Aleochara (Aleochara) coreana Bernhauer, 1926

Yamamoto, Shûhei & Maruyama, Munetoshi, 2016, Revision of the subgenus Aleochara Gravenhorst of the parasitoid rove beetle genus Aleochara Gravenhorst of Japan (Coleoptera: Staphylinidae: Aleocharinae), Zootaxa 4101 (1), pp. 1-68 : 10-20

Aleochara (Aleochara) coreana Bernhauer, 1926

( Figs 1 , 10 , 19 , 28 , 37 , 45–51 , 107–109 , 121 )

Aleochara coreana Bernhauer, 1926: 271 (original description; type locality: Korea, Pu?Ryong); Smetana, 2004b: 353 (as subgenus Aleochara ; catalogue of Palearctic species of Aleocharinae ); Park & Ahn, 2010a: 317 (as subgenus Aleochara ; in key to Far-Eastern species of subgenus Aleochara ); Park et al., 2011: 75 (as subgenus Aleochara ; redescription); Shibata et al., 2013: 105 (as subgenus Aleochara ; catalogue of Japanese species of Staphylinidae ); Schülke & Smetana, 2015: 490 (as subgenus Aleochara ; catalogue of Palearctic species of Aleocharinae ).

Aleochara lewisia Bernhauer, 1936: 324 [original description; type locality: Japan, Chiuzenji (Chûzenji, Nikkô-shi, Honshû)]; Likovský, 1973: 2 (as synonym of A. coreana ); Smetana, 2004b: 353 (as synonym of A. coreana ); Park et al., 2011: 75 (as synonym of A. coreana ); Shibata et al., 2013: 105 (as synonym of A. coreana ; catalogue of Japanese species of Staphylinidae ); Schülke & Smetana, 2015: 490 (as synonym of A. coreana ; catalogue of Palearctic species of Aleocharinae ). Synonymy confirmed.

Type material. A. coreana : Syntype, 1 male, “Para- / type [YRL] // koreana Brh / Cotypus. / Standinger don [HW] // Chicago NHMus / M. Bernhauer / Collection // Puryong / Korea [HW] // Brit. Mus /1956-709” (abdominal segments VIII–X and aedeagus dissected and mounted in Euparal by MM) ( BMNH); syntype, 1 male, “Co- / type [YRL] // Seishin / Korea // koreana Brh / Cotypus / Standinger [HW] // 1935.161 [HW]” (median lobe, broken, pulled off by somebody, mounted in Euparal by MM) ( BMNH); syntypes, 1 male, “Seishin / Korea // koreana Brh / Cotypus / Standinger [HW] // Chicago NHMus / M. Bernhauer / Collection” (median lobe pulled off by somebody, glued on PC) ( FMNH); syntype, 1 female, “Seishin / Korea // koreana Brh / Cotypus / Standinger [HW] // Chicago NHMus / M. Bernhauer / Collection” (spermatheca pulled off by somebody, glued on PC) ( FMNH); syntype, 1 male, “Seishin [HW] // Korea / lg. K. Rost. [HW] // koreana Brh / Cotypus / Standinger don [HW] // Chicago NHMus / M. Bernhauer / Collection // = COREANA Bernh. / det. Likovský 68’ [HW]” (median lobe pulled off by somebody, glued on PC with the body) ( FMNH).

A. lewisia : Syntype, 1 male, “Chiuzenji // Japan. / G. Lewis / B. M. 1926-369. // Aleochara / Lewisia Brnh. / M. Bernhauer det. / Type [HW]” (abdominal segments VIII-X and aedeagus dissected and mounted in Euparal by MM) (BMNH); syntype, 1 male, “Chiuzenji // Japan. / G. Lewis / B. M. 1926-369. // Lewisia Bernh. / Bernhauer det. / Cotype [HW: ‘ Lewisia Bernh.’, ‘Cotype’.] // BRIT. MUSEUM / DON. ARROW // Lewisia /Brnh. Cotype. / aleochara [HW] // Chicago NHMus M. Bernhauer / Collection // Aleochara / lewisia Bernhauer, 1936 / = coreana Bernhauer, 1936 / det. Z. Likovský 19 68 [HW: ‘Aleochara’, ‘ lewisia Bernhauer, 1936 ’, ‘= coreana Bernhauer, 1936 ’, only ‘[19] 68’.]” (median lobe pulled off by somebody, glued on PC with the body) (FMNH).

