OMOPHRON ROTUNDATUM CHAUDOIR, 1852 = rotundatum ( Bänninger, 1915 ) OMOPHRON SMARAGDUS ANDREWES, 1921 A review of genus Omophron Latreille, 1802 (Coleoptera: Carabidae) Palearctic fauna and distribution Valainis, Uldis Baltic Journal of Coleopterology 2010 10 2 105 128 6SLVC TIAN & DEUVE, 2000 TIAN & DEUVE 2000 Insecta Carabidae Omophron Animalia Coleoptera 12 117 Arthropoda species pseudotestudo  Description: lenght: 6.5 – 7.0 mm; width: 4.3 – 4.6 mm. Colour: testaceous, underside excluding sternites piceous, sternites brown, a patch on each side of pronotum somewhat coriaceous, the patch on median area of pronotum, with a narrow projection in middle to frons margin, and the elytral pattern dull green, epipleura of elytra and pronotum yellowish. Head moderately convex, clypeus quadrisetose, joint 1 of antennae bisetose at apex, 3-4 joints plurisetose along outer margin; penultimate joint of labial palpi plurisetose in the inner side, penultimate joint of maxillary palpi with several apical setae, distal joint glabrous; subocular ridge rugose in front. Pronotum moderately convex, transverse, somewhat bordered on basal half of lateral margin; green pattern narrow, reaching base, but not extending to front margin; punctuation on pronotum rather dense on the green patch except the central area, more coarsely in front and at base. A obvious break in the contour at the junction of pronotum and elytra. Elytra very convex, oval, bordered at base, 15- punctate-striae, striae very deep, punctures rather small. Intervals of elytra with an irregularly row of tiny punctures. Striae 3-4, 7-8, and 11 ended at before 1/4 from apex; intervals 14 and 15 not interrupted by the green patch Comparisons: Close allied to  O. testudoAndrew., but clypeal sulcus arcuate at middle, subocular ridge not extending near to gular suture, and intervals 14 and 15 not interrupted by the green patch. Comparing with  O. maculosumChaud., the green pattern is more dull, patches on pronotum narrower, but extending base of pronotum and body a little bigger.   Distribution:known only from typelocality – (Jianhong, Yunnan provincein South China).   Type locality:Jianhong, China ( holotypeis deposited in South China Agricultural University, Guangzhou, China).   References: Tian & Deuve, 2000    OMOPHRON ROTUNDATUM CHAUDOIR, 1852  = rotundatum( Bänninger, 1915)   Description:lenght: 6.0 – 6.5 mm; width: 3.8 - 4.4 mm. The patch on the pronotum usually does not reach the base margin, but in top it slightly streches over the middle part. On the basic margin of elytra the spots are small, reaching only the elytral striae8 – 9; sometimes they are hardly visible or completely reduced. The middle segment of the elytra pattern is interrupted or, more rarely, narrowed in the area about the elytral striae 7-9; the top segment is shortened, reaching only the elytral striae 7, 8. The specimens that have the elytra pattern similar to that of  O. limbatum(F.) are found very rarely. Elytral interval between elytral striae in the basic part usually are slightly convex. The vertex of the head between punctures has marked longitudinal plications. Puncturation on the sides of the sternites 1 and 2 is sparse, sometimes difficult to see, but it never disappears (contrary to the data presented by Bänninger (1956)). As to the body size and structural features, including also the form of aedeagus, the species is similar to  O. limbatum(F.). The dark pattern of the  O. rotundatumChaud.surface is less developed and the surface microsculpture usually is coarser than that of  O. limbatum(F.), especially of females. The most peculiar difference is the comparatively smaller and darker pattern on the pronotum. The elytral pattern