Andrena ( Micrandrena ) hebraica Pisanty & Wood, 2025

Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie & Wood, Thomas J., 2025, Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus, Arthropod Systematics & Phylogeny 83, pp. 573-655 : 573-655

publication ID

https://doi.org/10.3897/asp.83.e166986

DOI

https://doi.org/10.5281/zenodo.17516180

persistent identifier

https://treatment.plazi.org/id/F37F4D20-4ADA-511C-B97F-D2524DF4650E

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Andrena ( Micrandrena ) hebraica Pisanty & Wood
status

sp. nov.

3.1.2.12. Andrena ( Micrandrena) hebraica Pisanty & Wood sp. nov.

Figures 9, 17E, J View Figure 17 View Figure 9 , 21 J View Figure 21

Etymology.

Nominative feminine singular form of the Latin adjective “ hebraicus ” (= Hebrew).

Diagnosis.

Within the Andrena minutula species group, A. hebraica belongs to the species around A. spreta Pérez , which are characterized in the female by a clypeus which is non-rugose, weakly domed and partly to fully shagreened, a scutum which is shagreened to partly smooth, with fine, relatively sparse punctures, and terga which are mostly shagreened and impunctate. In the Levant and Cyprus, these include A. aphroditae sp. nov., A. minutuloides , A. spreta , A. tiaretta and A. tkalcui . Andrena hebraica differs from all the above species by the sculpturing of tergal disc 3 and to a lesser extent 2, which are apically shiny, instead of completely shagreened and matt. This difference is difficult to illustrate by photographs, but is clearly visible to the eye under magnification. In addition, the clypeus is more clearly domed and more strongly and extensively shagreened compared to A. spreta and A. tiaretta (Figs 8 B View Figure 8 , 17 E View Figure 17 ), the clypeus is more finely punctured and the scutum more densely punctured compared to A. minutuloides , and the flagella and wing veins are darker compared to A. tkalcui .

Within the Levantine and Cypriot members of the A. minutula species group, the male of A. hebraica is characterized by a flagellomere 1 which is subequal to slightly longer than 3, clypeus and scutum which are partly to fully shagreened and densely punctured, tergal discs 2 and 3 which are more strongly shagreened basally vs. apically, and characteristic genitalia lacking dorsal gonoxocite lobes and with gonostyli that converge apically. The genitalia are almost identical to those of A. povolnyi Warncke stat. nov. from Afghanistan, but they clearly differ from other similar species in the region of study as follows: lack of dorsal gonocoxite lobes (present in A. cervina and A. spreta ), gonostyli more strongly converging apically compared to A. spreta and A. aphroditae , gonostyli slenderer compared to A. aphroditae , gonostyli shorter and penis valves shorter and narrower compared to A. tiaretta (Figs 20 J View Figure 20 , 21 G, I, J View Figure 21 ).

Description.