Non-type material. JAPAN: [HOKKAIDÔ]: 1 unsexed, Mashû-ko Lake, Teshikaga-chô, 5.vii.1958, M. Miyake leg. (EUM); 1 male, Sawanchisappu, Teshikaga-chô, 24.vii.1976, T. Matsuda leg. (EUM); 2 unsexed, Rikubetsu, Rikubetsu-chô, 30.vii.–2.viii.2001, H. Yoshitomi leg. (KUM); 1 female, Ashoro Research Forest of Kyushu Univesity, Ashoro-chô, 1–6.ix.2009, S. Yamamoto leg. (KUM); 2 unsexed, Nukabiragensenkyô, Kamishihoro-chô, 12.vii.2014, S. Yamamoto leg., by spraying insecticide on rotten wood (KUM); 3 unsexed, Nukabiragensenkyô, Kamishihoro-chô, 17.vii.2014, S. Yamamoto leg., by spraying insecticide on rotten wood (KUM); 3 unsexed, Nukabira-ko Lake, Kamishihoro-chô, 12.vii.2014, S. Yamamoto leg. (KUM); 15 unsexed, Nukabira-ko Lake, Kamishihoro-chô, 13.vii.2014, S. Yamamoto leg. (KUM); 1 female, Shihoro-kôgen Heights, Shihoro-chô, 19–25.vi.2009, T. Watanabe leg., from flight interception trap (cTW); 1 male, 1 unsexed, Osarushi, Otofuke-chô, 20–26.vi.2009, T. Watanabe leg., from flight interception trap (cTW); 1 unsexed, Pososanno-sawa (entrance of Notsuka Tunnel), Hiroo-chô, 13.vii.2002, K. Haga leg. (KUM); 6 males, 1 female, 14 unsexed, Nopporoshinrin-kôen Park, Ôsawaguchi, Ebetsu-shi, 30.vi.–13.vii.1993, S. Hori leg., from pitfall trap (KUM); 2 unsexed, Nopporoshinrin-kôen Park, Ôsawaguchi, Ebetsu-shi, 2.vi.1999, M. Maruyama leg. (KUM); 1 male, Nopporoshinrin-kôen Park, Ebetsu-shi, 13–21.vi.2000, M. Ôhara leg., from flight interception trap (KUM); 1 unsexed, Nopporo, Ebetsu-shi, 13–21.vi.2000, S. Hori leg., from flight interception trap (KUM); 1 unsexed, Nopporo, Ebetsu-shi, 5–11.vii.2000, S. Hori leg., from pitfall trap (KUM); 3 males, 1 female, 9 unsexed, Nopporo, Ebetsu-shi, 14.vi.1986, M. Ôhara leg. (KUM); 1 male, 3 unsexed, Nopporo, Ebetsu-shi, 17.vi.1986, M. Ôhara leg. (KUM); 1 male, Nopporo, Ebetsu-shi, 16.v.1987, M. Ôhara leg. (KUM); 1 unsexed, Nopporo, Ebetsu-shi, 12.vi.1987, M. Ôhara leg. (KUM); 1 male, 1 female, Nopporo, Ebetsu-shi, 17.viii.1990, M. Ôhara leg. (KUM); 1 female, Shinoro 3-jô 10-chôme 4, Kita-ku, Sapporo-shi, 24.vi.2009, H. Kamezawa leg., under nest of snowy heron (cHK); 1 female, 4 unsexed, Maruyama Park, Sapporo-shi, 12.vi.1980, H. Takemoto leg. (KUM); 1 unsexed, Maruyama Park, Sapporo-shi, 12.vi.1980, K. Maeto leg. (KUM); 2 unsexed, Maruyama Park, Sapporo-shi, 24.vii.2000, T. Watanabe leg. (cTW); 3 unsexed, Mt. Maruyama, Sapporo-shi, 25.vi.1986, M. Tomokuni leg. ( NSMT); 2 unsexed, Maruyama, Sapporo-shi, 9.vii.1953, Y. Kurosawa leg., NSMT-I-C 34340 and 34182, respectively ( NSMT); 13 unsexed, Maruyama, Sapporo-shi, 18.vii.1956, M. Miyake leg. (EUM); 1 unsexed, Maruyama, Sapporo-shi, 18.vii.1956, S. Ueda leg. (EUM); 19 unsexed, Minami 30 jô, Nishi 11 chôme (Mt. Moiwa-yama, near Moiwa power plant and filtration plant), Minami-ku, Sapporo-shi, 11–13.vi.1986, K. Haga leg., from baited trap (silkworm pupa powder) (KUM); 5 unsexed, Mt. Moiwa-yama, Sapporo-shi, 30.viii.1979, T. Watanabe leg. (cTW); 2 unsexed, Mt. Moiwa-yama, Sapporo-shi, 16.vi.1983, Y. Hirano leg. (KUM); 2 females, Bankei, Sapporo-shi, 24.vi.1977, N. Nishikawa leg. (KUM); 1 unsexed, Misumai, Sapporo-shi, 7.vii.1977, S. Naomi leg. (KUM); 2 males, 6 unsexed, Mt. Hakken-zan, Sapporo-shi, 12–14.viii.2001, H. Sugaya leg., from flight interception trap (KUM); 1 unsexed, Mt. Hakken-zan, Jôzan-kei, Sapporo-shi, 20.vi.1985, M. Ôhara leg. (KUM); 1 male, 1 unsexed, Jôzan-kei, Sapporo-shi, 25.v.1990, M. Ôhara leg. (KUM); 3 unsexed, Hôhei-kyô, Sapporo-shi, 30.vi.1958, M. Miyake leg. (EUM); 1 unsexed, Hyakumatsuzawa, Sapporo-shi, 8.vi.1998, M. Maruyama leg. (KUM); 1 male, 1 female, Nagahashinaebo, Otaru-shi, 24.v.1991, M. Ôhara leg. ( OCM); 15 males, 2 females, 4 unsexed, Nagahashinaebo, Otaru-shi, 31.v.1991, M. Ôhara leg. ( OCM); 1 unsexed, Nagahashinaebo, Otaru-shi, 7.vi.1991, M. Ôhara leg. ( OCM); 2 males, 1 unsexed, Nagahashinaebo, Otaru-shi, 15.vi.1991, M. Ôhara leg. ( OCM); 1 male, 1 female, Nagahashinaebo, Otaru-shi, 21.vi.1991, M. Ôhara leg. ( OCM); 3 females, Nagahashinaebo, Otaru-shi, 21.vi.1991, Y. Sasaki leg. ( OCM); 2 males, 1 female, 3 unsexed, Nagahashinaebo, Otaru-shi, 21.v.1992, M. Ôhara leg., from baited trap ( OCM); 1 male, 4 unsexed, Nagahashinaebo, Otaru-shi, 28.v.1992, M. Ôhara leg., from baited trap ( OCM); 3 unsexed, Nagahashinaebo, Otaru-shi, 4.vi.1992, M. Ôhara leg., from baited trap ( OCM); 1 male, 2 females, 29 unsexed, Nagahashinaebo, Otaru-shi, 12.vi.1992, M. Ôhara leg., from baited trap ( OCM); 1 female, Nagahashinaebo, Otaru-shi, 18.vi.1992, M. Ôhara leg., from baited trap ( OCM); 4 males, 7 females, 4 unsexed, Nagahashinaebo, Otaru-shi, 25.vi.1992, M. Ôhara leg., from baited trap ( OCM); 1 male, Nagahashinaebo, Otaru-shi, 11.vii.1992, M. Ôhara leg., from baited trap ( OCM); 1 male, Okusawa-suigenchi, Shiraisawa, Otaru-shi, 5.vii.1996, M. Ôhara & Y. Sasaki leg., from malaise trap ( HUM); 7 unsexed, Asahi, Otaru-shi, 8.vii.1995, T. Watanabe leg. (cTW); 1 unsexed, Tomakomai Experimenal Forest of Hokkaido University, Tomakomai-shi, 22.vi.1978, H. Oetani leg. (KUM); 2 unsexed, Tomakomai-shi, 30.vi.1979, H. Nomura leg. (KUM); 1 unsexed, Mt. Orofure-yama, 22.vii.1962, Y. Shibata leg. (KUM); 8 unsexed, Kojyôhama Beach, Shiraoi-chô, 23.vii.1962, Y. Shibata leg. (KUM); 1 unsexed, Sôbetsu-chô, 1.viii.1987, K. Yahiro leg. (KUM); 1 unsexed, Moheji, Hokuto-shi, 1.viii.2002, T. Yoshida leg. (KUM); 1 unsexed, Miyatsu, Okushiri-chô (Okushiri-tô Is.), 3.ix.1986, Y. Kusui leg. (cTI); 1 unsexed, Mt. Higure-yama (near Ônuma), Nanae-chô, 28.vi.1986, M. Tomokuni leg. ( NSMT); 2 males, 1 female, 53 unsexed, Ninnikuzawa-gawa River, Nanae-chô, 29.viii.1998, S. Saitou leg. (KUM); [HONSHÛ]: 6 unsexed, Ishimochi, Higashidôri-mura, Aomori-ken, 22.viii.2003, Y. Kawahara leg. (KUM); 1 unsexed, Mt. Asahina-dake, Mutsu-shi, Aomori-ken, 21.viii.2003, Y. Kawahara leg. (KUM); 1 unsexed, Yunomata, Ohata, Mutsu-shi, Aomori-ken, 30.vii.1956, K. Morimoto leg. (EUM); 2 unsexed, Ôkawashirisawa, Kazamaura-mura, Aomori-ken, 20.viii.2003, Y. Kawahara leg. (KUM); 30 unsexed, Arasawa, Sai-mura, Aomori-ken, 2–21.viii.2003, Y. Kawahara leg. (KUM); 2 unsexed, Fuyube, Nakadomari-machi, Aomori-ken, 29.v.2000, A. Abe leg. (KUM); 1 unsexed, Katagariishi, Nakadomari-machi, Aomori-ken, 13.viii.1993, A. Abe leg. (KUM); 1 unsexed, Katagariishi, Nakadomari-machi, Aomori-ken, 1.viii.2007, A. Abe leg. (KUM); 6 unsexed, Shikanoko-taki Fall, Kanagi-chô, Goshogawara-shi, Aomori-ken, 30.viii.2003, Y. Kawahara leg. (KUM); 1 unsexed, Mt. Chôbô-san, Goshogawara-shi, Aomori-ken, 22.viii.2003, Y. Kawahara leg. (KUM); 1 male, 3 unsexed, Ôwasawa, Hirosaki-shi, Aomori-ken, 23.v.1998, A. Abe leg. (KUM); 1 unsexed, Mt. Kudoji-yama, Hirosaki-shi, Aomori-ken, 21.vi.1981, A. Abe leg. (KUM); 4 unsexed, Rangannomori, Ajigasawa-machi, Aomori-ken, 26–31.vii.2003, Y. Kawahara leg. (KUM); 1 unsexed, Matsukami, Fukaura-machi, Aomori-ken, 16.vi.1996, A. Abe leg. (KUM); 24 unsexed, Jûniko Lake, Aomori-ken, 11.vi.1988, N. Itô leg. ( DEI); 1 unsexed, Kuzube-sawa, Iwate-ken, 22.vi.1980, S. Tsutsui leg. (KUM); 1 unsexed, Mt. Gando-san, Yamagata-shi, Yamagata-ken, 25.vii.1957, J. Kamei leg. (cYH); 2 unsexed, Arakai-gawa River (alt. 950m) ~ Mt. Arakai-san (north ridgeline; alt. 1170m), Fukushima-ken, 19.vii.2008, K. Haga leg., from forest floor of natural beech forest by sweeping (KUM); 3 unsexed, Kashii-onsen, Nishigô-mura, Fukushima-ken, 22.vi.1969, Y. Shibata leg. (KUM); 1 female, 270 unsexed, Kashii-onsen, Nishigô-mura, Fukushima-ken, 16–17.vi.1998, M. Maruyama leg., from carrion trap (KUM); 2 unsexed, Takigawa trail (Mt. Komaga-take), Hinoemata-mura, Fukushima-ken, 11– 12.viii.2007, K. Haga leg., from carcass of snake, Rhabdophis tigrinus (Boie) (KUM); 1 male, 13 unsexed, Oike, Hinoemata-mura, Fukushima-ken, 15.vi.1986, A. Yamato leg. (KUM); 1 unsexed, Hinoemata-mura, Fukushimaken, 21.vi.1970, T. Go leg. (KUM); 2 unsexed, Ashio-machi, Ibaraki-ken, 17.vii.1988, K. Kusano leg. (KUM); 1 unsexed, Nagasawa, Yunishikawa, Nikkô-shi, Tochigi-ken, 4.vii.2009, H. Kamezawa leg., from a bird carcass (cHK); 1 unsexed, Ôsawa, Okunikkô, Nikkô-shi, Tochigi-ken, 17.vii.1974, O. Yamaji leg. (KUM); 1 unsexed, Gojûri-ko Lake, Nikkô-shi, Tochigi-ken, 12.v.1985, K. Emoto leg. (cTW); 1 unsexed, Nikkô, Tochigi-ken, 17.vii.1957, Y. Kurosawa leg. ( NSMT); 1 unsexed, Watarase, Tochigi-ken, 20.iii.1983, M. Tao leg. (KUM); 1 female, 17 unsexed, Marunuma, Katashina-mura, Gunma-ken, 18–20.vi.2014, S. Yamamoto leg., from carrion trap (KUM); 26 unsexed (3 specimens preserved in 99.5% EtOH for DNA sequencing), Marunuma, Katashina-mura, Gunma-ken, 19–20.vi.2014, S. Yamamoto leg., from carrion trap (KUM); 1 unsexed, Seno-sawa, Takaragawa, Minakami-machi, Gunma-ken, 1.viii.1998, H. Kamezawa leg., alt. 800m (cHK); 3 unsexed, Hôshi-onsen, Minakami-machi, Gunma-ken, 27.vii.1966, Y. Shibata leg. (KUM); 1 unsexed, Sawairi, Midori-shi, Gunma-ken, 8.v.2005, T. Horiguchi leg. (cTW); 1 unsexed, near Shima, Nakanojô-machi, Gunma-ken, 12.vi.1986, T. Matsumoto leg. (KUM); 1 unsexed, Ôto, Higashiazuma-machi, Gunma-ken, 22.v.2005, T. Horiguchi leg. (cTW); 1 unsexed, Ôto, Higashiazuma-machi, Gunma-ken, 25.vi.2005, T. Horiguchi leg. (cTW); 1 unsexed, Fujimi (Mt. Akagi-san), Seta, Gunma-ken, 6.v. –?. vi.1987, T. Tanaka leg. (KUM); 1 male, Minowa, Maebashi-shi, Gunma-ken, 28.v.2006, T. Horiguchi leg. (cTW); 4 unsexed, Kumano-taira, Annaka-shi, Gunma-ken, 10.vii.1958, T. Kikuchi leg. (KUM); 22 unsexed, Kirizumi-onsen, Annaka-shi, Gunma-ken, 25.v.1962, Y. Shibata leg. (KUM); 3 unsexed, Kirizumi-onsen, Annaka-shi, Gunma-ken, 18.vi.1957, Y. Shibata leg. (KUM); 1 unsexed, Kirizumi-onsen, Annaka-shi, Gunma-ken, 24.vi.1955, Y. Shibata leg. (KUM); 1 unsexed, Kirizumi-onsen, Annaka-shi, Gunma-ken, 20.vi.1957, Y. Shibata leg. (KUM); 3 unsexed, Kirizumi-onsen, Annaka-shi, Gunma-ken, 20.vi.1957, H. Yamazaki leg. (cYS); 1 unsexed, Mt. Haruna-san, Takasaki-shi, Gunma-ken, 25.vi.2005, T. Watanabe leg. (cTW); 2 unsexed, Hôroku-tôge Pass, Fujioka-shi, Gunma-ken, 12.vi.2005, T. Horiguchi leg. (cTW); 3 males, 11 unsexed, Shigasakatôge Pass, Kanna-machi, Gunma-ken, 17–19.vi.2008, T. Watanabe leg. (cTW); 3 males, 1 unsexed, Shigasaka-tôge Pass, Kanna-machi, Gunma-ken, 17–19.vi.2008, T. Watanabe leg., from flight interception trap (cTW); 1 unsexed, Usui-tôge Pass, 12.vii.1955, N. Gokan leg. (KUM); 2 unsexed, Hisawa-tôge Pass, Occhi, Ueno-mura, Gunma-ken, 14.v.2002, T. watanabe leg. (cTW); 1 male, 1 female, 1 unsexed, Yakyû-sawa, Narahara, Ueno-mura, Gunma-ken, 24.iv.2008, T. Watanabe leg. (cTW); 1 male, headwaters of Kanna-gawa River, Ueno-mura, Gunma-ken, 23.v.2009, T. Horiguchi leg. (cTW); 1 unsexed, Kawaura-keikoku, Chichibu-shi, Saitama-ken, 2.vi.2001, K. Toyoda leg., from cheese trap (KUM); 1 male, 4 unsexed, Irikawa-keikoku, Ôtaki, Chichibu-shi, Saitama-ken, 29.v.1997, M. Maruyama leg. (KUM); 1 unsexed, Tonbiiwa, Ryôkamikomori, Ogano-machi, Saitama-ken, 16.v.1994, T. Watanabe leg. (cTW); 1 unsexed (heavily damaged), Azeta, Sakura-shi, Chiba-ken, 26.iv.–2.v.1998, M. Sano leg., from carrion trap (KUM); 2 unsexed, Azeta, Sakura-shi, Chiba-ken, 10–17.v.1998, M. Sano leg., from carrion trap (KUM); 1 unsexed, Mt. Kiyosumi-yama, Kamogawa-shi, Chiba-ken, 22.vi.1989, S. Naomi leg.

