Aclista pseudosoror, Macek, 2007

Macek, Jan, 2007, Revision of Central European species of the Aclista scutellaris complex (Hymenoptera: Diapriidae), Acta Entomologica Musei Nationalis Pragae 47, pp. 211-228 : 220-223

Aclista pseudosoror sp. nov.

( Figs. 6 , 14, 22 , 30, 34)

Aclista soror: NIXON (1957) : 68 (nec Kieffer, 1909: 532), misinterpretation.

Type locality. Czech Republic, eastern Bohemia, Orlické hory Mts., Orlické Záhoří env., meander of Orlice river. Type material. HOLOTYPE: ♀, ‘CZ, Orlické hory Mts., 12.ix.1995, meandr Orlice (5764), J. Macek lgt. et det.’

( NMPC). ALLOTYPE: J, ‘CZ, Bohemia bor., Sokolov dist., Starý Jiří (strip mine) (5742), 6.vii.2001, J. Macek lgt et det.’ ( NMPC) . PARATYPES: CZECH REPUBLIC: BOHEMIA centr., Nižbor (5949), 26.vi.1982, ♀ ; Vůznice , river (5950), 16.viii.1984, ♀ ; Kokořínský důl Nature Reserve (5553), 20.viii.1982, ♀ ; Sázava (6155), 22.viii.1985, ♀ ; Ondřejov (6054), 18.viii.1988, ♀ ; Roztěž (6057), 17.viii.1985, ♀. BOHEMIA mer., Purkarec (6852), 12.vi.1982, ♀ ; Třeboň env, Hamr (7055), 31.v.1983, ♀ ; Písecké hory hills, Vysoký Kamýk hill (6751), 26.viii.1982, ♀. BOHEMIA or., Králický Sněžník Nature Reserve , Mlýnský potok brook (5866), 22.viii.2001, ♀ ; Orlické hory Mts.: Trčkov Nature Reserve (5664), 11.x.1995, ♀, 1.x.1997, ♀, 26.viii.-15.ix.1994, 2 ♀♀ ; Zdobnice (5764), 25.viii.1994, ♀ ; Orlické Záhoří (5764), 12.ix.1996, ♀ ; Kačerov (5764), 20.-28.vi.1984, ♀ ; Železné hory Mts.: Polom Nature Reserve (6160), 26.-30. vii.1999, ♀, Farkač lgt. BOHEMIA occ., Sokolov district : Starý Jiří opencast (5742), 6.vii.2001, Ježek et Chvojka lgt. MORAVIA mer., Podyjí National Park , Klapperův potok brook (7161), 11.viii.1992, 3 ♀♀ ; Baštův mlýn mill (7161), 12.viii.1991, J ; Braitava hill (7160), 14.viii.1992, 2 ♀♀ ; Lanžhot env., Ranšpurk Nature Reserve (7367), 8.viii.1991, ♀ ; Bílé Karpaty Mts., Velká Javořina Mt. (7171), 8.viii.1991, J. SLOVAKIA centr., Polana Nature Reserve , Hrončecký grúň hill (7382), 2.viii.1986, 2JJ ; Klenovský Vepor hill (7384), 29.vii.1986, J ; Slovenské Rudohorie Mts., Dobročský prales Nature Reserve (7284), 30.vii.1986,J. SLOVAKIA occ., Čachtice (7272), 11.viii.1991 ; all J. Macek lgt. (unless stated otherwise) et det. ( NMPC). GERMANY: BAVARIA, Spiegelau, 7.-11.ix.1995, ♀, Barták lgt., J. Macek det ( NMPC). AUSTRIA: UPPER AUSTRIA, Mühldorf – Hochreith, Scharenstein , 550-600 m a.s.l., 20.vii.1977, ♀ ; STYRIA: Gragger Schlucht, Neumarkt, 900-1000 m a.s.l., 10.vii.1971, J, elevation NO Further Teich pond, Mariahof, 870-890 m a.s.l., 2.vii.1971, J, all Fischer lgt., all J. Macek det. ( NHMW, NMPC). POLAND centr., Modrzewina Nature Reserve , 26.vii.1981, 1♀, 24.viii.1981, ♀, 17.-24.vi.1981, J ; Klembow, Debina Nature Reserve, 15.-24.x.1980, J, all Ekipa lgt., all J. Macek det. ( NMPC). SPAIN: Pyrenées or., Val d’Aran , 23.viii.1993, ♀, J. Macek lgt. & det. ( NMPC) .

Diagnosis. Mandibles prominent, sickle-shaped, widely overlapping; postmalar area smooth and surrounded by strong malar keel; malar keel not connected with occipital carina; antennae 14-segmented; flagellomeres submoniliform; pronotal shoulders prominent; epomia develo- ped; metascutellum rounded; inner plicae of propodeum converging, projecting posteriorly; radial cell closed, longer than marginal vein; petiole cylindrical and smooth, with longitudinal ribs; basal sculpture of macrotergite with long medial furrow and short irregular lateral striation; males with modified setae on the inner side of protibia.