Female. Body length: 5.5–6.5 mm. — Integumental colour: Body and legs predominantly black. Flagellum black, partly covered by grayish setae, distal flagellomeres occasionally brownish. Apical tarsomeres brown to black. Wings weakly infuscate, veins brown, stigma brownish centrally (Fig. 9 A View Figure 9 ). Tergal marginal zones black, apical margins black to brown (Fig. 9 D View Figure 9 ). — Pubescence: Body and leg hair mostly brightly coloured. Clypeus with sparse, short and thin, semi-erect white hairs. Supraclypeal area with short whitish hairs. Paraocular area and scape with dense, medium-lengthed whitish hairs (Fig. 9 A – C View Figure 9 ). Frons with sparse, medium-lengthed whitish hairs. Facial fovea in dorsal view with brownish hair on upper ⅓ of fovea, white hair on lower ⅔ (Fig. 9 B, C View Figure 9 ). Preoccipital ridge with long erect whitish hair. Genal area with whitish hair, short on dorsal part, medium to long on ventral part (Fig. 9 A – C View Figure 9 ). Scutum, scutellum and metanotum with moderately dense, erect whitish to yellowish hair, centrally minute to short, with some scattered longer hairs, underlying cuticle visible, peripherally short to medium-lengthed (Fig. 9 A, C View Figure 9 ). Mesepisternum with long whitish hair (Fig. 9 A View Figure 9 ). Propodeal corbicula incomplete, posterodorsal fringe with long whitish plumose hairs, corbicular surface with sparse, long simple whitish hairs. Leg hair mostly whitish, some golden reflections appear especially on tarsi. Flocculus incomplete, white. Tibial scopal hairs simple, whitish, becoming brownish near posterobasal edge of tibia (Fig. 9 A View Figure 9 ). Tergal discs centrally with sparse, inconspicuous minute thin hair, laterally with sparse short to medium white hair. Tergal marginal zones 2–4 with distinct bands of white hair, moderately dense and broadly interrupted on 2–3, sparser but almost continuous on 4. Terminal fringe golden to brownish, flanked by some long white hairs (Fig. 9 A, D View Figure 9 ). — Head: 1.2 times broader than long. Mandibles bidentate, moderately crossed. Galea finely shagreened (Fig. 9 B View Figure 9 ). Labral process fairly large, trapezoidal, not much broader than long. Clypeus weakly domed, surface entirely finely shagreened and matt, except for an occasional shinier apical margin, punctation moderately strong, distance between punctures 1–2 puncture diameters, an impunctate midline is not clearly discernible (Figs 9 B View Figure 9 , 17 E View Figure 17 ). Supraclypeal area finely longitudinally striated. Lowermost part of paraocular area smooth and shiny, densely and finely punctured. Frons and rest of paraocular area strongly longitudinally striated, some interspersed punctures can occasionally be discerned on frons. Flagellomere 1 as long as 2 + 3, 2 about as long as 3 (Fig. 9 B View Figure 9 ). Facial foveae narrow and elongate, extending from level of lower end of lateral ocellus to slightly above base of clypeus, upper part 0.4–0.5 times as broad as antennocular distance, lower ⅔ distinctly narrowed, here separated from compound eye by distinct strip of smooth cuticle. Distance of fovea from lateral ocellus about 2 ocellus diameters (Fig. 9 B, C View Figure 9 ). Lateral ocelli connected posteriorly by trough-shaped horizontal furrow. Ocelloccipital distance about 0.9 ocellus diameter. Preoccipital ridge weakly to distinctly carinate (Fig. 9 C View Figure 9 ). — Mesosoma: Dorsolateral angle of pronotum not elevated. Entire scutum strongly shagreened and matt, densely punctured, distance between punctures 0.5–1.5 puncture diameters, puncture strength varying among individuals. Scutellum similar, occasionally somewhat shinier (Figs 9 C View Figure 9 , 17 J View Figure 17 ). Mesepisternum strongly, finely reticulated, anterior part overlayed with dense, strong oblique punctures. Surface of propodeal corbicula finely reticulated, with few large punctures around hair bases. Posterior part of propodeum strongly differentiated into horizonal vs. vertical regions. Propodeal triangle delineated by lateral carinae, horizontal part strongly and coarsely rugose, radially rugose near base, rugose-areolate further down, vertical part very finely areolated (Fig. 9 C View Figure 9 ). Posterolateral part of propodeum strongly, finely rugose-areolate. Hind pretarsal claw with inner tooth. Recurrent vein 1 meets submarginal cell 2 at its middle or slightly proximal to it. Nervulus interstitial to weakly antefurcal (Fig. 9 A View Figure 9 ). — Metasoma: Tergal disc 1, basal part of disc 2 and occasionally of 3 strongly shagreened and matt, impunctate. Tergal disc 4, apical part of tergal disc 2 and at least apical part of disc 3 shiny, partly to fully smooth, the tight shagreening of basal areas here gradually disintegrating into irregular, dense fine punctation, distance between punctures up to 1 puncture diameter. Tergal marginal zones weakly depressed, impunctate, 2–3 centrally occupying 0.4–0.5 of tergum length, 1–2 distinctly shagreened, the following gradually more weakly so (Fig. 9 D View Figure 9 ). Pygidial plate normally developed. — Male. Body length: 5–6 mm. — Integumental colour: As in female (Fig. 9 E, H View Figure 9 ). — Pubescence: Clypeus with moderately dense, medium-lengthed, semi-erect white hairs, underlying cuticle visible (Fig. 9 E, F View Figure 9 ). Supraclypeal area with short erect whitish hair. Paraocular area with moderately dense, medium-lengthed erect hair, whitish to black on outer part bordering compound eye, whitish on inner part. Frons and paraocular area with sparse, erect short to medium hair of varying brightness, usually with predominantly dark hairs on dorsal parts (Fig. 9 F, G View Figure 9 ). Preoccipital ridge with long erect whitish hairs. Genal area often with few short to medium erect dark hairs around upper margin of compound eye, elsewhere with whitish hairs, short dorsally, gradually long ventrally (Fig. 9 E – G View Figure 9 ). Scutum, scutellum and metanotum with moderately dense erect whitish hair, centrally with mixed short and long hairs, underlying cuticle visible, peripherally medium to long. Mesepisternum and propodeum with very long white hairs (Fig. 9 E, G View Figure 9 ). Leg hair mostly white, with some subtle golden reflections mostly on tarsi (Fig. 9 E View Figure 9 ). Tergal discs centrally with minute inconspicuous bright hair, laterally and on base of disc 1 with moderately dense, medium-lengthed white hair. Tergal marginal zones 2–4 with strong, broadly interrupted bands of white hair, slightly extending onto following discs. Tergal marginal zone 5 with very sparse, continuous band of long whitish hair. Terminal fringe whitish (Fig. 9 E, H View Figure 9 ). — Head: About 1.2 times broader than long (Fig. 9 F View Figure 9 ). Mandibles bidentate, moderately crossed. Galea finely shagreened. Labral process trapezoidal to almost rectangular, apical margin concave. Clypeus moderately domed, strongly shagreened and matt basally, shiny and weakly shagreened apically, extent of shagreening highly variable among individuals, punctation moderately dense, distance between punctures 0.5–1.5 puncture diameters, without impunctate midline. Outer part of paraocular area bordering compound eye smooth and finely punctured. Supraclypeal area, frons and inner part of paraocular area longitudinally striated, occasionally with hints of punctures blended into the rugosity. Flagellomere 1 1.1–1.3 times longer than 3, distinctly shorter than 2 + 3, 2 distinctly shorter than 3 (Fig. 9 F View Figure 9 ). Ocelloccipital distance about 1 ocellus diameter. Preoccipital ridge moderately carinate (Fig. 9 G View Figure 9 ). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum and scutellum usually strongly shagreened and matt, rarely weakly shagreened and shiny, punctation distinct, distance between punctures 0.5–1.5 puncture diameters (Fig. 9 G View Figure 9 ). Mesepisternum finely alveolated, overlayed by weak oblique punctures. Anterolateral part of propodeum ventrally reticulated, dorsally gradually rugose-areolate. Posterior part of propodeum rugose-areolate, triangle weakly delineated, radially rugose near base, coarsely rugose-areolate further down, very finely so on vertical part (Fig. 9 G View Figure 9 ). Recurrent vein 1 meets submarginal cell 2 at its middle or slightly proximal to it. Nervulus interstitial to weakly antefurcal (Fig. 9 E View Figure 9 ). — Metasoma: As in female (Fig. 9 H View Figure 9 ). — Genitalia and hidden sterna: Gonocoxites apically truncate, without dorsal lobe. Gonostyli more or less finger-shaped, inner and outer margins with slight kink at mid-length, distal part more strongly converging apically. Penis valves moderately narrow basally, gradually tapering apically (Fig. 21 J View Figure 21 ). Sternum 8 columnar, broadening apically, apical margin blunt to slightly rounded.