(KUM); 1 unsexed, Yô-zawa, Akiruno-shi, Tôkyô-to, 11.vi.1959, Y. Shibata leg. (KUM); 1 unsexed, Akiruno-shi, Tôkyô-to, 16.vi.1959, T. Kikuchi leg. (KUM); 1 unsexed, Hikagesawa (Mt. Takao-san), Hachiôji-shi, Tôkyô-to, 16.vii.2006, S. Yamamoto leg. (KUM); 1 unsexed, Mt. Takao-san, Hachiôji-shi, Tôkyô-to, 15.iv.1959, Y. Shibata leg. (KUM); 1 unsexed, Mt. Takao-san, Hachiôji-shi, Tôkyô-to, 4.vi.1976, Niizato leg. (KUM); 1 male, 1 unsexed, Mt. Takao-san, Hachiôji-shi, Tôkyô-to, 17.vi.1976, Niizato leg. (KUM); 1 unsexed, near Ôchi-zawa, Machida-shi, Tôkyô-to, 3.vi.1985, Y. Shibata leg. (KUM); 3 unsexed, Mt. Isseki-san (alt. 1000–1100m), Mt. Ningyô-yama, Nippara, Okutama-machi, Tôkyô-to, 9.vii.2001, H. Kamezawa leg., from tree hollow in dry condition (cHK); 1 male, 4 unsexed, Mt. Isseki-san (alt. 800–1100m), Mt. Ningyô-yama, Nippara, Okutama-machi, Tôkyô-to, 2.vii.2013, H. Kamezawa leg., from tree hollow in dry condition (cHK); 1 unsexed, Nippara, Okutama-chô, Tôkyôto, 31.v.1964, K. Kurosa leg. (KUM); 1 unsexed, Nippara, Okutama-chô, Tôkyô-to, 25.vii.1966, Y. Kurosawa leg. ( NSMT); 1 unsexed, Serizawa, Zama-shi, Kanagawa-ken, 8.vii.1989, M. Nishikawa leg. (KUM); 3 unsexed, Zama-shi, Kanagawa-ken, 22.vii.1972, Y. Hirano leg. (KUM); 1 unsexed, Tsukui, Shônan, Kanagawa-ken, 18.viii.1949, R. Kiryû leg. (SCM); 1 female, Nakatsu, Aikô, Aikawa-machi, Kanagawa-ken, 7.vii.1959, R. Kiryû leg. (SCM); 2 unsexed, Mt. Kyôga-take, Aikawa-machi, Kanagawa-ken, 15–22.viii.2003, T. Watanabe leg. (cTW); 1 unsexed, Fudakake, Kiyokawa-mura, Kanagawa-ken, 16.vi.1984, Y. Hirano leg. (KUM); 1 unsexed, Mt. Jinbasan, Midori-ku, Sagamihara-shi, Kanagawa-ken, 17–23.vii.2004, T. Watanabe leg. (cTW); 1 male, 2 unsexed, Wada-tôge Pass, Midori-ku, Sagamihara-shi, Kanagawa-ken, 9–17.vii.2004, T. Watanabe leg. (cTW); 1 male, Mt. Ô-yama, Kanagawa-ken, 13.vii.1952, R. Kiryû leg. (SCM); 1 unsexed, Mt. Ô-yama, Kanagawa-ken, 5.v.1960, Y. Shibata leg. (KUM); 1 male, Mt. Ô-yama, Kanagawa-ken, 15.vii.1969, R. Kiryû leg. (SCM); 1 unsexed, Mt. Ôyama, Kanagawa-ken, 29.vi.1974, Y. Shibata leg. (KUM); 3 males, Mt. Ô-yama, Kanagawa-ken, 3–11.vii.2006, T. Watanabe leg. (cTW); 1 unsexed, Nagao-one mountain ridge, Kanagawa-ken, 1.vii.1984, T. Watanabe leg. (cTW); 1 male, 6 unsexed, Futamata, Hadano-shi, Kanagawa-ken, 30.iv.1996, T. Watanabe leg. (cTW); 6 unsexed, near Bodai, Hadano-shi, Kanagawa-ken, 8.vi.1969, Y. Shibata leg. (KUM); 1 unsexed, Ôkura, Hadano-shi, Kanagawaken, 26.v.2000, T. Watanabe leg. (cTW); 1 male, Mt. Fudô-yama, Nakai-machi, Kanagawa-ken, 3–12.vi.2006, T. Watanabe leg. (cTW); 1 unsexed, Yadorogi, matsuda-machi, Kanagawa-ken, 13.v.1984, T. Watanabe leg. (cTW); 1 male, 2 unsexed, Yadorigi, Matsuda-machi, Kanagawa-ken, 14.iv.1986, M. Tao leg. (KUM); 1 unsexed, Yadorigi, Matsuda-machi, Kanagawa-ken, 14.vi.1986, M. Tao leg. (KUM); 1 unsexed, Nishitanzawa, Kanagawa-ken, 26.v.1984, Y. Hirano leg. (KUM); 6 unsexed, Mt. Kanyûdô-yama, Yamakita-machi, Kanagawa-ken, 24.vi- 1.viii.2001, Y. Syu leg. (cTW); 3 unsexed, Shiraishi-zawa, Yamakita-machi, Kanagawa-ken, 13.v.1984, T. Watanabe leg. (cTW); 1 unsexed, Shiraishi-zawa, Yamakita-machi, Kanagawa-ken, 10.viii.2001, T. Watanabe leg. (cTW); 3 unsexed, Yôke-zawa, Yamakita-machi, Kanagawa-ken, 10.vii.1993, T. Watanabe leg. (cTW); 7 unsexed (preserved in 99.5% EtOH for DNA sequencing), Yôki-zawa deai, Yamakita-machi, Kanagawa-ken, 10.v.2014, Y. Shibata leg. (KUM); 1 unsexed, Hôki-zawa, Yamakita-machi, Kanagawa-ken, 13.v.1984, T. Watanabe leg. (cTW); 2 males, Ôtaki-zawa, Yamakita-machi, Kanagawa-ken, 9–17.vi.2005, T. Watanabe leg. (cTW); 2 unsexed, Jizôdaira, Yamakita-machi, Kanagawa-ken, 31.vi.2001, T. Watanabe leg. (cTW); 2 males, 2 unsexed, Jizô-daira, Yamakita-machi, Kanagawa-ken, 31.v–6.vi.2006, T. Watanabe leg. (cTW); 1 unsexed, Iden-zawa, Yamakitamachi, Kanagawa-ken, 31.v–6.vi.2006, T. Watanabe leg. (cTW); 1 male, Ômata-zawa, Yamakita-machi, Kanagawa-ken, 31.v.2006, T. Watanabe leg. (cTW); 1 unsexed, Yuzuku, Yamakita-machi, Kanagawa-ken, 1.v.1985, T. Watanabe leg. (cTW); 2 unsexed, Mt. Teppôgino-atama, Yamakita-machi, Kanagawa-ken, 29.vi.2001, T. Watanabe leg. (cTW); 1 unsexed, Mt. Teppôgino-atama, Yamakita-machi, Kanagawa-ken, 20.vi.2007, T. Watanabe leg. (cTW); 1 male, Mt. Teppôgino-atama, Yamakita-machi, Kanagawa-ken, 27.vi.2007, T. Watanabe leg. (cTW); 1 male, Mt. Teppôgino-atama, Yamakita-machi, Kanagawa-ken, 5–12.vii.2007, T. Watanabe leg., from flight interception trap (cTW); 2 unsexed, Mt. Teppôgino-atama, Yamakita-machi, Kanagawa-ken, 5–12.vii.2007, T. Watanabe leg. (cTW); 1 unsexed, Mikuni-tôge Pass, Yamakita-machi, Kanagawa-ken, 27.vi.2007, T. Watanabe leg. (cTW); 1 unsexed, Mt. Mikuni-yama, Yamakita-machi, Kanagawa-ken, 3.viii.2003, T. Watanabe leg. (cTW); 1 unsexed, Mt. Mikuni-yama, Yamakita-machi, Kanagawa-ken, 3–7.viii.2003, T. Watanabe leg. (cTW); 1 unsexed, Mt. Tanzawa-yama, Yamakita-machi, Kanagawa-ken, 3.vi.1962, S. Shinonaga leg. (cYS); 2 unsexed, Chôjagoya, Kanagawa-ken, 25.v.1986, T. Watanabe leg., under fallen leaves on streamlet (cTW); 1 unsexed, Chôjagoya, Kanagawa-ken, 1.viii.2000, T. Watanabe leg. (cTW); 7 unsexed, Chisujino-taki Fall, Hakone-machi, Kanagawaken, 1.vi.1969, Y. Shibata leg. (KUM); 1 unsexed, Tônosawa, Hakone-machi, Kanagawa-ken, 24.x.1971, Y. Hirano leg. (KUM); 1 male, 9 unsexed, Kamiyu, Hakone-machi, Kanagawa-ken, 8.viii.1984, K. Nemoto leg. (KUM); 4 unsexed, Sengokuhara, Hakone-machi, Kanagawa-ken, 29.vi.1969, Y. Shibata leg. (KUM); 1 male, Ôwakudani, Hakone-machi, Kanagawa-ken, 17–27.viii.2007, T. Watanabe leg. (cTW); 1 male, 1 female, Mt. Kami-yama, Hakone-machi, Kanagawa-ken, 1.vi.1974, Y. Shibata leg. (cYS); 6 unsexed, Mt. Kami-yama, Hakone-machi, Kanagawa-ken, 15.vii.1985, Y. Shibata leg. (KUM); 1 female, Mt. Kami-yama, Hakone-machi, Kanagawa-ken, 31.vii.1996, K. Kawada leg. (KUM); 2 unsexed, Kojiri, Hakone-machi, Kanagawa-ken, 11.vi.1972, Y. Hirano leg. (KUM); 3 males, 1 unsexed, Mt. Komaga-take, Hakone-machi, Kanagawa-ken, 