Genkyhag innebula, Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

3.1.1.2.1. † Genkyhag innebula sp. nov.

Figures 2C , 5

Etymology.

From Latin 'nebula, In nebula’, meaning "in the fog", in reference to the amber pieces where the specimens are found, that contains numerous terrigenous inclusions, sometimes partially hiding the specimens. It is a noun in apposition.

Material studied.

Holotype female NIGP203546; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China. Three paratypes male FAI-BI 11324a, male FAI-BI 11324b, FAI-BI-11324c of unknown sex; housed in the Fushun Amber Institute.

Type locality.

Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.

Age.

Upper Albian to lower Cenomanian, mid-Cretaceous.

Diagnosis.

As for genus.

Description.

Holotype female incompletely preserved (body length 3.12 mm), in a piece with multiple terrigenous inclusions. Paratypes preserved in a single piece: FAI-BI 11324a visible in dorsal and frontal views, FAI-BI 11324b in lateral view (body length ca. 3.45 mm), FAI-BI 11324c being one complete fore and hind wings and a partial fore wing. - Head globular, apparently entirely glabrous (length for holotype ca. 0.47 mm); frons shagreened, convex, divided by a faintly impressed sulcus; compound eye oval, without postocular carina; vertex convex, shagreened; clypeus short; subantennal groove present; toruli inserted half way between clypeus and eyes; antenna short, barely reaching propodeum; scape globular; pedicel slightly longer; flagellomeres cylindrical, longer than wide; flagellomeres 7-11 shorter than flagellomeres 1-6; flagellomere 12 longest; occipital carina crenulate. - Mesosoma slightly shorter than metasoma (length for holotype 1.12 mm; for paratype 11324a 1. 29 mm; for paratype 11324b 1.57 mm); pronotum not visible dorsally; mesoscutum shagreened, half as long as mesosoma, median mesoscutal sulcus crenulate (mesoscutum length for holotype 0.51 mm; for paratype 11324a 0.61 mm; width for holotype 0.65 mm; for paratype 11324a 0.71 mm); parapsidal lines present; axillae barely contiguous medially and inner grooves crenulate; mesoscutellum wider than long, shorter than mesoscutum; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; propodeum carinate, carinae delimitating row of small foveae in anterior region and larger foveae in posterior region. - Fore wing hyaline, covered with microtrichiae (length for holotype 1.84 mm; for paratype 11324a 2.10 mm; for paratype 11324b 2.11 mm; for paratype 11324c 2.86 mm); venation almost complete, Sc+R, M+Cu, A, Rs+M and M fully pigmented; Cu spectral; marginal cell very narrow, closed in straight line; R1 barely extending beyond marginal cell; medial cell rectangular and relatively broad. - Hind wing hyaline; venation reduced to Sc+R and R1, Sc+R almost not diverging from margin, R1 pigmented almost to tip; at least seven hamuli, located distal to middle of hind wing. - Legs with one mesotibial and two metatibial spurs; metafemur swollen; ventral surface of metatibia and metabasitarsus with row of comb-like setae, metabasitarsus at least three times as long as following metatarsomeres. - Metasoma elongate, oval, with sparse short setae (length for holotype 1.43 mm); tergites smooth; ovipositor shorter than metasoma (length for holotype 1.34 mm; OL/BL ratio 0.43).

Figures

Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H).

Figure 5. † Genkyhag innebula gen. et sp. nov., mid-Cretaceous, Kachin amber. A holotype female NIGP 203546, habitus in dorsal view; B paratype male FAI-BI 11324 a, habitus in dorsal view; C habitus in frontal view; D paratype male FAI-BI 11324 b, habitus in lateral view; E paratype FAI-BI 11324 c, fore and hind wings, with hamuli indicated by arrows. Scale bars: 1 mm (A - E). Abbreviations: AG = inner axillar groove; MMS = median mesoscutal sulcus; OC = occipital carina.