Distribution and habitat.

Mediterranean shrublands in Israel, the West Bank, Jordan and Lebanon. Mostly absent from the coastal plain, where it is replaced by A. tiaretta .

Flight period.

Late January to mid-May.

Flower records.

Collected mostly on Brassicaceae ( Hirschfeldia , Isatis, Ochtodium , Rapistrum , Sinapis ), also on Apiaceae ( Ferula ), Asteraceae ( Anthemis , Leontodon ), Campanulaceae ( Campanula ), Fabaceae ( Trifolium ) and Lamiaceae ( Nepeta , Salvia ).

Type material.

HOLOTYPE: ISRAEL ● 1 ♀; Nahal Keziv [Nahal Kziv] ; 33.0465°N 35.226°E; [ 197 m a. s. l.]; 26 Feb. 2021; G. Pisanty leg.; BOLD accession no. ANDIL 490-25; SMNHTAU 357574 GoogleMaps . – PARATYPES: ISRAEL ● 2 ♂; Ahu Binyamina Nature Reserve; 32°30’N 34°56’E; 8 Mar. 2017; A. Freidberg leg.; SMNHTAU 269485 , 269486 ● 1 ♀; Avivim; 25 Apr. 1983; A. Hefetz leg.; SMNHTAU 353608 ● 3 ♀; Banias; 18 Apr. 1992; R. Kasher leg.; SMNHTAU 353683 to 353685 ● 1 ♀; Bar’am; 4 Apr. 2014; O. Winberger leg.; SMNHTAU 183658 ● 1 ♀; Basmat Tab’un; 14 Apr. 1999; L. Friedman leg.; SMNHTAU 367678 ● 1 ♀; Beit Govrin; 15 Apr. 2017; T. Roth leg.; SMNHTAU 272634 ● 1 ♀; ibid.; 20 Apr. 2017; SMNHTAU 272874 ● 1 ♂; Beit Nir; 11 Mar. 2018; T. Roth leg.; SMNHTAU 290320 ● 1 ♀; ibid.; on Apiaceae ; RMNH ● 1 ♀; ibid.; on Anthemis ; SMNHTAU 290302 ● 3 ♀; Bene Yehuda – Golan; 31 Mar. 1991; R. Kasher leg.; on Trifolium purpureum ; SMNHTAU 353736 to 353738 ● 1 ♂; ibid.; SMNHTAU 353509 ● 1 ♂; Bet Guvrin; 10 Mar. 2004; A. Freidberg leg.; SMNHTAU 353483 ● 1 ♀; Biriyya Forest; 32.99– 33.00°N 35.52– 53°E; 19 May 2023; G. Pisanty leg.; SMNHTAU 429812 ● 1 ♀; Carmel; 12 Mar. 1940; H. Bytinski-Salz leg.; SMNHTAU ● 1 ♂; Deganya A, Bet Gordon; 3 Mar. 1942; Y. Palmoni leg.; on Diplotaxis & Sinapis alba ; SMNHTAU 180731 ● 3 ♀; ‘ En Tina NR; 33.08417°N 35.642°E; 70–80 m a. s. l.; 24 Mar. 2019; A. Dorchin, T. Roth & A. Sviri leg.; SMNHTAU 310136 , 310227, 310279 ● 1 ♂; ‘ En Ziwan, camping cite [site]; 955 m a. s. l.; 16 Mar. 2020; L. Friedman leg.; SMNHTAU 332994 ● 1 ♀; Gal’on; 4 Apr. 2018; T. Roth leg.; on Isatis lusitanica ; SMNHTAU 290448 ● 1 ♀; ibid.; on Hirschfeldia incana ; SMNHTAU 290473 ● 1 ♀; Gal’on, 2 kmNW; 31.649°N 34.837°E; 5 Apr. 2015; G. Pisanty leg.; SMNHTAU 209389 ● 1 ♀; Gamla; 29 Apr. 1988; I. Yarom leg.; SMNHTAU 353715 ● 3 ♀; Gilat Forest; 31.341°N 34.654°E; 1 Mar. 2023; G. Pisanty leg.; sweep; SMNHTAU 422802 to 422804 ● 1 ♀; Gilat Forest; 31.35°N 34.66°E; 2 Mar. 2022; G. Pisanty leg.; p. trap; SMNHTAU 386118 ● 1 ♂; Gilat Research Center, 500 m NNE; 31.341°N 34.6693°E; 2 Mar. 2022; G. Pisanty leg.; pan trap; SMNHTAU 386292 ● 1 ♂; Gilat Research Center, 500 m NNE, fallow field; 31.3405°N 34.670°E; 2 Mar. 2022; G. Pisanty leg.; sweep; SMNHTAU 385044 ● 1 ♂; ibid.; NHMUK ● 1 ♂; Gonen, 21 km NW Zefat; 10 Mar. 1990; R. Kasher leg.; on Sinapis alba ; SMNHTAU 353508 ● 1 ♀; HaGolan, N Nahal