24.vii–1.viii.2008, T. Watanabe leg. (cTW); 1 unsexed, Sukumogawa, Hakone-machi, Kanagawa-ken, 8.vi.1991, T. Fujimori leg. (EUM); 1 unsexed, Hakone-machi, Kanagawa-ken, 22.viii.1982, M. Tao leg. (KUM); 1 male, Asami, Yuzawa-machi, Niigata-ken, 28.v.1995, K. Haga leg., from larch forest (KUM); 1 female, Mt. Sanri-yama, Fukui-ken, 26.vii.1974, H. Sasaji leg. (KUM); 1 male, 1 unsexed, Kamihikawa-tôge Pass, Kôshû-shi, Yamanashi-ken, 15–18.vii.1982, S. Naomi leg. (KUM); 6 unsexed, Kamihikawa-tôge Pass, Kôshû-shi, Yamanashi-ken, 14.vi.1986, T. Watanabe leg., under rubbish (cTW); 1 male, 3 unsexed, Kamihagiwara, Kôshû-shi, Yamanashi-ken, 9–15.viii.2006, T. Watanabe leg. (cTW); 1 unsexed, middle of Hikawa-rindô forestry road, Daibosatsu-tôge Pass, Yamanashi-ken, 28.v.1980, K. Haga leg. (KUM); 1 male, Mts. Daibosatsu-rei, Yamanashi-ken, 12.iv.1982, S. Ômomo leg. (KUM); 3 males, 4 unsexed, Sasago, Ôtsuki-shi, Yamanashi-ken, 20–27.2001, Y. Syu leg. (cTW); 1 male, Murokubo-rindô forestry road, Dôshi-mura, Yamanashi-ken, 13–21.viii.2009, T. Watanabe leg., from flight interception trap (cTW); 1 unsexed, Shiroi-daira, Dôshi-mura, Yamanashi-ken, 17.vii.2001, T. Watanabe leg. (cTW); 6 males, 7 females, 4 unsexed, Masutomi-onsen, Hokuto-shi, Yamanashi-ken, 9–10.vii.1998, M. Maruyama leg., alt. 1200m, from carrion trap (KUM); 1 unsexed, Sawara-ike Pond, Nirasaki-shi, Yamanashi-ken, 8–9.vii.1998, M. Maruyama leg., alt. 1200m (KUM); 1 male, Gozaishi-onsen, Nirasaki-shi, Yamanashi-ken, 10.vi.1989, K. Hosoda leg. (cTI); 1 unsexed, Gozaishi-onsen, Nirasaki-shi, Yamanashi-ken, 4.vi.1990, K. Hosoda leg. (cTI); 1 unsexed, Aokigahara (Mt. Fuji-san), Yamanashi-ken, 20.vii.1982, S. Naomi leg. (KUM); 2 males, 2 unsexed, Aokigahara, Fujikawaguchiko-machi, Yamanashi-ken, 11–17.v.2012, T. Watanabe leg., from flight interception trap (cTW); 1 male, Aokigahara, Fujikawaguchiko-machi, Yamanashi-ken, 17–24.v.2012, T. Watanabe leg., from flight interception trap (cTW); 3 males, Aokigahara, Fujikawaguchiko-machi, Yamanashi-ken, 17.v–7.vi.2012, T. Watanabe leg., from chicken baited trap (cTW); 1 male, 2 unsexed, Aokigahara, Fujikawaguchiko-machi, Yamanashi-ken, 24.v.2012, T. Watanabe leg. (cTW); 287 unsexed, Shibire-ko Lake, Ichikawamisato-chô, Yamanashi-ken, 25–26.vi.2011, S. Morita leg. (KUM); 2 males, Kazumizu-rindô forestry road, Narusawa-mura, Yamanashi-ken, 22–29.vi.2011, T. Watanabe leg., alt. 1600m, from flight interception trap (cTW); 1 male, Kazumizu-rindô forestry road, Narusawa-mura, Yamanashi-ken, 29.vi–6.vii.2011, T. Watanabe leg., alt. 1600m, from flight interception trap (cTW); 1 male, Kazumizu-rindô forestry road, Narusawa-mura, Yamanashi-ken, 16– 25.vii.2012, T. Watanabe leg., alt. 1800m, from flight interception trap (cTW); 1 unsexed, Kazumizu-rindô forestry road, Narusawa-mura, Yamanashi-ken, 25.vii–1.viii.2012, T. Watanabe leg., alt. 1800m, from flight interception trap (cTW); 1 female, Kazumizu-rindô forestry road, Narusawa-mura, Yamanashi-ken, 1.viii.2012, T. Watanabe leg., alt. 1800m (cTW); 1 female, 1 unsexed, Kazumizu-rindô forestry road (1800m alt.), Narusawa-mura, Yamanashi-ken, 1–8.viii.2012, T. Watanabe leg., from flight interception trap (cTW); 1 unsexed, Kazumizu-rindô forestry road, Narusawa-mura, Yamanashi-ken, 10–17.viii.2011, T. Watanabe leg., alt. 1800m (cTW); 1 unsexed, Mt. Fuji-san, Narusawa-mura, Yamanashi-ken, 2.vi.1986, T. Inagawa leg. (KUM); 1 unsexed, Mt. Fuji-san, Narusawa-mura, Yamanashi-ken, 30.vii.1986, S. Komata leg. (KUM); 1 unsexed, Mt. Fuji-san (south side), 25– 28.vii.1971, K. Tanaka & K. Suga leg., alt. 1000m, from baited trap (KUM); 2 unsexed, Mt. Fuji-san (south side), 28–31.vii.1971, K. Tanaka & K. Suga leg., alt. 1000m, from baited trap (KUM); 1 unsexed, Mt. Fuji-san (south side), 7–13.viii.1971, K. Tanaka leg., alt. 1000m, from baited trap (KUM); 2 unsexed, Mt. Fuji-san (south side), 25–28.vii.1971, K. Tanaka & K. Suga leg., alt. 1400m, from baited trap (KUM); 1 unsexed, Mt. Fuji-san (south side), 27.vi.1971, K. Tanaka leg. (KUM); 1 unsexed, Wadanono-mori, Hakuba-mura, Nagano-ken, 31.vii.2003, T. Watanabe leg. (cTW); 1 unsexed, Karuizawa, Nagano-ken, 23.vii.1963, K. Kurosa leg. (KUM); 1 unsexed, Jigokudani, Yamanouchi-machi, Nagano-ken, 8.vii.1982, S. Naomi leg. (KUM); 1 unsexed, Togakushi, Nagano-shi, Nagano-ken, 29.vii.1965, K. Tanizawa leg. (cYH); 1 unsexed, Kakuma-keikoku, Ueda-shi, Nagano-ken, 6.vii.2002, T. Watanabe leg. (cTW); 1 unsexed, Shimashima-dani, Nagano-ken, 7.vii.1969, S. Kinoshita leg. (EUM); 1 unsexed, Kemi, Azumino-shi, Nagano-ken, 3–4.iii.2002, A. Sugimura leg. (KUM); 1 unsexed, Tobira, Iriyamabe, Matsumoto-shi, Nagano-ken, 3–4.viii.2002, A. Sugimura leg., alt. 1000m (KUM); 1 male, 4 unsexed, Fujiidani, Matsumoto-shi, Nagano-ken, 4.v.1987, N. Itô leg. (cYH); 1 unsexed, Fujiidani, Matsumoto-shi, Naganoken, 4.v.1987, N. Itô leg. (KUM); 1 male, 9 unsexed, Pension-village, Hara-mura, Nagano-ken, 9.vii.1999, A. Shiho leg. (KUM); 1 male, 1 unsexed, Mt. Nyûkasa-yama, Fujimi-machi, Nagano-ken, 17.vi.2007, S. Yamamoto leg. (KUM); 1 unsexed, Mt. Kyôga-take, Nagano-ken, 7.viii.1997, Y. Hayashi leg. (cYH); 1 female, Okeya, Ôshika-mura, Nagano-ken, 26.iv.2008, H. Kamezawa leg. (cHK); 1 unsexed, Mt. Shirasawa-yama, Ôshika-mura, Nagano-ken, 29.v.1999, H. Kamezawa leg., alt. 700–800m (cHK); 1 male, 1 unsexed, Koshibugawa, Ôshika-mura, Nagano-ken, 21–23.viii.2007, T. Watanabe leg., alt. 1000m (cTW); 1 male, 1 unsexed, Kisofukushima, Naganoken, 31.v.1987, Y. Hayashi (cYH); 1 unsexed, Mikuni-tôge Pass, Shizuoka-ken, 28.vi.1992, Y. Notsu leg. (EUM); 2 males, Subashiri (Mt. Fuji-san, 1 gôme), Oyama-chô, Shizuoka-ken, 20–26.v.2011, T. Watanabe leg., alt. 1400m, from flight interception trap (cTW); 2 males, Subashiri-guchi (Mt. Fuji-san), Shizuoka-ken, 9–15.vi.2011, T. Watanabe leg., alt. 1400m, from flight interception trap (cTW); 2 males, Midoro, Gotenba-shi, Shizuoka-ken, 15– 20.v.2011, T. Watanabe leg. (cTW); 1 male, Mizugatsuka, Suyama, Susono-shi, Shizuoka-ken, 