Amiga arnaca indianacristoi Nakahara & Marin

Nakahara, Shinichi, Lamas, Gerardo, Tyler, Stephanie, Marin, Mario Alejandro, Huertas, Blanca, Willmott, Keith R., Mielke, Olaf H. H. & Espeland, Marianne, 2019, A revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilioarnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae), ZooKeys 821, pp. 85-152 : 102-105

Amiga arnaca indianacristoi Nakahara & Marin subsp. n. Figs 2o, p, 6, 7

Description and diagnosis.

MALE: forewing length 18.5-21 mm (n = 6): Differs from the nominotypical subspecies in the following respects: dorsal hindwing bluish lilac reflection restricted to posterior one-thirds of hindwing and not extending into discal cell; ventral ground color appearing somewhat paler; discal band, postdiscal band, submarginal band on ventral forewing and ventral hindwing narrower; feeble pearly reflection on the ventral forewing (distal side of postdiscal band) absent or insignificant.

FEMALE: forewing length 20-22 mm (n = 3): Differs from the nominotypical subspecies by the characters provided for the male, except for the first DHW character.

Variation.

The area of the dorsal hindwing bluish-lilac reflection is variable, restricted only to the distal side of cells Cu1 and Cu2 in some specimens, whereas extending up to the area around M3 in some specimens. One female (FLMNH-MGCL 1036235) lacks the dorsal hindwing bluish-lilac reflection, while it is present in three other females (FLMNH-MGCL-264685, 1036240, 1036239).

Types.

HOLOTYPE ♂: // VENEZUELA: CARABOBO Hda. María Teresa (IAN), W of Las Trincheras 400 m, moist forest 28.vii.1981 Lee D. Miller, sta. VE16 // Allyn Museum Acc. 1981-23 // FLMNH-MGCL Specimen 264660 // (FLMNH).

PARATYPES (15 ♂, 4♀): Venezuela: Aragua: Rancho Grande Biological Station, [10°20'58"N, 67°41'3"W], 1100 m, (Heppner, J. B.), Jul 1981, 1 ♀ [FLMNH-MGCL-1036240], (FLMNH); km 22 Maracay–Ocumaré rd., Rancho Grande, 1100-1200 m, (Lichy, R.), 2 Aug 1943, 1 ♂ [FLMNH-MGCL-262940], (FLMNH); Maracay, Pozo del Diablo, [10°17'N, 67°37'W], 420-440 m, (Miller, L. D.), 22 Jul 1981, 1 ♂ [FLMNH-MGCL-264654], 1 ♂ [FLMNH-MGCL-264656], (FLMNH), 27 Jul 1981, 1 ♂ [FLMNH-MGCL-264655], (FLMNH); Maracay–Ocumare hwy., 1100-1200 m, (Lichy, R.), 4 Sept 1942, 1 ♀ [FLMNH-MGCL-1036235], (FLMNH); Carabobo: nr. Puerto Cabello, San Esteban, [10°26'N, 68°1'W], (Hahnel de Sagan), 1er trimestre 1877, 1 ♀ [BMNH(E)#1420075], (NHMUK), Jun, Jul 1877, 1 ♂ [BMNH(E)#1420068], 1 ♂ [BMNH(E)#1420069], (NHMUK); Yuma, [10°6'N, 67°42'W], 550 m, (Miller, L. D.), 16 Dec 1981, 1 ♂ [FLMNH-MGCL-1036241], 1 ♀ [FLMNH-MGCL-1036239], (FLMNH); Cojedes: El Baul, Hato Pinero, (Brenner, J.), Feb 1968, 1 ♂ [FLMNH-MGCL-1036242], (FLMNH); Distrito Federal: Caracas, [10°30'N, 66°55'W], 1200 m, (Forster, W.), 29 Jun 1954, 1 ♂, (ZSM); Miranda: 10 miles S of Caracas, [10°23'46"N, 66°53'26"W], 1311 m, 30 Mar 1970, 1 ♂ [woods], (USNM); [Caracas], Macizo Naiguatá, 720-800 m, (Lichy, R.), 6 Dec 1942, 1 ♂ [FLMNH-MGCL-262941], (FLMNH); Massif du Naiguatá, (Lichy, R.), 5 Jul 1945, 1 ♂ [FLMNH-MGCL-264683], (FLMNH); mist forest 20 km SE Caracas, [10°21'24"N, 66°46'18"W], 500-750 m, (Miller, L. D. & J. Y., Dukes, D.), 1 Sep 1990, 1 ♂ [FLMNH-MGCL-264657], 1 ♂ [FLMNH-MGCL-264659], (FLMNH); Río Chacaito, [10°25'N, 66°55'W], 980-1080 m, (Lichy, R.), 12 Sep 1936, 1 ♂ [FLMNH-MGCL-264658], (FLMNH).

Etymology.

This new species-group name is proposed in recognition of our friend and colleague, Indiana Cristóbal Ríos-Málaver, known as "Indiana Cristo", who studied the butterflies of the area where this taxon occurs. Indiana Cristo has contributed to Neotropical lepidopterology in various ways, especially through social media, where he is bringing lepidopterology to the public. This species-group name is treated as a latinized masculine noun in the genitive case.

Distribution

(Fig. 6). This taxon occurs in the Venezuelan Cordillera de la Costa and northwestern Cordillera de Mérida, and possibly also into the Serranía de Perijá.

Remarks.

We have examined a single specimen from Monagas, Venezuela (FLMNH-MGCL-264682), with a phenotype that corresponds to this subspecies. Thus, this taxon’s range may extend further east than that indicated above. However, given that we have examined only a single specimen from the area, combined with the fact that specimens from the island of Trinidad, whose butterfly fauna has strong biogeographic affinities with Monagas (e.g., Neild 1996), clearly represent the nominotypical subspecies, there still remains the possibility of this specimen being mislabeled.