Achilia nipponobythoides, Sabella & Cuccodoro & Kurbatov, 2019

Sabella, Giorgio, Cuccodoro, Giulio & Kurbatov, Sergey A., 2019, A revision of the Chilean Brachyglutini - Part 4. Revision of Achilia Reitter, 1890: A. puncticeps and A. approximans species groups, with description of seven new species (Coleoptera: Staphylinidae: Pselaphinae), Revue suisse de Zoologie 126 (1), pp. 127-145 : 141

publication ID	https://doi.org/ 10.5281/zenodo.2619528
DOI	https://doi.org/10.5281/zenodo.3706249
persistent identifier	https://treatment.plazi.org/id/03E587B3-0D34-3301-EB0D-E0E06D43F99C
treatment provided by	Carolina
scientific name	Achilia nipponobythoides
status	sp. nov.

Treatment

Achilia nipponobythoides View in CoL n. sp.

Figs 5 View Figs 1-6 , 15 View Figs 15-18 , 20, 28 View Figs 19-28 , 36, 38, 40 View Figs 35-40 , 54-56 View Figs 51-56 , 58 View Fig

Holotype: MHNG (# MHNG-ENTO-13845); 1 ♂; SOUTHERN CHILI: Región Los Lagos: Osorno prov.: Puyehue National Park, Antillanca road; 500-1000 m; 18-20.XII.1984; S. & J. Peck; car netting.

Paratypes (3): SOUTHERN CHILI: Región Los Lagos: Llanquihue prov.: FMNH (FMHD #97-11); 1 ♂; Vicente Perez Rosales National Park, SW slope Volcàn Osorno, km 10.1 to La Burbuja; 41° 08.30’S 72° 32.15’W; 925 m; 03-27.I.1997; A. Newton & M. Thayer 988; Nothofagus dombeyi & Podocarpus nubigena w/Valdivian rainforest understory, flight intercept trap. – Osorno prov.: FMNH; 1 ♂; Puyehue National Park, Antillanca road; 720-1000 m; 18-24.XII.1982; A. Newton & M. Thayer; Nothofagus ssp. forest , screen sweeping at dusk. – FMNH; 1 ♂; Vicente Perez Rosales National Park, N slope Volcàn Osorno, road to Ref. La Picada; 41° 03.25’S 72° 30.18’W; 660 m; 16.XII.2002; A. Solodovnikov, A. Newton & M. Thayer 1067; Nothofagus dombeyi w/conifers dense Chusquea bamboo understory, flat area, Berlese, leaf & log litter.

Description: Body 1.45-1.60 mm long, reddish-brown, with darker abdomen, reddish elytra, antennae and legs, and yellowish palpi. Head with protruding eyes about as long as convex temples. Pronotum slightly wider than head; median antebasal fovea smaller than lateral ones. First abdominal tergite with basal striae extending to about one-third of paratergal length, and separated at base by about one-third of tergal width.

Male: Head as in Figs 36, 38 and 40 View Figs 35-40 ; posterior part raised, slightly punctuate and pubescent; anterior part at middle flattened, densely punctuate and pubescent, expanded just above eyes in two round shallow depression more densely punctuate and finely pubescent, and with strongly convergent anterior margins giving a triangularshaped forehead; vertexal sulcus absent. Antennae ( Figs 15 View Figs 15-18 , 54-56 View Figs 51-56 ) with scape and pedicel distinctly longer than wide; antennomere III as wide as long; antennomere IV slightly wider than long; antennomere V slightly longer than wide; antennomere VI-VII wider than long with protruding lateral margins; antennomere VII transverse; antennomere VIII as wide as long; antennomere IX with mesal margin projecting in bilobed lamina with large apical setae; antennomere X strongly transverse, apical half of mesal margin projecting and bearing a tuft of setae, one of which particularly long and thin; antennomere XI elongate and longer than VIII-X combined, bearing long and thin subbasal seta inserted in deep impression. Metaventrite slightly raised at middle, this area with median depression. Legs with trochanters very elongate; protrochanters ( Fig. 20 View Figs 19-28 ) bearing tuft of setae; mesotrochanters ( Fig. 28 View Figs 19-28 ) unarmed; profemora and mesofemora slightly thickened; distal half of metatibiae slightly sinuate. Abdominal tergites unmodified. First abdominal ventrite with distinct median carina extending from posterior margin to about one-third of ventrite length. Aedeagus ( Fig. 5 View Figs 1-6 , dorsal longitudinal struts not shown) 0.30 mm long; medial sclerites apically pointed, and associated on each side with one pointed sclerite.

Female: Unknown.

Collecting data: Collected in December and January in Nothofagus ssp. mixed forests by flight intercept traps and screen sweeping at dusk, and also by car netting and sifting.

Distribution: Achilia nipponobythoides n. sp. is known from Southern Chile ( Fig. 58 View Fig : blue triangles) only in the Region of Los Lagos (Llanquihue and Osorno provinces).

Comments: Achilia nipponobythoides n. sp. is easily distinguished from the other members of the genus by the very peculiar morphology of the male head ( Figs 36, 38, 40 View Figs 35-40 ), features of the antennae ( Figs 15 View Figs 15-18 , 54-56 View Figs 51-56 ), and the copulatory pieces of the aedeagus ( Fig. 5 View Figs 1-6 ).