31.v–5.vi.2013, T. Watanabe leg., from baited trap (silkworm pupa powder) (cTW); 1 unsexed, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 13–18.v.2010, T. Watanabe leg. (cTW); 2 males, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 10– 17.vi.2010, T. Watanabe leg., from flight interception trap (cTW); 1 male, 1 unsexed, Nishiusuzuka, Fujinomiyashi, Shizuoka-ken, 8–16.vii.2010, T. Watanabe leg., from flight interception trap (cTW); 2 unsexed, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 16–22.vii.2010, T. Watanabe leg., from flight interception trap (cTW); 1 unsexed, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 17–24.viii.2010, T. Watanabe leg., from flight interception trap (cTW); 1 male, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 22.viii.2010, T. Watanabe leg., (cTW); 1 unsexed, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 22–28.viii.2012, T. Watanabe leg. (cTW); 1 male, 2 unsexed, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 28.viii–5.ix.2012, T. Watanabe leg., from baited trap (silkworm pupa powder) (cTW); 1 female, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 5.ix.2012, T. Watanabe leg. (cTW); 1 female, Nishiusuzuka, Fujinomiya-shi, Shizuoka-ken, 5–11.vi.2013, T. Watanabe leg., from carrion trap (cTW); 1 male, Mt. Takahachi-yama, Fujinomiya-shi, Shizuoka-ken, 16–22.vii.2011, T. Watanabe leg., from fermented banana trap (cTW); 2 males, Ôbuchi, Fuji-shi, Shizuoka-ken, 10–17.vi.2010, T. Watanabe leg., alt. 950m, from flight interception trap (cTW); 2 unsexed, Ôbuchi, Fuji-shi, Shizuoka-ken, 10–17.vi.2010, T. Watanabe leg., alt. 950m, from baited trap (cTW); 1 unsexed, Ôbuchi, Fuji-shi, Shizuoka-ken, 8–16.vii.2010, T. Watanabe leg., alt. 950m, from flight interception trap (cTW); 1 male, 1 female, 1 unsexed, Ôbuchi, Fuji-shi, Shizuoka-ken, 17– 24.viii.2010, T. Watanabe leg., alt. 950m, from flight interception trap (cTW); 1 male, 1 female, Ôbuchi, Fuji-shi, Shizuoka-ken, 11–18.vi.2013, T. Watanabe leg., alt. 950m, from baited trap (silkworm pupa powder) (cTW); 1 unsexed, Itô-shi, Shizuoka-ken, 4.iv.1969, Y. Kusui leg. (cYH); 2 unsexed, Hacchô-ike Pond, Izu-shi, Shizuokaken, 3.viii.1985, T. Watanabe leg. (cTW); 2 unsexed, Hacchô-ike Pond, Izu-shi, Shizuoka-ken, 24.v.2002, T. Watanabe leg. (cTW); 1 unsexed, Mts. Amagi-san, Shizuoka-ken, 19.vii.1955, Y. Shibata leg. (KUM); 2 unsexed, Mts. Amagi-san, Shizuoka-ken, 21.viii.1955, Y. Shibata leg. (KUM); 1 unsexed, Taguchi, Shitara-chô, Aichi-ken, 7.ix.2001, H. Kamezawa leg., alt. 400m (cHK); 1 unsexed, Mt. Kurai-yama, Takayama-shi, Gifu-ken, 21.vii.2003, Y. Takai leg. (KUM); 1 unsexed, Midoriyokokura-rindô forestry road, Ibigawa-chô, Gifu-ken, 15–16.vi.2000, A. Sugimura leg. (KUM); 1 unsexed, Torigoe-tôge Pass, Nagahama-shi, Shiga-ken, 5–12.viii.2001, A. Sugimura leg., alt. 900m (KUM); 1 unsexed, Mii-dera Temple, Ôtsu-shi, Shiga-ken, 14.vi.1989, R. Kuranishi leg. (KUM); 1 unsexed, Hanase-tôge Pass, Sakyô-ku, Kyôto-shi, Kyôto-fu, 2.vi.1958, Y. Hayashi leg. (cYH); 1 unsexed, Kibune, Sakyô-ku, Kyôto-shi, Kyôto-fu, 7.viii.1983, K. Masaki leg. (cTI); 1 unsexed, Kumogahata, Kita-ku, Kyôto-shi, Kyôto-fu, 28.vi.1987, T. Watanabe leg. (cTW); 1 unsexed, Kumogahata, Kita-ku, Kyôto-shi, Kyôto-fu, 27.viii.1983, K. Masaki leg. (cTI); 3 unsexed, Mt. Chôrô-san, Kyôtanba-chô, Kyôto-fu, 22.v.1994, Y. Hayashi leg. (cYH); 1 unsexed, Mt. Oaihi, Kyôto-fu, 26.v.1962, M. Yasui leg. (cYH); 1 unsexed, Kasuga, Nara-shi, Nara-ken, 22.vii.1970, Y. Hayashi leg. (cYH); 1 unsexed, Mt. Misen, Tenkawa-mura, Nara-ken, 3.viii.1985, N. Itô leg. (KUM); 1 unsexed, Tenkawa-mura, Nara-ken, 9.vii.1990, A. Saito leg. (KUM); 1 unsexed, Tônomine, Sakurai-shi, Nara-ken, 13.vii.1985, T. Ito leg. (KUM); 1 unsexed, Mt. Ôdaigahara-yama, Nara-ken, 5.viii.1959, H. Konishi leg. (cYH); 1 unsexed, Mt. Ôdaigahara-yama, Nara-ken, 7.viii.1988, N. Itô leg. (KUM); 4 unsexed, Mt. Ôdaigaharayama, 2.viii.1967, H. Nomura leg. (cYH); 1 female, 1 unsexed, Hirai, Kozagawa-chô, Wakayama-ken, 16.vii.1999, M. Maruyama leg., from carrion trap (KUM); 2 unsexed, Mt. Gomadan-zan, Tanabe-shi, Wakayama-ken, 4.viii.1993, I. Matoba leg. (KUM); 1 unsexed, Mt. Kôya-san, Kôya-chô, Wakayama-ken, 10.vii.1995, I. Matoba leg. (KUM); 2 unsexed, Akazai-keikoku, Shisô-shi, Hyôgo-ken, 22.vii.1979, T. Ito leg., from berlase apparatus (cYH); 1 unsexed, Akazai-keikoku, Shisô-shi, Hyôgo-ken, 15.vii.1979, Y. Hayashi leg. (cYH); [SHIKOKU]: 58 unsexed, Mt. Rokuro-yama, Sanagôchi-son, Tokushima-ken, 5.v–1.vii.2000, M. Yoshida leg., alt. 830m, from fowl trap (KUM); 1 male, 14 unsexed, Mt. Takamaru, Kamikatsu-chô, Tokushima-ken, 28.v.–18.vi.2005, M. Yoshida leg., alt. 1260m, from fowl trap (KUM); 2 females, 9 unsexed, Mt. Kumosa-yama, Naka-chô, Tokushima-ken, 13– 15.vii.2001, A. Sugimura leg. (KUM); 9 unsexed, Mt. Kumosa-yama, Tokushima-ken, 6–7.vi.1987, K. Nemoto leg. (KUM); 14 unsexed, Mt. Takashiro-yama, Kisawa, Naka-chô, Tokushima-ken, 29.v–19.vii.2004, M. Yoshida leg., alt. 1430m, from fowl trap (KUM); 2 unsexed, Mt. Kôtsu-san, Yamakawa-chô, Yoshinogawa-shi, Tokushimaken, 29.vi.2011, M. Nishikawa leg., alt. 1050m, from flight interception trap (EUM); 1 unsexed, Mt. Kôtsu-san, Yamakawa-chô, Yoshinogawa-shi, Tokushima-ken, 5.vi.1999, K. Aita leg. (EUM); 3 unsexed, Mt. Kôtsu-san, Yoshinogawa-shi, Tokushima-ken, 12.vii.2003, H. Fujimoto leg., from flight interception trap (KUM); 25 unsexed, Ôhira, Misato, Yoshinogawa-shi, Tokushima-ken, 25.v–7.vi.2003, alt. 820m, M. Yoshida leg., from fowl trap (KUM); 12 unsexed, Mt. Maruzasa-yama, Ichiu-son, Tokushima-ken, 11.vi–6.viii.2000, M. Yoshida leg., alt. 1410m, from fowl trap (KUM); 4 unsexed, Mt. Ichinomori, Tokushima-ken, 3–10.viii.2002, M. Yoshida leg., from fowl trap (KUM); 1 unsexed, Minokoshi (Mt. Turugi-san), Tokushima-ken, 18.vi.1998, M. Yoshida leg., alt. 1350m (KUM); 1 unsexed, Minokoshi (Mt. Tsurugi-san), Tokushima-ken, 29.vii.1960, T. Shibata leg. (cYH); 1 male, 1 unsexed, Minokoshi (Mt. Tsurugi-san), Tokushima-ken, 