is as presented in Fig. 2.3. The southeast border of the distribution of the species is not clear. O. L. Krizhanovskij (1982) doubted correctness of the data ( Andrewes, 1929) on the distribution of this species in Southern Chinaand Wietnam; he considered those data referring to  O. saigonenseChaud.There are the data ( Tian & Deuve, 2000) also on the finding of  O. rotundatumChaud.in HainanIsland, but, as the authors themselves have pointed out, the morphologic features of the definite specimen do not correspond to the description of the speices given by Andrewes (1929). Still, having compared the specimen with the holotypeof the species ( Chaudoir, 1852), the authors have included it into the species  O. rotundatumChaud.   Distribution:PAR:AB; AF; AR; IR; IQ; IS; KZ; LB; PA; SY; TD; TM; TRa; UZ; CH: HAI; XIN; IN: HP  Processed material:Caucasus, Arexeststhal, Leder Reitter ( 2 Males) (ETHZ);  Afganistan:  Panjao,  25.07.1948( 1 Male, Haarlovleg.) (BMNH); El Hedjaz, 1915 ( 2 Females, Mellingenleg.) (BMNH); Tr. Casp., Aschabad( 1 Female) (BMNH); Transkasp., Kuschka( 1 Female) (BMNH);  Bamyan Province, Panjab( 1 Female, N. Harlovleg.) (ZMUC);  Armenia:  Caucasus, Kasikoporan( 1 Male, 4 Females) (ETHZ); Aresh Distr., Geok-Tapa( 1 Male) (ETHZ); Geok-Tapa, A. Schelkownikowleg. ( 2 Males, 1 Female); Geok-Tapa, L. Mesmin leg. ( 1 Male, 3 Females); Geok-Tapa, Clermontleg. ( 3 Females) (ETHZ); Marmarik River valley, Tarkjarluvill.,   1900 m, 40 o37’ N, 44 o31’ E,  23.06.1997( 1 Male, Melnikleg.) (MPUC);  Azerbaijan:  Ganja, ( 16 Males, 26 Females) (ETHZ);  Lankaran, Talish,  02.06.1936(3, Arnoldileg.) (ZIN); NakhichevanAutonomous Republic,  Ordubad Province, Arakas05.1882 (1, Xristoerleg.) (ZIN); Nahich,  27.06.1933(1, Znajkoleg.);  China:  Tian-Schan( 1 Male) (ETHZ);  Iran: Elburs– Gebirge, Nord – Persien, Rudbar, 800 m, B.v. Bodemeyer ( 1 Male, 1 Female) (ETHZ); Kopet. Dagh ( 1 Female) (ETHZ);  Kerman Province, Sargad, 28.08. –  04.09.1898(21, Zarubnij U.leg.) (ZIN);  Sistan and Baluchestan Province, Sistan, 10. –  11.04.1901(1, Zarubnijleg.) (ZIN); 24. –  26.04.1901(2, Zarubnijleg.),  Sistan and Baluchestan province, Samukan, 1893 (3) (ZIN);  Iraq:  Bagdad, 1915 ( 2 Males, 3 Females) (BMNH); Mesopotamia, Mosul, 05.1909 ( 1 Female, Hauserleg.) (BMNH); Assur, Mesopot, Pietschmann, Mesopotexp. Nat., O. V, ( 1 Male, 1910) (ETHZ); Mesopotamia, Mossui, ( 4 Males, 4 Females) (ETHZ);  Kazakhstan:  Aktobe Province, Kargalinsk,  23.06.1908(1, Malchevskijleg.) (ZIN);  West Kazakhstan Province, Aksaj,  13.04.1909(2) (ZIN);  Kyrgyzstan,  Fergana, Tashkumar, Kara- Suu,  700 m,  22.07.1991( 1 Male, Danilevskijleg.) (MPUC);  Chuy Province, Bishkek, 11-  12.07.1913(1, Chernavinleg.) (ZIN);  Zhambyl Province, Taraz(1, E. Fischerleg.) (ZIN);  Tajikistan:  Kurgan – Tube, Dzhilikul,  18 V 2000( 2 Females) (DUBC);  Sughd Province, Khujand, 06.1977 ( 1 Male, Kukuruzaleg.) (MPUC);  Khatlon Province, Saricashma,  29.06.1986( 1 Male, Semenovleg.) (MPUC); Darvoz district, Pamirmount., Vanch rivervall., 2000 m, 28.05.1980( 1 Female, Mihailov leg.) (MPUC);  Khatlon Region, Kulob08.1933 (1, Popovleg.) (ZIN);  Dushanbe,  28.06.1939(1, Jevrikivaleg.),  14.06.1939(1, Kizanovskijleg) (ZMUM);  Turkmenistan:  Mary Province, Mary, Coll., Hauser, ( 2 Males, 05.1900) (ETHZ);  Mary, Coll., Hauser, ( 1 Female, 04.1900) (ETHZ); Transcaspien, Kuscha?, Koshantchikoffleg. ( 2 Males, 3 Females, 25.