Alouatta palliata (Gray, 1849)

Fonda, Federica, Vezzani, Liam, Mena Aguilar, Luis Ángel, Venegas-Elizondo, Carlos Andrés, Bolaños Brenes, Alexander, Lopez, Dayron Manuel, Romeo, Giuseppe, Sonetti, Dario & Dal Zotto, Matteo, 2025, Mammal diversity and tourism influence in an under-investigated region of Costa Rica, ZooKeys 1260, pp. 37-74 : 37-74

Alouatta palliata (Gray, 1849)

Fig. 2 D Mantled howler

Material examined.

Costa Rica • Puntarenas Province, Karen Mogensen Wildlife Refuge ; 09°52'10"N, 085°03'20"W; 294 m alt.; 15.01.2018; direct observation .

Identification.

Medium-sized primate with dark fur and a brown to orange mantle on sides and rump, particularly marked in males. Face is black. Males show particularly visible white testicles. Frequently recorded thanks to its call, resembling a powerful roar in males ( Wainwright 2007; Reid and Gόmez Zamora 2022). In line with the other monkey that occurs in the Wildlife Refuge ( C. imitator , see above) A. palliata is relatively common and widespread in the study area and in several areas of Costa Rica, but is considered globally vulnerable ( Beal et al. 2020; IUCN 2025).

Figures

Figure 2. Mammals directly observed at the Karen Mogensen Wildlife Refuge. A. Dasypus novemcinctus; B. Dicotyles tajacu; C. Cebus imitator; D. Alouatta palliata; E. Nasua narica; F. Dasyprocta punctata. Photographs by M. Dal Zotto.