MTrL 1–4: 0.22; MTrL 5: 0.15); AedL: 0.72; TL: 3.40–3.60 (holotype: 3.50).

Habitus as in Fig. 3 . Head, maxillary palpomeres, antennomeres, pronotum and abdomen brown; lateral and basal portions of pronotum and elytra yellow-brown (apical portions of elytra paler); basal portions of antennomeres 1–3 and legs yellow. Head with distinct microscupture, denser and transverse on clypeus, transverse, sparser and sometimes finer in middle, and isodiametric on infraorbital ridges; neck with fine and transverse microreticulation; pronotum with transverse microsculpture, finer in middle and sometimes missing in mediobasal third; abdomen with very dense isodiametric meshes. Head with moderately dense punctation, finer in basal portion of vertex, denser and larger in middle and infraorbital ridges, with moderately wide mediobasal portions in front of ocelli without any sculpture; middle portion of neck with sparse and fine punctation; punctation of pronotum slightly sparser than that in middle of head, but distinctly coarser, deeper and sparser in middle and basal portions, with or without very sparse punctures in mediobasal third; scutellum without visible punctures; punctation of elytra about as that on pronotum, but markedly sparser, deeper and coarser in basal and apical portions, finer and sparser in middle, on each elytron forming very tangled and vague five to six longitudinal rows of punctures; abdominal tergites with very sparse and fine punctation.

Head 1.5–1.6 times as broad as long, with slightly elevated middle portion and infraorbital ridges, with deep anteocellar foveae, reaching level of apical third of eye, and obtuse postocular ridges. Apical segment of maxillary palpomere twice as long as preceding segment. Antennomere 2 distinctly narrower than basal antennomere, 3 slightly longer and narrower than 2, 4–5 shorter and broader than 3, 6–8 slightly broader than 5, 9–10 about as long as broad, slightly broader than 8, apical antennomere from middle gradually narrowed toward rounded apex.

Pronotum 1.8 times as broad as long, more or less evenly narrowed both anteriad and posteriad; anterior angles not protruded anteriad; median portion without median impressions, lateral portions slightly impressed.

Elytra about as long as broad or slightly longer, significantly more than twice as long as pronotum, reaching apical margin of abdominal tergite IV, with rounded apical margins.

Metatarsus 1.8 times as long as metatibia.

Male. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII sinuate. Aedeagus with narrow basal bulb, gradually narrowed toward wide, truncate apex; parameres significantly exceeding apex of median lobe, slightly widened in preapical portion and narrowed apically, with one short and two preapical setae; internal sac very long, spirally folded in basal portion ( Fig. 65 ). Lateral aspect of aedeagus as in Fig. 66 .

Female. Apical margins of abdominal tergite VIII and sternite VIII truncate or rounded.

Comparative notes. Regarding the shape of the narrow aedeagus with widely truncate apex and long parameres, it is similar to the Middle Asian A. tichomirovae sp. n., from which it can be distinguished by the darker and smaller body, shorter antennomeres 6–10, less protruding anterior angles of the pronotum, and narrower apical portions of the parameres.

Distribution. The new species is at present known only from the type locality in Shalui Shan, Sichuan, China ( Fig. 93 ).

Natural history. The specimens were found in spruce forest under bark and sifted from mushrooms and needle litter at an elevation of 4300 m a.s.l.

Etymology. The specific epithet is a combination of Latin adjective cornatus (horn-like, horn-shaped) and the Latin noun penis (penis). It refers to the shape of apical portion of the aedeagus.

Figures

FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond).

FIGURES 1–5. Habitus of species of Anthobium: 1—A. algidum (Austria, Innsbruck), 2—A. tectum (Russia, Buryatia), 3—A. cornatipenis (paratype), 4—A. himalayense (holotype), 5—A. multipunctatum (holotype). Scale bar: 1.0 mm.

FIGURES 63–68. Aedeagus of Anthobium tectum (Figs. 63–64; Finland, Kuhmo), A. cornatipenis (Figs. 65–66; holotype) and A. frigidum (Figs. 67–68; syntype): 63, 65, 67—parameral view, 64, 66, 68—lateral view. Scale bar: 0.1 mm.