Yadbir; 33.052°N 35.646°E; 120–130 m a. s. l.; 24 Mar. 2019; A. Dorchin, T. Roth & A. Sviri leg.; SMNHTAU 310202 ● 1 ♀; HaGolan, S Nahal Neshef; 33.092°N 35.644°E; 100 m a. s. l.; 24 Mar. 2019; Dorchin, T. Roth & A. Sviri leg.; SMNHTAU 310151 ● 1 ♀; Haifa Univ.; 10 Apr. 1988; I. Yarom leg.; SMNHTAU 353713 ● 1 ♂; Hanita; 27 Mar. 1976; D. Gerling leg.; SMNHTAU 353469 ● 3 ♀; Har Ahino’am; 32.502– 6°N 35.408– 14°E; 440 m a. s. l.; 3 Mar. 2025; G. Pisanty leg.; SMNHTAU 463990 , 463994, 464001 ● 4 ♂; ibid.; SMNHTAU 463946 , 463947, 463951, 463952 ● 2 ♂; ibid.; OLML ● 1 ♂; ibid.; ZMB ● 1 ♀; ibid.; on yellow Asteraceae ; SMNHTAU 463935 ● 3 ♀; Har Karmila; 31.78– 79°N 35.02– 04°E; 5 Mar. 2017; G. Pisanty leg.; SMNHTAU 268883 , 268884, 268886 ● 1 ♂; ibid.; SMNHTAU 268882 ● 1 ♀; Har Meron; 32.9949°N 35.4166°E; 21 Apr. 2017; G. Pisanty leg.; SMNHTAU 270981 ● 1 ♀; Har Tuv; 3 Mar. [19] 54; Kadman leg.; SMNHTAU 353581 ● 1 ♂; Hare Gilboa’, Har Ahino’am; 32.502°N 35.414°E; 450 m a. s. l.; 23 Feb. 2012; L. Friedman leg.; SMNHTAU 118603 ● 1 ♀; HareGilboa’, Merav, HarAvinadav; 32°27.7’N 35°26’E; 420 m a. s. l.; 11 Apr. 2011; A. Freidberg leg.; SMNHTAU 91746 ● 1 ♂; ibid.; BOLD accession no. ANDIL 512-25; SMNHTAU 91745 ● 2 ♀; Harel; 10 Mar. 2020; T. Roth leg.; SMNHTAU 346514 , 346557 ● 1 ♀; ibid.; 21 Feb. 2020; on Isatis lusitanica ; SMNHTAU 338454 ● 1 ♂; Hof Rosh Haniqra N. R.; 33.076– 85°N 35.105– 9°E; 18 Mar. 2025; G. Pisanty leg.; SMNHTAU 464967 ● 1 ♀; Horbat Kefar Lakhish; 31.575°N 34.853°E; 15 Mar. 2021; G. Pisanty leg.; pan trap; SMNHTAU 358991 ● 2 ♀; ibid.; 31.575°N 34.8532°E; 5 Mar. 2021; sweep; SMNHTAU 358851 , 358861 ● 1 ♂; ibid.; SMNHTAU 358864 ● 1 ♀; Hulda; 23 Mar. 2017; T. Roth leg.; SMNHTAU 272678 ● 2 ♂; Jer [usalem?]; 27 Feb. 1940; SMNHTAU ● 1 ♀; ibid.; 10 Apr. 1940 ● 1 ♀; Jerusalem; 28 Jan. 1940; H. Bytinski-Salz leg.; SMNHTAU ● 1 ♀; ibid.; 10 Feb. 1940 ● 1 ♀; ibid.; 7 Mar. 1940 ● 1 ♀; ibid.; 24 [or 29?] Mar. 1940 ● 1 ♀; ibid.; 18 Apr. 1940 ● 1 ♀; Jerusalem, Mt. Scopus; 19 Apr. 1945; SMNHTAU ● 1 ♀; Jerusalem, Mt. Scopus – U. [niversity?]; 15 Mar. 1946; SMNHTAU ● 1 ♀; Jish; 29 Mar. 2014; O. Winberger leg.; SMNHTAU 183606 ● 1 ♂; Kabri; 17 Mar. 1973; M. Kaplan leg.; SMNHTAU 353463 ● 1 ♀; Kefar Giladi S; 1 Apr. 1997; R. Kasher leg.; SMNHTAU 353756 ● 1 ♂; ibid.; SMNHTAU 353550 ● 2 ♀; ibid.; 12 Apr. 1997; SMNHTAU 353767 , 353768 ● 1 ♂; ibid.; SMNHTAU 353549 ● 1 ♀; Kfar Masaryk; 32.87°N 35.13°E; 4 Apr. 2023; E. L. A. E. leg.; pan trap; SMNHTAU 462866 ● 1 ♀; Kfar Menahem; 1 Mar. 2008; G. Pisanty leg.; SMNHTAU 31528 ● 2 ♀; Kfar Menahem; 18 Mar. 2018; T. Roth leg.; on Sinapis alba ; SMNHTAU 276683 , 276686 ● 1 ♀; ibid.; RMNH ● 1 ♂; ibid.; on Ferula communis ; SMNHTAU 290271 ● 1 ♂; ibid.; on Campanula rapunculus ; SMNHTAU 276679 ● 2 ♀; Kfar Shammay; 13 Apr. 1988; I. Yarom leg.; SMNHTAU 353717 , 353718 ● 1 ♀; Kiryat Gat; 19 Apr. 1977; A. Freidberg