Gasterosteus aculeatus Linnaeus, 1758

Denys, Gaël P. J., Daszkiewicz, Piotr, Urtizberea, Frank & Bernatchez, Louis, 2022, Diadromous fishes from Saint-Pierre and Miquelon archipelago: diagnoses, taxonomy, nomenclature and distribution, Cybium 46 (4), pp. 385-413 : 396-398

Gasterosteus aculeatus Linnaeus, 1758 – NATIVE ( Fig. 3 )

Ga s te ros teu s aculeatu s Linnaeus, 1758: 295 [type locality: Europe; syntypes: LSL 29-31 ( Wheeler, 1985: 43; Fig. 3A )].

Ga s te ros teu s bi s pin os u s Walbaum, 1792: 450 (type locality: New York, USA; no types known) .

Ga s te ros teu s te r aculeatu s Lacepède, 1801: 295, 296 (unneeded new nomen for Ga s te ros teu s aculeatu s Linnaeus, 1758: 295).

Ga s te ros teu s biaculeatu s Shaw, 1803: 608 (type locality: New York, USA; unneeded new name for Ga s te ros teu s bi s pin os u s Walbaum, 1792: 450) .

Ga s te r acanthu s cataph r actu s Pallas, 1814: 229 (type locality: Kamchatka, Russia; no types known) .

Ga s te ros teu s gymnu r u s Cuvier (ex Willughby), 1829: 170 [type locality: England and northward France; no types known; based on Pi s ciculu s aculeatu s of Willughby (1686: 341)] .

Ga s te ros teu s leiu r u s Cuvier in Cuvier & Valenciennes, 1829: 481, 487, Pl. 98 (fig. 4) [type localities: France; Germany; syntypes: MNHN-IC-0000-7088, MNHN-IC-0000-7151 ( Monod, 1973: 281)] .

Ga s te ros teu s t r achu r u s Cuvier in Cuvier & Valenciennes, 1829: 481, 487, Pl. 98 (fig. 1) (type localities: France, Germany; syntypes:?NMW 6918) .

Ga s te ros teu s s emia r matu s Cuvier in Cuvier & Valenciennes, 1829: 493 [type localities: Le Havre and River Braie near Abbeville, France; Syntypes: MNHN-IC-0000-2347, MNHN-IC-0000-2348 and MNHN-IC-0000-7078 ( Bertin and Estève, 1950: 34-35)] .

Ga s te ros teu s s emil or icatu s Cuvier in Cuvier & Valenciennes, 1829: 494 [type localities: Somme River, around Paris, Oise, La Rochelle, Normandy, Caen, Hable d’Ault, Le Treport, Berlin; syntypes: whereabouts unknown; the lot of specimens MNHN-IC-0000-2347 was considered as syntypes of G. s emil or icatu s according to Bertin (1925: 154), however, as they came from Le Havre so one of the type locality of G. s emia r matu s and confirmed as well by Monod (1973: 281), this designation might be a mistake].

Ga s te ros teu s a r gy ro p o mu s Cuvier in Cuvier & Valenciennes, 1829: 498 [type locality: Florence, Italy; syntypes: MNHN-IC-0000-0677 ( Bertin and Estève, 1950: 35)] .

Ga s te ros teu s b r achycent r u s Cuvier in Cuvier & Valenciennes, 1829: 499, Pl. 98 (fig. 2) [type locality: Florence, Italy; syntypes: MNHN-IC-0000-0694 and MNHN-IC-0000-7097 ( Bertin and Estève 1950: 35), SMF 1181] .

Ga s te ros teu s tet r acanthu s Cuvier in Cuvier & Valenciennes, 1829: 499 [type locality: Florence, Italy; holotype: MNHN-IC-0000-1821 ( Bertin and Estève, 1950: 35)] .

Ga s te ros teu s o b o la r iu s Cuvier in Cuvier & Valenciennes, 1829: 500 (type locality: Kamchatka, Russia; no types known) .

Ga s te ros teu s n o veb or acen s i s Cuvier in Cuvier & Valenciennes, 1829: 502, Pl. 98 (fig. 3) (type locality: New York, USA; holotype: MNHN-IC-0000-7074 ( Bertin and Estève, 1950: 35)].

Ga s te ros teu s nige r Cuvier in Cuvier & Valenciennes 1829: 503 [type locality: Newfoundland, eastern Canada; syntypes: MNHN-IC-0000-3057 ( Bertin and Estève, 1950: 36); the nomen Ga s te ros teu s nige r Cuvier, 1829: 170 is erroneous] .

Ga s te ros teu s s pinul os u s Yarrell (ex Jenyns), 1835: 350 (type locality: Edinburgh, Scotland; no types known) .

Ga s te ros teu s l or icatu s Reinhardt, 1837: 114, 193 (type locality: Greenland; no types known) .

Ga s te ros teu s dimidiatu s Reinhardt , 1837: 114, 137, 193 (type locality: Greenland; syntypes: whereabouts unknown) .

Ga s te ros teu s p o nticu s Nordmann, 1840: 380, 613, Pl. 3 (fig. 11) [type locality: Tauria (Crimea), Black Sea; syntypes: MNHN-IC-0000-7098 ( Bertin and Estève 1950: 363), NMW 77244].

Ga s te ros teu s bia r matu s Nordmann, 1840: 381 (type locality: Lagoons near Tarkanckut, Crimea; no types known).

Ga s te ros teu s biaculeatu s DeKay, 1842 (ex Mitchill): 64, Pl. 3 (fig. 9) (type locality: New York, USA; no types known; junior homonym of Ga s te ros teu s biaculeatu s Shaw, 1803: 608) .

