Miarus Schoenherr, 1826

Skuhrovec, Jiri, Gosik, Rafal, Caldara, Roberto, Tosevski, Ivo, Letowski, Jacek & Szwaj, Ewelina, 2018, Morphological characters of immature stages of Palaearctic species of Cleopomiarus and Miarus and their systematic value in Mecinini (Coleoptera, Curculionidae, Curculioninae), ZooKeys 808, pp. 23-92 : 48-49

Genus Miarus Schoenherr, 1826

Description.

Measurements (in mm). Body length: 3.80-8.39. Body width (metathorax or abdominal segments I–II) 1.55-2.04. Head width: 0.57-0.83.

General. Body slender, C-curved, rounded in cross section.

Colouration. From black to dark brown head. All thoracic and abdominal segments yellowish, with some asperities.

Vestiture. Setae on body thin, in different colouration, distinctly different in length; piliform.

Head capsule. Head almost rounded, sometimes slightly flattened laterally, endocarinal line present and distinct, more than half the length of frons. Frontal sutures on the head narrow and loosened, but distinct, and ever extended to the antennae. One stemma (st), in the form of a pigmented spot with convex cornea. Dorsum of the epicranium with four or five setae; des3 located anteriorly on epicranium close border with frontal suture. Frons with three or four setae; fs2 absent. Head also with two les and two ves. Epicranial area with two or three pes and more or without sensilla.

Antennae located at the end of the frontal suture on each side, membranous and distinctly convex basal article bearing one very long conical sensorium; basal membranous article with 1-4 sensilla.

Clypeus transverse-shaped, approximately 2.5-3.5 times as wide as long with two cls, and one sensillum (clss) between setae; all very close to margin with frons.

Mouthparts. Labrum with three piliform lms; anterior margin bisinuate. Epipharynx with three finger-like als; with two ams; and 0-2 mes; labral rods (lr) elongated. Mandibles distinctly broad, bifid, teeth of unequal height; slightly truncate; both setae piliform and located apically. Maxilla stipes with one stps, two pfs and one mbs and one sensillum; mala with six finger-like dms, in two sizes, first or first and second dms very long as pfs, next medium length; five vms; all vms distinctly shorter than dms. Maxillary palpi with two palpomeres; basal palpomere with one short mxps and two sensilla; distal palpomere with one sensillum and a group of micro cuticular apical processes. Prelabium oval-shaped, with one prms; ligula with sinuate margin and 2-3 ligs; pre mental sclerite feebly visible. Labial palpi with two palpomeres; each of the palpomeres with one sensillum, distal palpomere with cuticular apical processes. Postlabium with three pms, all located laterally.

Thorax. Prothorax slightly smaller than meso- and metathorax. Spiracle bicameral, placed between the pro- and mesothorax (see, e.g., Skuhrovec et al. 2015). Prothorax with ten prns; two ps; and one eus. Mesothorax with one prs, three pds; one as; two long and one short ss; one eps; one ps; and one eus. Chaetotaxy of metathorax almost identical to that of mesothorax. Each pedal area of the thoracic segments well separated, with 5-6 pda.

Abdomen. Abdominal segments I–III of almost equal length, next abdominal segments decreasing gradually to the terminal parts of the body. Abdominal segment X reduced to four anal lobes of unequal size, the lateral lobes being distinctly the largest, the dorsal and the ventral lobe very small. Anus located terminally. Eight spiracles, bicameral, all spiracles placed medially or anteromedially and functional. Abdominal segments I–VIII with one prs (sometimes abdominal segment VIII without); three pds, pds2 the longest one; one long and one minute ss; two long eps; one ps; one lsts; and two eus. Abdominal segment IX with 3-4 ds; 1-3 ps; and two sts. Abdominal segment X with one minute seta present or absent.