-  26.06.1912) (ETHZ);  Mary Province, Badkhiz, Morgunavkavill.,  06.05.1976( 1 Male, Chernjaxovskijleg.) (MPUC);  Mary Province, Badkhiz, Turangagorge,  19.05.1970( 3 Males, Tihomirovaleg.) (MPUC);  Ashgabat, Kaaxkinsk, Dushakvill.,  400 m,  21.06.1992( 2 Males, Danilevskijleg.) (MPUC);  Cardzuj Province  20.05.1905(1, Sumakovleg.) (ZIN),  02.06.1905(1, Sumakovleg.) (ZIN),  15.07.1905(1, Fisher A.leg.) (ZIN);  Ahal Province, Kopetdah Chandir river, Tubligorge  25.07.1935(1, Arnoldileg.) (ZIN); Eylandt(1) (ZIN);  Lebap Province, Repetek,  19.06.1905(1, Sumakovleg.) (ZIN);  Uzbekistan :  Bukhara Province, Bukhara, Hissar Mountains, ( 12 Males, 7 Females) (ETHZ);  Fergana Province, Margelan( 1 Female) (ETHZ); Bukhara( 2 Males, 1 Female) (BMNH);  Bukhara Province, Bukhara, Hissar Mountains(1) (ZIN);  Bukhara Province, Bukhara(2, Staudingerleg.) (ZMUM); Kaukasus, Elisabethpol( 1 Female) (BMNH); Asia Centr., Hissar, Buchara( 1 Male) (BMNH); Surhundarja, Termez, Dzhar– Kurgan, Katta– Kum,  21. IV 2004( 7 Males, 16 Females) (DUBC);  Bukhara Province, Bukhara( 1 Male) (MPUC);  Surkhandarya Province, Termez, Ugkizilbskreservoir,  28.08.1990( 1 Female, Semenovleg.) (MPUC);  Tashkent Province, Tashkent,  06.05.1925( 1 Female, Kirpovleg.) (MPUC); Tashkent Province, Tashkent (2) (ZIN);  Samarqand Province, Toxta-Karagar, 03-  07.07.1996( 1 Female, Veriginleg.) (ZIN);  Surkhandarja Province, Shirabad  28.05.1912(1, A. Kirichenkoleg.) (ZIN).   Type locality:„ Mesopotamia” ( holotypeis deposited in Muséum National d’Histoire Naturelle, Paris, France).   References: Andrewes, 1929; Bänninger, 1915; Bänninger, 1918; Bänninger, 1956; Chaudoir, 1852; Chaudoir, 1868; Gestro, 1892; Hurka, 2003; Kryzhanovskij, 1982; Kryzhanovskij et. al., 1995; Semenov-Tian-Shanskij, 1926; Tian & Deuve, 2000; Valainis, 2009; Zaitzev, 1916;    OMOPHRON SMARAGDUSANDREWES, 1921   Description:lenght: 6.0 – 7.0 mm; width: 4.0 - 4.4 mm. Colour: testaceous, sterna and front margin of prothorax piceous; back and sides of head, a wide transverse patch on prothorax, extending to base and covering its central third, extending aiso to a broad point at middle of apex, and an elytral pattern rather bright green.  Headflat, front nearly smooth, sides and back rather closely striate-punctate, clypeus uneven, quadrisetose, indistinctly bordered, widely emarginate, suture with an angle at middle, lobes of mentum rounded, epilobes not projecting, subocular ridges extending little beyond fissure, area in front of them finely rugose-striate.  Prothoraxmoderately convex, rather coarsely coriaceous, coarsely punctate along base, more finely on disk and along front margin, base only slightly sinuate near hind angles, sides a little rounded at about middle and faintly sinuate before angles, median line fine but distinct, not reaching extremities. Elytra oval, convex, base unbordered, sides rather strongly dilated behind shoulders; 15-punctate-striate, the punctures very distinct, intervals rather flat on disk, more convex at sides, a row of minute punctures down the centre of each. A fine microsculpture is present. The elytral pattern is as presented in Fig. 3.1.  Comparisons:In general appearance strikingly like O. maculosusChaud.. Head less rugose, more conspicuously punctuate, the punctures less confluent; prothorax very obviously punctuate and much less roughly coriaceous (in O. maculosusthe punctures hardly show except along front margin), the green patch larger, the testaceous colour at base and sides separated (or almost to); elytra rather more strongly rounded behind shoulder, the minute puncturation of intervals much less evident, striae rather deeper and more strongly punctuate, front and middle patches not interrupted on interval 13.   Distribution:PAR: IN: UP, UTH  Processed material:   India:Uttarkhand, Kumaon, Ranikhet ( 1 Male, Championleg.) ( Holotype) (BMNH), ( 1 Male, 1 Female, Championleg.) ( Cotypes) (BMNH); Uttarkhand W. Almora, Kumaon, ( 1 Female, Championleg.) (BMNH); Uttarkhand Kuamon, Saryu Valley,  5000 ft.