10.vi.2000, M. Yoshida leg., from baited trap (two weeks) (KUM); 1 female, 32 unsexed, Minokoshi (Mt. Tsurugi-san), Higashiiya, Miyoshi-shi, Tokushima-ken, 5.vi–3.vii.2006, M. Yoshida leg., alt. 1350m, from fowl trap (KUM); 28 unsexed, Minokoshi (Mt. Tsurugi-san), Higashiiya, Miyoshi-shi, Tokushima-ken, 11.vi–17.vii.2006, M. Yoshida leg., alt. 1350m, from fowl trap (KUM); 1 unsexed, Minokoshi (Mt. Tsurugi-san), Higashiiya, Miyoshi-shi, Tokushima-ken, 5.vi–3.vii.2006, M. Yoshida leg., alt. 1700m, from fowl trap (KUM); 1 male, 29 unsexed, Ôtsurugi (Mt. Tsurugi-san), Higashiiya, Miyoshi-shi, Tokushima-ken, 5.vi–3.vii.2006, M. Yoshida leg., alt. 1800m, from fowl trap (KUM); 1 male, 1 female, 10 unsexed, Otome-dani, Nagoro, Higashiiya, Miyoshi-shi, Tokushima-ken, 17.vii–30.x.2006, M. Yoshida leg., alt. 1260m, from fowl trap (KUM); 2 unsexed, Mt. Tsurugi-san, Tokushima-ken, 25.vi.1966, Y. Imai leg. (cYH); 1 unsexed, Mt. Tsurugi-san, 11.vii.1976, T. Suzuki leg. (EUM); 1 unsexed, Mt. Daisen-zan, Mannô-chô, Kagawaken, 21.vi.2003, H. Fujimoto leg., from flight interception trap (KUM); 1 unsexed, Mt. Daisen-zan, Mannô-chô, Kagawa-ken, 3.vii.2008, H. Fujimoto leg. (KUM); 1 unsexed, Mt. Daisen-zan, Mannô-chô, Kagawa-ken, 21.vi.2011, H. Fujimoto leg. (KUM); 1 unsexed, Umatate, Shikokuchûô-shi, Ehime-ken, 4.viii.2003, J. Ogawa leg. (EUM); 14 unsexed, Mado-tôge Pass, Saijô-shi, Ehime-ken, 9.vi.1984, T. Nagata leg. (EUM); 1 unsexed, Mt. Takanawa-san, Matsuyama-shi, Ehime-ken, 19.v.1956, M. Miyake leg. (EUM); 1 unsexed, Mt. Takanawa-san, Matsuyama-shi, Ehime-ken, 14.vii.2005, Y. Satô leg. (EUM); 1 unsexed, west ravine of Mt. Shiratsue, Matsuyamashi, Ehime-ken, 11.v.1969, M. Sakai leg. (EUM); 13 unsexed, west ravine of Mt. Shiratsue, Matsuyama-shi, Ehime-ken, 14.vi.1984, T. Nagata leg. (EUM); 1 unsexed, Fukumigawa-chô, Matsuyama-shi, Ehime-ken, 1– 4.ix.1986, M. Hiraoka leg. (EUM); 12 unsexed, Tateiwakomenono, Matsuyama-shi, Ehime-ken, 10.v.1977, N. Yashiro leg., from chicken trap (EUM); 1 unsexed, Tateiwakomenono, Matsuyama-shi, Ehime-ken, 23.v.1978, I. Amano leg. (EUM); 1 unsexed, Tateiwakomenono, Matsuyama-shi, Ehime-ken, 11.vi.1993, L. Li leg. (EUM); 2 unsexed, Tateiwakomenono, Matsuyama-shi, Ehime-ken, 20.vi.1995, S. Yoshimichi leg. (EUM); 2 males, 8 unsexed, Tateiwakomenono, Matsuyama-shi, Ehime-ken, 1–2.vi.1996, K. Yamanaka leg., from baited trap (EUM); 2 unsexed, Tateiwakomenono, Matsuyama-shi, Ehime-ken, 15.v.2005, Y. Satô leg. (EUM); 1 male, 1 female, Tateiwakomenono, Matsuyama-shi, Ehime-ken, 29.v.2005, Y. Katayama leg. (EUM); 3 unsexed, west ravine of Mt. Shiratsue, Ehime-ken, 20.vi.1986, T. Nagata leg., alt. 780m (scrub and Japanese cedar, st. 3) (EUM); 1 male, Wakimizu, Kamibayashi, Tôon-shi, Ehime-ken, 12.vi.2006, Y. Satô leg. (EUM); 1 unsexed, Wakimizu, Kamibayshi, Tôon-shi, Ehime-ken, 10.vii.2006, Y. Satô leg. (EUM); 1 unsexed, Wakimizu, Kamibayashi, Tôonshi, Ehime-ken, 22.vii.2006, Y. Satô leg. (EUM); 1 unsexed, Mt. Saraga-mine, Tôon-shi, Ehime-ken, 21.v.2002, Y. Kikuhara leg. (EUM); 1 unsexed, Omogo-kei (Mt. Ishizuchi-san), Tsuchigoya, Kumakôgen-chô, 3–7.viii.1986, M. Ebihara leg., from chicken trap (st.55) (EUM); 2 unsexed, Omogo-kei (Mt. Ishizuchi-san), Tsuchigoya, Kumakôgen-chô, 3–7.viii.1986, M. Ebihara leg., from chicken trap (st.58) (EUM); 13 unsexed, Masakinomori, Tengu-kôgen Heights, Kumakôgen-chô, Ehime-ken, 13–14.vii.1999, H. Kamezawa leg. (cHK); 1 unsexed, Mt. Ishizuchi-san, Ehime-ken, 20.vii.1960, T. Kikuchi leg. (KUM); 2 unsexed, Furuiwaya, Kumakôgen-chô, Ehimeken, 25.v.1986, S. Hisamatsu leg., alt. 575m, from chicken trap (EUM); 1 unsexed, Mt. Saraga-mine, Kumakôgenchô, Ehime-ken, 17.vii.2005, Y. Satô leg. (EUM); 2 males, 53 unsexed, Mt. Saraga-mine, Kumakôgen-chô, Ehimeken, 18.vi.2006, Y. Satô leg. (EUM); 2 unsexed, Mt. Saraga-mine, Ehime-ken, 26.vi.1959, M. Satô leg. (EUM); 2 unsexed, Odamiyama, Uchiko-chô, Ehime-ken, 11–13.vi.1981, S. Naomi leg. (KUM); 4 unsexed, Odamiyama, Uchiko-chô, Ehime-ken, 19.vii.1993, M. Sakai leg. alt. 800m (EUM); 1 unsexed, Odamiyama, Uchiko-chô, Ehime-ken, 14.v.1994, L. Li leg. (EUM); 1 unsexed, Odamiyama, Uchiko-chô, Ehime-ken, 1–2.vii.1995, M. Zhao leg. (EUM); 1 unsexed, Befu, Monobe, Kôchi-ken, 4.vi.1994, I. Yamashita leg. (EUM); 2 unsexed, Befu, Monobe, Kôchi-ken, 12.vi.1994, I. Yamashita leg. (EUM); 2 unsexed, Befu, Monobe, Kôchi-ken, 28.v.1994, I. Yamashita leg. (EUM); 2 unsexed, Takano, Yusuhara-chô, Kôchi-ken, 24–25.v.1997, K. Kasahara leg., from baited trap (EUM); 1 unsexed, Kuroson, Shimanto-shi, Kôchi-ken, 20.vii.1957, M. Miyake leg. (EUM); 5 unsexed, Komi, Kumakôgen-chô, 15–16.vii.1994, K. Okada & M. Mino leg., from baited trap (EUM); [KYÛSHÛ]: 1 male, 1 unsexed, Umami, Kama-shi, Fukuoka-ken, R. Noda leg., from flight interception trap (KUM); 1 unsexed, Hacchôtôge Pass, Kama-shi, Fukuoka-ken, 17–19.vii.2003, R. Noda leg., from flight interception trap (KUM); 1 unsexed, Mt. Hiko-san, Soeda-machi, Fukuoka-ken, 12.v.1969, M.-T. Chûjô leg. (KUM); 1 unsexed, Fukuoka-ken, 28.vii.1931, Yasumatsu & Cho leg. (KUM); 1 unsexed, Uchiyama, Tsushima-shi (Tsu-shima Is.), Nagasaki-ken, 6.v.2009, S. Yamamoto leg. (KUM); 1 unsexed, Rougaudo, Minamiaso-mura, Kumamoto-ken, 10.vii.2001, I. Ôtsuka leg. (KUM); 4 unsexed, Mt. Shiratori-yama, Momigi, Izumi-mura, Yatsushiro-shi, Kumamoto-ken, 6.vii.2003, R. Noda leg. (KUM); 9 unsexed, Mt. Shiratori-yama, Momigi, Izumi-mura, Yatsushiro-shi, Kumamotoken, 6.vii.2003, R. Noda leg., from flight interception trap (KUM); 1 male, Momigi, Izumi-machi, Yatsushiro-shi, Kumamoto-ken, 30–31.viii.2008, R. Noda leg., from flight interception trap (KUM); 2 unsexed, Hagi, Yatsushiroshi, Kumamoto-ken, 24.vi.1982, S. Imasaka leg. (KUM); 1 unsexed, Mt. Ichifusa-yama, Mizukami-mura, Kumamoto-ken, 24.vii.1969, H. Nomura leg. (cYH); 1 unsexed, Mt. Kuro-dake, Yufu-shi, Ôita-ken, 29.vii.1979, S. Imasaka leg. (KUM); 1 unsexed, Ko-ike Lake, Oike-chô, Miyakonojô-shi, Miyazaki-ken, 5.v.1981, K. Andô leg. (cYH).