leg.; SMNHTAU 353617 ● 2 ♀; Kokhav HaYarden, moat of castle; 26 Mar. 2001; L. Friedman leg.; SMNHTAU ● 1 ♀; Lahav; 27 Feb. 1974; A. Freidberg leg.; SMNHTAU ● 2 ♀; Lakhish; 18 Feb. 2013; T. Shapira leg.; SMNHTAU 133191 , 133245 ● 1 ♂; ibid.; SMNHTAU 133247 ● 1 ♀; ibid.; 19 Mar. 2013; SMNHTAU 150243 ● 1 ♀; Lakhish, 3 km NE; 31.575°N 34.870°E; 11 Mar. 2016; G. Pisanty leg.; SMNHTAU 236050 ● 1 ♀; ibid.; 31.578°N 34.870°E; 19 Feb. 2016; SMNHTAU 234220 ● 1 ♂; ibid.; SMNHTAU 234219 ● 2 ♀; ibid.; 26 Feb. 2016; SMNHTAU 234630 , 234640 ● 1 ♂; ibid.; 4 Mar. 2016; BOLD accession no. ANDIL 426-25; SMNHTAU 235274 ● 1 ♀; ibid.; 31.579°N 34.871°E; on Sinapis ; SMNHTAU 235195 ● 1 ♀; Lehavim, 400 m SE; 31.368°N 34.829°E; 27 Jan. 2021; G. Pisanty leg.; SMNHTAU 354989 ● 1 ♂; Ma’ale Gamla; 32.885– 92°N 35.681– 4°E; 28 Feb. 2022; G. Pisanty leg.; SMNHTAU 385113 ● 2 ♀; Majdal Shams; 33.262°N 35.755°E; 1100 m a. s. l.; 27 Apr. 2020; G. Pisanty leg.; SMNHTAU 334295 , 334296 ● 1 ♀; Malkiyya; 27 Apr. 2014; N. Atkin leg.; SMNHTAU 182993 ● 6 ♀; Ma’yan Barukh; 33.232°N 35.611°E; 29 Mar. 2023; E. L. A. E. leg.; pan trap; SMNHTAU 462796 , 462799, 462808, 462809, 462815, 462816 ● 1 ♂; Me’arat Yishah, 0.5 km E; 32.718°N 35.007°E; 22 Feb. 2019; G. Pisanty leg.; BOLD accession no. ANDIL 466-25; SMNHTAU 321645 ● 2 ♀; Merom-Golan, N Golan Height; 30 Mar. 1997; R. Kasher leg.; ZMB ● 1 ♀; ibid.; 3 Apr. 1997; SMNHTAU 353752 ● 1 ♀; ibid.; 1000 m a. s. l.; 23 Apr. 1997; SMNHTAU 353751 ● 1 ♂; Monfort; 33°02.635’N 35°13.271’E; 4 Mar. 2010; A. Freidberg leg.; SMNHTAU 47055 ● 1 ♂; Montfort, Nahal Keziv; 28 Feb. 2018; G. Pisanty leg.; BOLD accession no. ANDIL 451-25; SMNHTAU 286331 ● 1 ♂; Moradot HaGolan NR, Nahal Neshef, Site B 3; 33.094°N 35.650°E; 184 m a. s. l.; 24 Apr. 2019; A. Dorchin, T. Roth & O. Halbershtat leg.; BOLD accession no. ANDIL 461-25; SMNHTAU 308947 ● 1 ♂; Moradot HaGolan NR, Tel ‘ Arfiyya, Plot A 2; 33.105°N 35.657°E; 264 m a. s. l.; 7 Apr. 2019; A. Dorchin, A. Sviri & O. Halbershtat leg.; BOLD accession no. ANDIL 460-25; SMNHTAU 308860 ● 1 ♀; Mt. Carmel, En Hod; 4 Apr. 1999; S. P. M. Roberts leg.; TJWC ● 1 ♀; Mt. Meron; 900 m a. s. l.; 14–15 Apr. 1999; S. P. M. Roberts leg.; TJWC ● 1 ♀; MtMeron; 1000 m a. s. l.; 13 Apr. 1988; I. Yarom leg.; SMNHTAU 353720 ● 2 ♂; Nachshon; 20 Feb. 2020; T. Roth leg.; on Leontodon tuberosus ; SMNHTAU 338538 , 338539 ● 1 ♀; Nachson; 20 Feb. 2020; T. Roth leg.; SMNHTAU 346691 ● 1 ♂; ibid.; SMNHTAU 346688 ● 8 ♀; Nahal Dishon; 1 Apr. 1991; R. Kasher leg.; SMNHTAU 353741 , 353742, 353744 to 353748, 353750 ● 1 ♂; ibid.; SMNHTAU 353749 ● 1 ♀; ibid.; on Ochtodium aegyptiacum ; SMNHTAU 353739 ● 1 ♂; Nahal Hazav, BitronotRuhama; 31°32’N 34°42’E; 5 Apr. 2005; A. Freidberg leg.; SMNHTAU 353496 ● 5 ♀; Nahal Maresha; 31.577°N 34.858°E; 220 m a. s. l.; 15 Mar. 2021; G. Pisanty leg.; sweep; SMNHTAU 359305 , 359306, 359315, 359316, 359319 ● 2 ♂; Nahal Mezar; 32.75– 7°N 35.69°E; 1 Mar. 2022; G. Pisanty leg.; sweeping; SMNHTAU 385149 , 385168 ● 2 ♂; ibid.; RMNH ● 1 ♀; Nahal Shemarya; 31.341°N 34.6567°E; 2 Mar. 2022; G. Pisanty leg.; pan trap; SMNHTAU 386162 ● 1 ♀; ibid.; OLML ● 3 ♀; ibid.; 31.341°N 34.6568°E; sweeping; OLML ● 1 ♀; ibid.; 31.3412°N 34.657°E; M. Ben-Yosef leg.; SMNHTAU 386227 ● 1 ♀; Nahal Zilzal; 31.7°N 35.03°E; 25 Feb. 2019; G. Pisanty leg.; SMNHTAU 321739 ● 1 ♂; ibid.; SMNHTAU 321751 ● 1 ♂; Nahshon; 25 Mar. 2010; G. Pisanty leg.; pan trap; BOLD accession no. ANDIL 507-25; SMNHTAU 60450 ● 1 ♀; Nahshon; 26 Mar. 2018; T. Roth leg.; on Sinapis alba ; SMNHTAU 290562 ● 1 ♀; Nahshon junction; 9 Mar. 2017; T. Roth leg.; SMNHTAU 276554 ● 1 ♀; ibid.; BOLD accession no. ANDIL 448-25; SMNHTAU 274104 ● 2 ♂; ibid.; SMNHTAU 274072 , 274119 ● 1 ♀; Nahshonim; 25 Apr. 2023; S. Asis leg.; SMNHTAU 414794 ● 1 ♂; Netanya, Irus Ha’Argaman NR; 24 Feb. 2021; ITI Bee Course leg.; SMNHTAU 366434 ● 1 ♀; Netiv Halamed He; 26 Feb. 2009; G. Pisanty leg.; SMNHTAU 27558 ● 1 ♀; Netiv Halamed Heh; 27 Mar. 2017; T. Roth leg.; SMNHTAU 272976 ● 1 ♀; Park Britannia; 24 Apr. 2011; T. Koznichki leg.; pan trap; SMNHTAU 354431 ● 1 ♀; ibid.; 24 Mar. 2016; T. Chaprazaro leg.; SMNHTAU 251264 ● 1 ♀; Park Britanya; 7 Apr. 2010; T. Koznichki leg.; pan trap; SMNHTAU 59443 ● 1 ♀; Pura Nature Reserve; 31°29.7’N 34°46.8’E; 200 m a. s. l.; 18 Mar. 2010; A. Freidberg leg.; SMNHTAU 48496 ● 1 ♂; Pura Nature Reserve; 31.496°N 34.778°E; 27 Mar. 2015; G. Pisanty leg.; BOLD accession no. ANDIL 419-25; SMNHTAU 208231 ● 1 ♂; Qedma; 17 Feb. 2010; G. Pisanty leg.; pan trap; SMNHTAU 31849 ● 1 ♀; R. [amat] Hanadiv; 4 Mar. 1990; R. Kasher leg.; SMNHTAU 353631 ● 1 ♂; Ramat HaNadiv; 32.54– 6°N 34.94– 6°E; 16 Mar. 2023; G. Pisanty leg.; SMNHTAU 425831 ● 1 ♀; Ravid; 32.847°N 35.468°E; 29 Mar. 2023; E. L. A. E. leg.; pan trap; SMNHTAU 462768 ● 1 ♀; Rd. 918, Nahal Gonen; 33.111°N 35.645°E; 76 m a. s. l.; 24 Mar. 2019; A. Dorchin, T. Roth & A. Sviri leg.; SMNHTAU 310180 ● 8 ♀; Rekhes HaSullam; 33.091°N 35.112°E; 80 m a. s. l.; 18 Mar. 2025; G. Pisanty leg.; SMNHTAU 464898 to 464900, 465057 to 465061 ● 3 ♀; ibid.; L. Friedman leg.; SMNHTAU 464905 to 464907 ● 2 ♀; Revadim; 15 Mar. 2018; T. Roth leg.; on Sinapis alba ; NHMUK ● 2 ♀; ibid.; 27 Mar. 2018; SMNHTAU 290959 , 290960 ● 3 ♀; ibid.; RMNH ● 7 ♀; ibid.; on Ferula communis ; SMNHTAU 290935 , 290936, 290945 to 290947, 290990, 290993 ● 1 ♀; ibid.; RMNH ● 1 ♀; Road 869, 2.12 Km NE Ma’ale Gamla Jcn, Plot D 8; 32.895°N 35.676°E; – 51 m a. s. l.; 11 Apr. 2019; A. Dorchin, Y. Mersman & O. Halbershtat leg.; SMNHTAU 307424 ● 1 ♀; Road 87, 260 m E Yehudiyya Jcn, Plot D 6; 32.902°N 35.650°E; – 160 m a. s. l.; 11 Apr. 2019; A. Dorchin, Y. Mersman & O. Halbershtat leg.; SMNHTAU 307447 ● 2 ♀; Rosh Ha’ayin Forest; 32.1013°N 34.9695°E; 28 Mar. 2022; G. Pisanty leg.; sweeping; SMNHTAU 