( 3 Males, 3 Females, Championleg.) Uttarkhand, Kuamon( Comparedwith type) ( 1 Female) (ETHZ); Uttarkhand, Ranikhet( 1 Female, Championleg.) (BMNH); Uttarkhand, West Bhatoe, Kumaon,  4000 ft.( 2 Females, Championleg.) (BMNH); Inde Anglaise Pedong, Region de Darjecling, 1935 ( 1 Female) (BMNH)   Type locality:  India, Kumaon, Ranikhet( holotypeis deposited in BMNH).   References:Andrewes, 1921; Andrewes, 1929; Hurka, 2003 China Guangzhou South Agricultural University 12 117 1 holotype 1948-07-25 Haarlov & El Hedjaz & Mellingen & Tr. Casp. & Transkasp. & Kuschka Aschabad Panjao 13 118 5 4 1 N. Harlov Armenia Panjab 13 118 1 1 Bamyan Geok-Tapa, A. & Schelkownikow & Geok-Tapa, L & Geok-Tapa & Clermont Armenia Caucasus Aresh Distr. Kasikoporan 13 118 16 11 5 Bamyan 1997-06-23 Melnik Azerbaijan 1900 40.616665 Azerbaijan 1161 44.516666 13 118 1 1 Azerbaijan Ganja 13 118 42 26 16 Ganca 1936-02-06 Arnoldi Azerbaijan Talish 13 118 1 Lankaran 1933-06-27 Xristoer & Znajko China Nahich Arakas 13 118 1 Ordubad [340,694,1212,1237] Iran Tian-Schan 13 118 1 1 1898-04-09 Zarubnij U. Iran Sargad 13 118 1 Kerman 1901-04-26 1901-11-04 1901-04-26 Zarubnij Iran Sistan 13 118 1 Sistan va Baluchestan Iraq Sistan and Baluchestan province 13 118 1 Sistan va Baluchestan Mesopotamia & Mosul & Hauser & Assur & Mesopot & Pietschmann & Nat., O. V Kazakhstan Mossui Mesopotamia 13 118 15 8 7 Baghdad [555,1316,124,1684] 1908-06-23 Malchevskij Kazakhstan Kargalinsk 13 118 1 Aktobe Province 1909-04-13 Kyrgyzstan Aksaj 13 118 1 Batys Qazaqstan Oblysy 1991-07-22 Danilevskij Kyrgyzstan Fergana 700 Suu Tashkumar 13 118 1 1 Batys Qazaqstan Oblysy 1913-12-07 Chernavin Kyrgyzstan Chuy Province 13 118 1 Bishkek Shaary E. Fischer Tajikistan Taraz 13 118 1 Zhambyl Province 2000-05-18 Tajikistan Dzhilikul Kurgan - Tube 13 118 2 2 Zhambyl Province Kukuruza Tajikistan Khujand 13 118 1 1 Sughd 1986-06-29 Semenov & Pamir Tajikistan Saricashma Vanch river Darvoz district 13 118 1 1 Khatlon Popov Tajikistan Kulob 13 118 1 Khatlon 1939-06-14 1939-06-28 1939-06-14 Jevrikiva & Kizanovskij Turkmenistan Dushanbe 13 118 1 Dushanbe Turkmenistan Hauser Coll. 13 118 2 2 Mary Welayaty 1912-06-26 Transcaspien & Koshantchikoff Turkmenistan Coll. Kuscha Hauser 13 118 6 4 2 Mary Welayaty 1976-06-05 Chernjaxovskij Turkmenistan Morgunavka Badkhiz 13 118 1 1 Mary Welayaty 1970-05-19 Tihomirova Turkmenistan Turanga Badkhiz 13 118 3 3 Mary Welayaty 1992-06-21 Danilevskij Turkmenistan 400 Dushak Kaaxkinsk 13 118 2 2 Ashgabat 1905-02-06 1905-07-15 1905-02-06 Sumakov & Fisher A. Turkmenistan Cardzuj Province 13 118 1 Cardzuj Province 1935-07-25 Arnoldi & Eylandt Turkmenistan Tubli Kopetdah Chandir river 13 118 1 Ahal Welayaty 1905-06-19 Sumakov Uzbekistan Repetek 13 118 1 Lebap Welayaty Uzbekistan Hissar Mountains Bukhara 13 118 19 7 12 Bukhara Province Uzbekistan Bukhara Margelan 13 118 4 2 2 Fergana Province Uzbekistan Hissar Mountains 13 118 1 Bukhara Province 2004-04-21 Staudinger Uzbekistan Bukhara Elisabethpol Kaukasus 13 118 25 17 8 Bukhara Province Uzbekistan Bukhara 13 118 1 1 Bukhara Province 1990-08-28 Semenov Uzbekistan Ugkizilbsk Termez 13 118 1 1 Surkhandarya Province 1925-06-05 Kirpov Uzbekistan Tashkent 13 118 1 1 Toshkent Viloyati 1996-07-07 Verigin Uzbekistan Toxta-Karagar 14 119 1 1 Samarqand Province 1912-05-28 A. Kirichenko Uzbekistan Shirabad 14 119 1 Surkhandarja Province France Museum National d'Histoire Naturelle Paris 14 119 1 holotype Champion & Cotypes & Uttarkhand W. Almora & Kumaon & Uttarkhand Kuamon & Uttarkhand & Kuamon & Compared & Ranikhet & Inde Anglaise Pedong & de Darjecling India Region de Darjecling 1524 Saryu Valley West Bhatoe 14 119 15 10 5 holotype India Kumaon Ranikhet 14 119 1 holotype