Other material. CHINA: 1 male, “Manchuria / Chandaochedzi // Chicago NHMus / M. Bernhauer / Collection” (median lobe was pulled off by somebody, glued on PC with the body) ( FMNH).

Note: There has been no record of A. coreana from China for a long time ( Smetana, 2004b; Luo & Zhao, 2012), but it was recently added to the Chinese fauna (see Schülke & Smetana, 2015).

Diagnosis. Aleochara coreana is externally very similar to the other Japanese congeners, especially A. curtula , A. postica , and A. parens , but it can be discriminated from them by the combined following characteristics: body size medium to relatively large ( A. parens often small-sized); dorsal surface rather finely punctured (not A. postica ); elytra uniformly yellowish to reddish brown without lateral dark maculations (not A. curtula and some of A. postica ); median lobe of male aedeagus bulbous (not A. postica ), no hook-like shaped at apex (not A. curtula ), with complex sclerites reaching to almost basal margin of median lobe (not A. parens ), and by; non-deformed female spermatheca (not A. postica ), with only small apical invagination of spermathecal head and coarsely striate inner walls (not A. curtula ). Restricted to only mainland and adjacent large islands in Japan.

Redescription. Measurements (in mm, n = 20): BL = 6.40 (4.96–8.66); HL = 0.80 (0.72–0.93); HW = 0.92 (0.79–1.16); PL = 1.18 (1.03–1.34); PW = 1.61 (1.42–1.82); EL = 0.94 (0.81–1.09); EW = 1.87 (1.27–2.21).

Body ( Figs 1 , 10 ): fusiform, robust, medium to large in size; dorsal surface moderately glossy and pubescent, without any types of micro-reticulation, rather finely punctured. Color ( Figs 1 , 10 ): usually uniformly black to blackish brown, but elytra variable in color, generally yellowish to reddish brown uniformly including lateral margins, without any types of dark maculations; antennae with antennomeres I–III pale to dark brown, antennomere IV much darker, antennomeres V–XI slightly darker than those of I–III but covered with minute whitish setae densely; mouth parts and legs pale yellowish brown to dark reddish brown; pubescence yellowish brown.

Head ( Fig. 19 ): semicircular, as long as width (HW/HL = 1.16, n = 20), widest at posterior part of eyes; setae on vertex rather dense but inconspicuous, directed anteriomedially; punctation medium in size, scattered rather densely; vertex longitudinally lack of punctation at middle, but small to medium in size. Eyes: medium in length, occupying approximately half length of head, slightly protruding laterally.