Ga s te ros teu s ne o b or acen s i s DeKay, 1842: 66, Pl. 6 (fig. 17) (type locality: New York, USA; no types known) .

Ga s te ros teu s nemau s en s i s Crespon, 1844: 283 (type locality: Nîmes, France; no types known) .

Ga s te ros teu s quad r i s pin os a Crespon , 1844: 283 (type locality: Vistre River and ditches around Nîmes, France; syntypes: whereabouts unknown) .

Ga s te ros teu s cuvie r i Girard in Storer , 1850: 254, Pl. 7 (fig. 1) (type localities: Bras d’Or and Red Bay, Labrador, Canada; no types known) .

Ga s te ros teu s mic ro cephalu s Girard, 1854: 133 [type locality: Tule Lake, San Joaquin Valley, California, USA; syntypes: ANSP 14275-87, ANSP 14288-89 (missing), MNHN-IC-0000-0302, UMMZ 146720 ( Bertin and Estève 1950: 36; Böhlke, 1984: 103)].

Ga s te ros teu s william so ni Girard, 1854: 133 [type locality: upper part of Santa Clara River, Soledad Canyon, Los Angeles County, California, USA ( Miller, 1960); syntypes: MNHN-IC-0000-0303 ( Bertin and Estève, 1950: 36), USNM 340].

Ga s te ros teu s plebeiu s Girard , 1854: 147 [type locality: salt marshes about Presidio, California, USA; Syntypes: FMNH 6038, MCZ 6706, MNHN-IC-0000-0298 ( Bertin and Estève 1950: 36-37)] .

Ga s te ros teu s in o pinatu s Girard , 1854: 147 [type locality: a fresh water lagoon about 1 mile back of Presidio, California, USA; syntypes: MCZ 6701, MNHN-IC-0000- 0300 ( Bertin and Estève, 1950: 37)] .

Ga s te ros teu s s e rr atu s Ayres , 1855: [2] col. 6 [type locality: San Francisco, California, USA; holotype: apparently lost even if Monod (1973: 282) lists a syntype at MNHN] .

Ga s te ros teu s dekayi Ayres, 1855: 48 (objective synonym of Ga s te ros teu s biaculeatu s DeKay; junior homonym of Ga s te ros teu s dekayi Agassiz, 1850).

Ga s te ros teu s in s culptu s Richardson , 1855: 356, Pl. 25 (type locality: Northumberland Sound, western Greenland; holotype:? BMNH) .

Ga s te ros teu s pugetti Girard, 1856: 135 (type locality: Fort Steilacoom , Puget Sound, Washington, USA; no types known) .

Ga s te ros teu s inte r mediu s Girard, 1856: 135 (type locality: Cape Flattery, Washington, USA; no types known) .

Ga s te ros teu s elegan s Blanchard , 1866: 234, Fig. 37 [type locality: between Cadillac and Langon, Gironde, southwestern France; syntypes: MNHN-IC-0000-6673 ( Bertin and Estève 1950: 38)] .

Ga s te ros teu s neu s t r ianu s Blanchard , 1866: 220, Fig. 28 [type locality: Brooks in Harfleur and Gournay, France; syntype: MNHN-IC-0000-7114 ( Bertin and Estève, 1950: 37)] .

Ga s te ros teu s baill o ni Blanchard , 1866: 231, Fig. 34 [type locality: Abbeville, France; syntypes: MNHN-IC-0000- 7151, MNHN-IC-0000-7153 ( Bertin and Estève, 1950: 37)] .

Ga s te ros teu s a r gentati ss imu s Blanchard , 1866: 232, Figs. 35-36 [type locality: Brooks near Avignon, France; syntypes: BMNH 1883.7.4.5-10, MNHN-IC-0000-6675, MNHN-IC-0000-6678 ( Bertin and Estève, 1950: 38), USNM 21140] .

Ga s te ros teu s alge r ien s i s Sauvage, 1874: 17 [type locality: Algeria; syntypes: MNHN-IC-0000-5123 ( Bertin and Estève, 1950: 38)].

Ga s te ros teu s s upp os itu s Sauvage, 1874: 11 (type locality: New York, USA; based on Ga s te ros teu s n o veb or acen s i s Dekay, 1842: 66)

Ga s te ros teu s texanu s Sauvage, 1874: 15, Pl. 1 (fig. 5-5a) [type locality: Texas, USA; holotype: MNHN-IC-0000- 7076 ( Bertin and Estève, 1950: 38)].

Ga s te ros teu s atkin s ii Bean , 1879: 67 [type locality: Schoodic Lakes, Maine, USA; Syntypes: USNM 2249; misspelled atkin s i ( Froese and Pauly, 2021)] .

Ga s t ros teu s h o l o gymnu s Regan , 1909: 436 (type locality: Rome, Italy; syntypes: BMNH 1852.8.17.19-22; original genus should have been Ga s te ros teu s) .

Ga s te ros teu s s antaeannae Regan , 1909: 437 (type locality: Colton, Santa Ana River, California, USA; syntypes: BMNH 1894.2.22.15-17) .

Ga s te ros teu s william so ni jap o nicu s Franz, 1910: 19, Pl. 3 (fig. 10) [type locality: Misaki, Japan; syntypes: ZSM destroyed in World War II ( Neumann 2006: 275)].

Ga s te ros teu s aculeatu s var. me ss inicu s Stephanidis, 1971: 202, Fig. 10 (not available, infrasubspecific; type locality: Pamissos River near Messini, Greece) .

Figures

Figure 3. – Ga�te���teu� aculeatu� Linnaeus, 1758: syntypes LSL 29-31; scale bar = 1cm; with the permission of the Linnean Society of London (A); MNHN-IC-2022-0022, 33.7 mm SL, Saint-Pierre and Miquelon (B).