Aeolodontini Johnson et al. 2020a

Young, Mark T., Wilberg, Eric W., Johnson, Michela M., Herrera, Yanina, Brandalise, Marco de Andrade, Brignon, Arnaud, Sachs, Sven, Abel, Pascal, Foffa, Davide, Fernández, Marta S., Vignaud, Patrick, Cowgill, Thomas & Brusatte, Stephen L., 2024, The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha), Zoological Journal of the Linnean Society 200 (2), pp. 547-617 : 594-595

Aeolodontini Johnson et al. 2020a (Zoological Code)

Aeolodontini Johnson et al. 2022b: 3 (PhyloCode)

RegNum registration number 758.

Type genus: Aeolodon von Meyer, 1832 .

Etymology

‘Pertaining to Aeolodon ’. The stem Aeolodont - is based on the genus Aeolodon . - ini is a Latin masculine plural suffix for ‘pertaining to’. Under Article 29.2 of the Zoological Code, the suffix - ini denotes a tribe rank within the family-group.

Geological range

Middle Jurassic (middle Callovian) to Late Jurassic (early Tithonian) ( Johnson et al. 2020a).

PhyloCode phylogenetic definition

The largest clade within Teleosauroidea containing Aeolodon priscus (von Sömmerring 1814) but not Steneosaurus megistorhynchus ( Eudes-Deslongchamps 1866) and Teleosaurus cadomensis ( Lamouroux 1820) . Definition from Johnson et al. (2022b).

Reference phylogeny

Fig. 3 .

Zoological Code diagnosis and PhyloCode diagnostic apomorphies

Teleosauroid crocodylomorphs with the following unique combination of characters (10): external nares has a distinct ‘8’- shape (97.1); narial fossa expanded, also creating an ‘8’-shape (shared with the unnamed Mystriosaurus subclade) (102.2); basioccipital tuberosities reduced (459.0); first two premaxillary alveoli do not form a couplet but are positioned on the anterior margin of the premaxilla (579.1); first two premaxillary alveoli are oriented laterally when seen in palatal view (580.2); first two premaxillary alveoli are in the same transverse plane (shared with Pholidosauridae ) (582.1); premaxilla lateral margins sub-rectangular, with the third premaxillary alveoli being clearly lateral to the second premaxillary alveoli (583.1); final premaxillary tooth is evidently anterolateral to the first maxillary tooth (shared with Pholidosauridae and derived Goniopholididae ) (593.2); ulna olecranon process greatly expanded (shared with machimosaurine machimosaurids) (768.1); tibia evidently shorter than the femur (shared with machimosaurine machimosaurids and geosaurine metriorhynchids) (809.3–4).

Composition

Aeolodon , Bathysuchus , Mycterosuchus , and Sericodon .

Comments

Authorship: The nomen Aeolodontini was first used by Johnson et al. (2022b: 3); however, it was exclusively defined under the PhyloCode. Herein, we ensure that the tribe is valid under the Zoological Code. Note, the nominal author is Johnson et al. (2020a), who validly established Aeolodontinae. The nominal author of a family-group is the author who first erected a family-group taxon that is valid (in fulfilment of Article 11), and in accordance with the Zoological Code Principle of Coordination applied to family-group names (Article 36.1).

Steneosauridae: The name Aeolodontinae is not used herein. In Johnson et al. (2022b), we also did not phylogenetically define Aeolodontinae under the PhyloCode. We have explicitly left the clade unnamed in our ergotaxonomies, as we are awaiting the Steneosaurus - type species application to the ICZN to be resolved (Young et al., in review a), and the potential resurrection of the Steneosaurus -based family-group.

Figures

Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/.