Antennae ( Fig. 37 ): short, moderately shorter than head and pronotum combined; thick, setose, with antennomere IV moderately transverse, but antennomeres V to X strongly transverse in approximately same width each; antennomere XI symmetrical, obtusely rounded at apex; approximate relative length of antennomeres from basal to apex: 20 (including base): 9: 11: 4: 4.5: 5: 5: 5: 5: 5: 14.

Pronotum ( Fig. 1 ): transversely oval (PW/PL = 1.36, n = 20), strongly narrowing anterad, moderately longer than sutural length of elytra, widest around below of basal half, basal margin weakly rounded; pubescence dense, directed laterally and posterolaterally; punctation small and shallow but covered with densely and uniformly.

Mesoventrite ( Fig. 28 ): inter coxal process rather narrowly elongate, with rounded apex, completely reaching to inter coxal process of metaventrite.

Metaventrite ( Fig. 28 ): inter coxal process of metaventrite small but rather obtusely pointed apically.

Elytra ( Fig. 1 ): together, transverse (EW/EL = 1.99, n = 20), rather small, slightly smaller than pronotum, widest at middle; fine pubescence scattered densely, diverging posteriorly on each elytron; dorsal surface rough and impressed moderately; posterolateral corners of each elytron not sinuate.

Legs ( Fig. 1 ): simple, short, moderately slender; fore and midtibia, densely covered with undeveloped spines, respectively.

Abdomen ( Fig. 1 ): first three visible tergites deeply impressed transversely at base; dorsal and ventral surface moderately covered with thin and long setae.

Male: tergite VIII ( Fig. 45 ): posterior margin smooth, not serrate at all, broadly shallowly emarginate medially; dorsal surface covered with setae rather densely, with approximately 11 macrosetae. Sternite VIII ( Fig. 47 ): posterior margin broadly rounded, with a row of minute sensory setae densely (see Yamamoto & Maruyama, 2012: 10), and with a complex-cluster of very minute setae just behind the row of minute setae; ventral surface covered with setae densely, with approximately 12 macrosetae. Median lobe of aedeagus ( Figs 49–50 ): [lateral view ( Fig. 49 )]: robust, gradually narrowing apically; apical lobe pointed, not hook-shaped at apex; [parameral view ( Fig. 50 )]: bulbous, with basal part widely rounded; apical lobe abruptly narrowing moderately near apex, with thin and sharply pointed apex; a series of complex sclerites, reaching to almost basal margin of median lobe.

Female: tergite VIII ( Fig. 51 ): posterior margin as in male; dorsal surface covered with setae densely, with approximately 11 macrosetae. Sternite VIII ( Fig. 48 ): posterior margin very weakly rounded broadly or almost truncate; ventral surface covered with setae densely, with approximately 13 macrosetae; no complex-cluster of minute setae situated near posterior margin. Spermatheca ( Fig. 51 ): C-shaped; apical invagination of spermatheca very small, inconspicuous, shallowly inward; spermathecal head and neck weakly elliptical shape together; attachment of spermathecal duct inconspicuous; basal part of spermathecal stem widely curved, much longer than spermathecal head and neck combined; chitinized portion of spermathecal stem moderately long, as long as spermathecal head and neck combined; each part of spermatheca uniformly and weakly sclerotized; inner wall of spermathecal head and neck coarsely striate, that of basal part not reticulate at all.

Distribution. Japan (Hokkaidô, Honshû, Shikoku, Kyûshû; Okushiri-tô Is., Tsu-shima Is.); North Korea, South Korea, China.

Host records. No host record is available.

Remarks for type material. According to Park et al. (2011), Aleochara lewisia is represented by only the holotype and is regarded as a synonym of A. coreana . There are, however, two syntypes in BMNH and FMNH. Also, according to the original description, it seems that multiple type specimens were used. The syntype of A. lewisia in BMNH is considerably smaller than the syntypes of A. coreana (e.g., Fig. 107 ), and the shape of the aedeagus is slightly different from that of A. coreana . However, the general structures of the median lobe and the paramere are almost the same as those of A. coreana . In contrast, the syntype of A. lewisia in FMNH is medium in size, and male genitalic structures were identical to those of A. coreana . The syntype of A. lewisia in BMNH is considered to be an individual variation of A. coreana .

Type materials of A. coreana were collected from Seiskin and Pu Ryong, Korea.

Comments. This is probably the most common Aleochara species in Japan. Numerous specimens were found across all of mainland Japan ( Fig. 121 ). Many A. coreana specimens have been identified erroneously as either A. curtula or A. parens . Japanese records of A. coreana are limited to only A. lewisia , which was already synonymized with A. coreana .

Figures

FIGURES 1 – 9. Habitus of Japanese species of Aleochara (Aleochara) with the excluded taxa (dorsal view): 1, Aleochara (Aleochara) coreana; 2, A. (A.) curtula; 3, A. (A.) lata; 4, A. (A.) parens; 5, A. (A.) postica with yellowish elytra from Hokkaidô, Japan; 6, A. (A.) postica with blackish brown elytra from Amami-ôshima Is., Japan; 7, A. (A.) yaeyamensis n. sp.; 8, A. (Xenochara) niponensis sp. rev.; 9, Paraleochara kochi n. comb. Scale bars: 1.5 mm.

FIGURES 10 – 18. Lateral habitus of Japanese species of Aleochara (Aleochara) with the excluded taxa: 10, Aleochara (Aleochara) coreana; 11, A. (A.) curtula; 12, A. (A.) lata; 13, A. (A.) parens; 14, A. (A.) postica with yellowish elytra from Hokkaidô, Japan; 15, A. (A.) postica with blackish brown elytra from Amami-ôshima Is., Japan; 16, A. (A.) yaeyamensis n. sp.; 17, A. (Xenochara) niponensis sp. rev.; 18, Paraleochara kochi n. comb. Scale bars: 1.5 mm.

FIGURES 19 – 27. Head of Japanese species of Aleochara (Aleochara) with the excluded taxa (dorsal view): 19, Aleochara (Aleochara) coreana; 20, A. (A.) curtula; 21, A. (A.) lata; 22, A. (A.) parens; 23, A. (A.) postica with yellowish elytra from Hokkaidô, Japan; 24, A. (A.) postica with blackish brown elytra from Amami-ôshima Is., Japan; 25, A. (A.) yaeyamensis n. sp.; 26, A. (Xenochara) niponensis sp. rev.; 27, Paraleochara kochi n. comb. Scale bars: 0.3 mm.

FIGURES 28 – 36. Meso- and metaventrite of Japanese species of Aleochara (Aleochara) with the excluded taxa (dorsal view): 28, Aleochara (Aleochara) coreana; 29, A. (A.) curtula; 30, A. (A.) lata; 31, A. (A.) parens; 32, A. (A.) postica with yellowish elytra from Hokkaidô, Japan; 33, A. (A.) postica with blackish brown elytra from Amami-ôshima Is., Japan; 34, A. (A.) yaeyamensis n. sp.; 35, A. (Xenochara) niponensis sp. rev.; 36, Paraleochara kochi n. comb. Scale bars: 0.5 mm.

FIGURES 37 – 44. Antenna of Japanese species of Aleochara (Aleochara) with the excluded taxa (right, dorsal view): 37, Aleochara (Aleochara) coreana; 38, A. (A.) curtula; 39, A. (A.) lata; 40, A. (A.) parens; 41, A. (A.) postica with yellowish elytra from Hokkaidô, Japan; 42, A. (A.) yaeyamensis n. sp.; 43, A. (Xenochara) niponensis sp. rev.; 44, Paraleochara kochi n. comb. Scale bars: 0.5 mm.

FIGURES 45 – 51. Terminalia of Aleochara (Aleochara) coreana: 45, male tergite VIII, dorsal view; 46, female tergite VIII, dorsal view; 47, male sternite VIII, ventral view; 48, female sternite VIII, ventral view; 49, median lobe of male aedeagus, lateral view; 50, ditto, parameral view; 51, female spermatheca. Scale bars: 0.5 mm (45 – 48); 0.2 mm (49 – 50); 0.1 mm (51).

FIGURES 107 – 119. Type materials of Japanese species of Aleochara (Aleochara) with the excluded taxa (dorsal view): 107, syntype of Aleochara coreana; 108, syntype of A. lewisia; 109, ditto, re-mounted; 110, lectotype of A. discoidea; 111, lectotype of A. parens; 112, paralectotypes of A. parens; 113, holotype of A. postica; 114, lectotype of A. claviger; 115, syntype of A. nitouensis; 116, ditto; 117 – 118, lectotype of A. niponensis; 119, holotype of A. kochi. Deposited in BMNH: 107 – 114, 117 – 118. Deposited in FMNH: 115 – 116, 119.

FIGURES 121 – 122. Collecting localities of Aleochara (Aleochara) spp. in Japan, based on the specimens examined in this study: 121, Aleochara (Aleochara) coreana; 122, A. (A.) curtula.