Description. Female. Length: 2.75 mm. Dark brown; head black; antennae and appendages fulvous.

Head strongly transverse in dorsal view with prominent antennal sockets; antennal sockets between toruli with shallow cleft; malar area smooth, postmalar area developed and smooth, in part surrounded by strong arcuate malar keel; occipital carina incomplete, finely crenulate; vertex convex; temples converging posteriorly; ocelli large; transverse diameter of eye a little shorter than malar space; eyes pubescent; head in lateral view higher than long; head in frontal view subtriangular, face lustrous; antennal sockets smooth; subantennal furrows absent; epistomal sulcus developed; tentorial pits surrounded by deep hollows; mouth aperture shorter than malar space; clypeus slightly convex in lateral view, smooth; mandibles strongly sickle-shaped, widely overlapping. Antennae 14-segmented; flagellum with semidecumbent short pubescence; proximal flagellomeres subcylindrical, distal ones subquadrate.

Mesosoma convex, as wide as head; pronotal collar indistinct, pronotal shoulders angular; epomia strongly developed; lateral pronotum strongly impressed, smooth and intensively lustrous, not perceptible from above; rim connecting pronotal spiracle with tegula distinct; mesoscutum convex; notauli percurrent, diverging posteriorly; scutellum convex, smooth, with large rectangular fovea; axillar depressions smooth, with a very fine tuft of pubescence; mesopleuron smooth with large impression in middle; metascutellum tuberculate with short medial keel; lateral sides of metanotum smooth; propodeum subrectangular, sparsely pubescent with distinct, slightly convex, upraised posterior ledge; medial keel of propodeum simple; both inner and outer plicae converging and projecting posteriorly. Wings translucent; marginal vein distinctly shorter than parastigma; radial cell closed, longer than marginal vein; stigmal vein slightly oblique to marginal vein, shorter than marginal vein.

Metasoma. Petiole cylindrical, smooth, with longitudinal ribs, 1.5 times as long as wide; gaster fusiform, sparsely pubescent; base of macrotergite slightly wider than width of petiole; basal sculpture of macrotergite with strong fan-shaped striation and fine medial furrow; praepygidial segments narrow and closely abutted; pygidium very short and upcurved.

Male. Differs from the female by the cylindrical flagellomeres, modified flagellomere 1, and heterogeneous pubescence of the fore tibia.

Differential diagnosis. Aclista pseudosoror sp. nov. is closely related to A. pseudobitensis sp. nov., differing by the following characters: i) apical flagellomeres subquadarate (females); ii) occipital keel not crenulate (both sexes); iii) gaster stout (female); iv) praepygidial segments closely abutted (female); v) radial cell 1.5 times as long as marginalis (both sexes); vi) emargination of flagellomere 1 large, covering more than half of the segment.

Variability. No significant variation was observed in the available material examined.

Bionomics. Poorly known; flight period May to October.

Etymology. The species name refers to Nixon’s misinterpretation of A. soror .

Distribution. England and Sweden ( NIXON 1957, as A. soror auct.); Austria, Czech Republic, Germany, Poland, Slovakia, and Spain.

Figures

Figs. 1-11. 1-8 – metasoma. 1 – Aclista angusta (Kieffer, 1909); 2 – A. bitensis (Kieffer, 1909); 3 – A. crassistilus (Kieffer, 1909); 4 – A. neglecta (Kieffer, 1907); 5 – A. pseudobitensis sp. nov.; 6 – A. pseudosoror sp. nov.; 7 – A. scutellaris (Thomson, 1859); 8 – A. szelenyii (Móczár, 1938). 9-10 – head, frontal view. 9 – A. angusta; 10 – A. bitensis; 11 – A. crassistilus. Scale bars: 2.0 mm (Figs. 1-8) and 0.5 mm (Figs. 9-10).

Figs. 12-24. 12-16 – head, frontal view. 12 – Aclista neglecta (Kieffer, 1907), 13 – A. pseudobitensis sp. nov.;14 – A. pseudosoror sp. nov.; 15 – A. szelenyii (Móczár, 1938); 16 – A.scutellaris (Thomson, 1859). 17-24 – fore wing, radial cell region. 17 – A. angusta (Kieffer, 1909); 18 – A. bitensis (Kieffer, 1909); 19 – A. crassistilus (Kieffer, 1909); 20 – A. neglecta; 21 – A. pseudobitensis sp. nov.; 22 – A. pseudosoror sp. nov.; 23 – A. scutellaris; 24 – A. szelenyii. Scale bars: 1.0 mm.