Ceratophila

Tang, William, Skelley, Paul & Pérez-Farrera, Miguel Angel, 2018, Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae), Zootaxa 4508 (2), pp. 151-178 : 155-156

publication ID

https://doi.org/ 10.11646/zootaxa.4508.2.1

publication LSID

lsid:zoobank.org:pub:5E2BC894-1919-4F63-8EF5-BAAC91913388

DOI

https://doi.org/10.5281/zenodo.5957999

persistent identifier

https://treatment.plazi.org/id/F95B87D7-FFB1-951D-FF49-F8E7D7FA5E38

treatment provided by

Plazi

scientific name

Ceratophila
status

 

Key to adult Ceratophila

Because of variable and overlapping morphological character states with some species, it is desirable, and in some cases necessary, to have a series of specimens representing color variations and both sexes to properly identify a

species. In the following key, text in brackets before the species name are the Mexican states of occurrence and the known host species of Ceratozamia .

1. Pronotum not explanate laterally, surface convexly curved to lateral carinae; lateral carinae in lateral view distinctly thicker anteriorly, anterior thickness 2× posterior thickness ( Fig. 1D View FIGURE 1 ); pronotal disc lacking longitudinal groove extending anteriorly from lateral basal pore in margin ( Fig. 1B View FIGURE 1 ); metatibia triangularly dilated toward apex ( Fig. 1F View FIGURE 1 )......................................................................................... [subgenus Ceratophila, new subgenus] ... 2

- Pronotum explanate laterally, surface broadly concave near lateral carinae; lateral carina weakly thickening anteriorly, anterior thickness <1.5× that at base ( Fig. 1E View FIGURE 1 ); pronotal disc with longitudinal groove extending anteriorly from lateral basal pore in margin ( Fig. 6D View FIGURE 6 ); metatibia not triangularly dilated toward apex ( Fig. 1G View FIGURE 1 )............... [ Vovidesa , new subgenus] … 5

2(1). Head with broad medial transverse ridge anterior of a distinct transverse basal groove in males ( Fig. 2D View FIGURE 2 ), ridge and groove weak in females, ridge interrupting supraocular line in both male and female; head broad, width = 0.74–0.75× pronotal width; anterior clypeal margin emarginate; pronotal hypomeron densely punctate ( Fig. 2E View FIGURE 2 ); [Veracruz— C. euryphyllidia View in CoL ].................................................................................... C. (C.) chemnicki , new species

- Base of head at most with indistinct shallow transverse groove, supraocular line complete ( Fig. 3D View FIGURE 3 ); head narrower, width = 0.66–0.68× pronotal width; anterior clypeal margin truncate or weakly convex; pronotal hypomeron with punctures sparse and minute or lacking ( Fig. 3E View FIGURE 3 ).............................................................................. 3

3(2). Dark elytral markings (when present) laterally covering most of disc ( Fig. 4A View FIGURE 4 ); pronotum generally more elongate, pronotal length/pronotal width (PL/PW) = 0.83–0.90; [Chiapas— C. alvarezii View in CoL , C. mirandae View in CoL , C. norstogii View in CoL , C. vovidesii View in CoL ]......................................................................................... C. (C.) picipennis , new species

- Dark elytral markings (when present) only along suture and laterally ( Fig. 5A View FIGURE 5 ); pronotum generally shorter, PL/PW = 0.80– 0.84................................................................................................ 4

4(3). Color of pronotum and elytra similar ( Fig. 5A View FIGURE 5 ); pronotum generally more quadrate; male genitalia with penile struts relatively long, length of penile strut relative to median lobe 2:1; setae at apex of parameres with greatest length <dorsal width of parameres ( Fig. 5F View FIGURE 5 ); [Veracruz— C. tenuis View in CoL ]......................................... C. (C.) sanchezae , new species

- Pronotum reddish brown contrasting with light brown of elytra ( Fig. 3A View FIGURE 3 ); pronotum generally more trapezoidal, narrowing anteriorly; male genitalia with penile struts shorter, length of penile struts relative to median lobe 3:2; setae at apex of parameres with greatest length ± dorsal width of parameres ( Fig. 3H View FIGURE 3 ); [Oaxaca— C. mixeorum ]...................................................................................................... C. (C.) gregoryi , new species

5(1). Male meso- and metatibiae each with medial subapical emargination ( Fig. 1G View FIGURE 1 ); female abdominal segments weakly alutaceous, punctures and setae visible except on lateral quarters.................................................... 6

- Male tibiae lacking medial subapical emargination; female abdominal segments strongly alutaceous ( Fig. 6B View FIGURE 6 ), punctures obscured across middle, setae reduced [Chiapas— C. alvarezii View in CoL , C. mirandae View in CoL , C. norstogi , C. vovidesii View in CoL ]............................................................................................... C. (V.) chiapensis , new species

6(5). Submentum of male and female with sparse patch of setae-bearing punctures, male with long setae projecting anteriorly ( Fig. 7E View FIGURE 7 ); male genitalia more dorsoventrally compressed, median lobe nearly cylindrical; [Veracruz— C. mixeorum ]........................................................................................ C. (V.) mixeorum , new species

- Submentum of male and female with dense patch of setae-bearing punctures, male with long setae projecting laterally ( Figs. 8 View FIGURE 8 E–F); male genitalia laterally compressed, median lobe laterally flattened and distinctly curved in lateral view; [Veracruz— C. tenuis View in CoL ]....................................................................... C. (V.) vazquezi , new species

Kingdom

Plantae

Phylum

Tracheophyta

Class

Cycadopsida

Order

Cycadales

Family

Zamiaceae

Genus

Ceratophila

Kingdom

Plantae

Phylum

Tracheophyta

Class

Cycadopsida

Order

Cycadales

Family

Zamiaceae

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