388615 , 388616 ● 1 ♀; Sasa; 11 Apr. 2014; O. Winberger leg.; SMNHTAU 183027 ● 3 ♀; Sha’alvim; 16 Mar. 2017; T. Roth leg.; SMNHTAU 272535 , 276419, 276430 ● 2 ♂; ibid.; SMNHTAU 272532 , 272534 ● 1 ♀; Shoham; 26 Mar. 2014; E. Morgulis & A. Freidberg leg.; SMNHTAU 179267 ● 1 ♀; Shoham Forest Park; 32.00°N 34.963°E; 28 Mar. 2022; G. Pisanty leg.; sweeping; SMNHTAU 388587 ● 1 ♂; Snir - Hermon Field Study Center; 27 Mar. 1997; R. Kasher leg.; SMNHTAU 353558 ● 1 ♀; Snir, Hermon Field Study Center; 13 Mar. 1997; R. Kasher leg.; SMNHTAU 354450 ● 1 ♂; ibid.; 1 Apr. 1997; SMNHTAU 353560 ● 1 ♀; ibid.; 3 Apr. 1997; SMNHTAU 353788 ● 1 ♀; ibid.; 18 Apr. 1997; SMNHTAU 353781 ● 1 ♀; ibid.; 21 Apr. 1997; SMNHTAU 353782 ● 1 ♂; Tal Shachar; 20 Mar. 2017; T. Roth leg.; BOLD accession no. ANDIL 445-25; SMNHTAU 273153 ● 1 ♂; Tanur; 15 Mar. 1975; F. Kaplan leg.; SMNHTAU 354489 ● 1 ♀; Tel Aviv; 32.0990°N 34.8042°E; 10–24 Apr. 2024; E. L. A. E. leg.; Malaise trap; SMNHTAU 462740 ● 2 ♀; Tel ‘ Eton; 31.492°N 34.925°E; 3 Apr. 2022; G. Pisanty leg.; sweeping; SMNHTAU 388491 , 388493 ● 1 ♀; Tel Sokho; 31°41’N 34°58’E; 290 m a. s. l.; 10 Apr. 2016; L. Friedman leg.; SMNHTAU 239592 ● 1 ♀; Tel Zafit; 31 Mar. 2018; T. Roth leg.; on Rapistrum rugosum ; SMNHTAU 290882 ● 7 ♀; Tiberias; 32.765°N 35.523°E; 27 Mar. 2023; E. L. A. E. leg.; pan trap; SMNHTAU 462969 , 462971, 462973, 462977, 462978, 462983, 462990 ● 1 ♀; W. Habiz [Nahal Kziv], Mt. Meron; 8 Apr. 1972; Zultan leg.; SMNHTAU 353615 ● 2 ♀; Ya’ar Adulam; 2013; Y. Berner leg.; on Nepeta curviflora ; SMNHTAU 151548 , 151555 ● 2 ♀; Ya’ar ‘ Adullam; 20 Apr. 2011; T. Koznichki leg.; pan trap; SMNHTAU 354424 , 354425 ● 1 ♀; Ya’ar Kedoshim; 23 Mar. 2014; N. Shamir leg.; SMNHTAU 152640 ● 1 ♀; ibid.; 13 Apr. 2014; SMNHTAU 152648 ● 1 ♀; ibid.; 2 Mar. 2014; on Sinapis alba ; SMNHTAU 152576 ● 1 ♀; ibid.; 21 Mar. 2014; SMNHTAU 152566 ● 1 ♀; ibid.; 30 Mar. 2014; SMNHTAU 152545 ● 1 ♀; ibid.; 6 Mar. 2014; on Sarcopoterium spinosum ; SMNHTAU 152571 ● 1 ♀; Ya’ar Kedoshim; 7 Mar. 2017; Y. Farago leg.; SMNHTAU 272264 ● 1 ♂; Ya’ar Nehosha; 18 Mar. 2015; T. Chaprazaro leg.; SMNHTAU 184896 ● 1 ♀; Ya’ar Nehusha; 25 Apr. 2011; T. Koznichki leg.; pan trap; SMNHTAU 354436 ● 1 ♀; Ya’ar Odem NR; 33.205°N 35.736°E; 27 Apr. 2020; G. Pisanty leg.; pan trap; BOLD accession no. ANDIL 479-25; SMNHTAU 334583 ● 1 ♂; ibid.; BOLD accession no. ANDIL 478-25; SMNHTAU 334551 ● 1 ♀; Ya’ar Yish’i; 26 Apr. 2011; T. Koznichki leg.; pan trap; SMNHTAU 81398 ● 1 ♀; Yarkon; 27 Mar. 1996; T. Kimchi leg.; SMNHTAU ● 1 ♀; Yerushalayim [ Jerusalem]; 13 Apr. 2014; I. Arar; on Anthemis ; SMNHTAU 184498 ● 1 ♀; ibid.; on Sinapis arvensis ; SMNHTAU 184551 ● 1 ♀; Yiftach; 5 Apr. 2014; O. Winberger leg.; SMNHTAU 183181 ● 1 ♀; ibid.; 17 Apr. 2014; N. Atkin leg.; on Sinapis ; SMNHTAU 183159 ● 1 ♀; Zekharya; 3 Apr. 1988; I. Yarom leg.; SMNHTAU 353712 ● 1 ♀; Ziv’on; 17 Apr. 2012; L. Friedman leg.; SMNHTAU 124704 ● 1 ♀; Ziv’on, 1 kmSW; 33.019°N 35.407°E; 5 Apr. 2016; G. Pisanty leg.; BOLD accession no. ANDIL 427-25; SMNHTAU 239289 ● 1 ♂; Zomet Haela; 4 Apr. 1999; O. Greif leg.; SMNHTAU 353481 ● 1 ♂; Zomet Oren; 8 Apr. 1988; I. Yarom leg.; SMNHTAU 353506 ● 1 ♂; Zomet Oren, 10 km S Haifa; 8 Apr. 1988; C. O’Toole leg.; “ Andrena petraensis War. ” paratype label; OUMNH ● 1 ♀; כרמל [Carmel]; 1 May [19] 72; Kuty Yefenof leg.; SMNHTAU 353593 . – JORDAN ● 2 ♀; S of At Tafila; 27–30 Mar. 2013; M. Snižek leg.; OUMNH. – LEBANON ● 1 ♀; Beqaa, Anjar, 1 km E, reforestation area; 33.7311°N 35.9478°E; 1000 m a. s. l.; 7 Apr. 2023; T. Wood leg.; TJWC ● 1 ♀; Beqaa, Beqaa valley, Qaraoun, 3.5 km W of Madjal Balhis; 33.5377°N 35.7038°E; 900 m a. s. l.; 5 Apr. 2023; T. Wood leg.; BOLD accession no. WPATW 1108 -23; TJWC ● 1 ♂; Beqaa, Beqaa valley, Qob Elias, valley 500 m NW; 33.7989°N 35.8192°E; 900 m a. s. l.; 3 Apr. 2023; T. Wood leg.; TJWC. – WEST BANK ● 2 ♀; Berakha, 1 kmS, ‘ Amassa Spring; 595 m a. s. l.; 6 Mar. 2015; L. Friedman leg.; SMNHTAU 206346 , 206406 ● 1 ♂; ibid.; BOLD accession no. ANDIL 416-25; SMNHTAU 206347 ● 2 ♀; Deir Nidham Nat. Res.; 32.00°N 35.07°E; 420–530 m a. s. l.; 30 Mar. 2014; L. Friedman leg.; SMNHTAU 176991 , 176994 ● 3 ♀; Har-Gilo; 14 Apr. 1991; R. Kasher leg.; 10 min sweeping; SMNHTAU 353688 to 353690 ● 1 ♀; ibid.; 17 Apr. 1991; SMNHTAU 353691 ● 1 ♂; ibid.; 14 Apr. 1991; on Salvia fructosa [ fruticosa]; SMNHTAU 353507 ● 1 ♀; Har Gilo, 5 km SW Jerusalem; 850 m a. s. l.; 30 Mar. 1989; C. O’Toole leg.; “ Andrena petraensis War. ” paratype label; OUMNH ● 1 ♀; Herodyon; 31°40’N 35°14’E; 31 Mar. 2009; M. Guershon leg.; SMNHTAU 354461 ● 2 ♀; Itamar, 5 kmE, ThreeSeas’ Lookout; 860 m a. s. l.; 5 Mar. 2021; L. Friedman leg.; SMNHTAU 357727 , 357733 ● 1 ♀; Jeriho; 20–28 Apr. [19] 27; Dr. Enslin leg.; SMNHTAU ● 2 ♀; Nahal Teqoa’; 31°38’N 35°14’E; 650 m a. s. l.; 31 Mar. 2009; L. Friedman leg.; SMNHTAU 353699 , 353701 ● 1 ♀; ibid.; M. Guershon leg.; SMNHTAU 353700 ● 2 ♀; Qedumim, 1 kmSW, Mizpe ‘ Ami; 390 m a. s. l.; 13 Mar. 2015; L. Friedman leg.; SMNHTAU 206944 , 206947 ● 1 ♂; Shechem [Nablus]; 1 Mar. 1973; M. Kaplan leg.; SMNHTAU 353648 ● 1 ♀; Shilo, Nahal Shilo; 10 Apr. 2012; L. Friedman leg.; SMNHTAU 125335 ● 3 ♀; W. Faria [Nahal Tirza]; 1 Mar. 1973; M. Kaplan leg.; SMNHTAU 353626 , 353628, 354462 ● 2 ♂; ibid; SMNHTAU 353570 , 353574 ● 1 ♀; Wadi Faria; 8 Mar. 1973; A. Freidberg leg.; SMNHTAU ● 1 ♀; Zur Natan 800 mNE; 32.247°N 35.023°E; 23 Mar. 2020; G. Pisanty leg.; BOLD accession no. ANDIL 477-25; SMNHTAU 333737 .

RMNH

National Museum of Natural History, Naturalis

NHMUK

Natural History Museum, London

OLML

Oberösterreichisches Landesmuseum

ZMB

Museum für Naturkunde Berlin (Zoological Collections)

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Andrenidae

Genus

Andrena

SubGenus

Andrena