Sciasmomyia meijerei Hendel, 1907

Shi, Li, Gaimari, Stephen D. & Yang, Ding, 2013, Revision of Sciasmomyia Hendel (Diptera: Lauxaniidae), with eight new species, Zootaxa 3691 (4), pp. 401-435 : 420-423

Sciasmomyia meijerei Hendel, 1907

( Figs 58–70 )

Sciasmomyia meijerei Hendel 1907: 234 . Type locality: Vietnam, Tonkin. Type: Lectotype male, designated herein (NHMW). Hendel 1908: 18 (listing), plate 3, figs. 43–44 (head), 45 (wing). Shewell 1977: 193 (catalog entry, distribution). Sciasmomyia demeijeri. Misspelling, Hendel 1925: 107 (in key).

Diagnosis. Face with blackish brown median stripe and two brown Y-shaped lateral stripes. Mesonotum with pair of brown spots anteriorly, two grayish median stripes along inside of dc rows (sometimes these two grayish stripes indistinct); anepisternum yellow with a large brown spot near anterior margin and a brown spot near upper margin; katepisternum yellow with a blurry grayish brown middle spot near upper margin. Wing with crossvein r-m incompletely surrounded by hyaline area, except above and below; hyaline spot in cell r4+5 above dorsal tip of crossvein dm-cu. Abdomen yellow to brownish yellow, tergites 2–6 (and 7 in female) each with pale brown to black posterior margin. Male with syntergosternite 7+8 degenerate, spiracle present on membrane; epandrium blackish brown and slender, but broad and strongly clavate apically in lateral view; hypandrium with lateral arms very narrow, hypandrial apodeme much shorter than lateral arms in ventral view; phallus broad with four pairs of lateral processes. Female with hypogynial valve slender with setulae except for bare triangular area at tip, curved upwards apically in lateral view.

Description. MALE. Body length 5.8–6.5 mm, wing length 6.5–6.6 mm. FEMALE. Body length 5.2–5.6 mm, wing length 5.9–6.6 mm.

Head ( Fig. 59 ) yellow. Frons with two wide brown stripes extending to vertex; ocellar triangle black; oc longer than posterior or. Face with blackish brown median stripe and two brown Y-shaped lateral stripes, brown spot at ventral corner, a median spot and two blackish brown transverse stripes on ventral margin; parafacial yellow except for inner margin black, and with brown elliptical spot at ventral corner. Gena about 1/2 height of eye, with a large brown spot. Antenna yellow to brownish yellow, 1st flagellomere 2.0 times longer than high; arista with longest setula nearly as long as 1/4 height of 1st flagellomere. Proboscis brownish yellow to pale brown. Palpus brown.

Thorax ( Fig. 60 ) yellow. Mesonotum with a pair of brown spots anteriorly, two grayish median stripes along inside of dc rows (sometimes these two grayish stripes indistinct); anepisternum yellow with large brown spot near anterior margin and brown spot near upper margin; katepisternum yellow with blurry grayish brown middle spot near upper margin. Legs mostly yellow; fore femur yellow to brownish yellow or only posteroventral area pale brown, with brown inner basal spot and brown ventral spot or half ring on inner side of apical 1/3 (sometimes this spot indistinct); hind femur entirely yellow or pale brown on basal 3/4 of dorsal surface; all basitarsi with a basal ring, tarsomere 2 brown on apical 1/2 and tarsomeres 3–5 entirely brown. Fore femur with 4–5 pv and 5–6 pd, mid femur with 4 weak a, hind femur with 1 weak preapical ad. Wing ( Figs 62–63 ) pale brown with hyaline spots, all of which display white on that part of vein; series of longitudinal hyaline spots along costal margin extending through R2+3, all of which are constricted at middle; series of irregular hyaline spots along R4+5 and along CuA1; crossvein rm pale brown, incompletely surrounded by hyaline area (except above and below); hyaline spot in cell r4+5 above dorsal tip of crossvein dm-cu; costa with 2nd (between R1 and R2+3), 3rd (between R2+3 and R4+5) and 4th (between R4+5 and M1) sections in proportion of 5.3:1.3:1; ultimate and penultimate sections of M 1 in proportion of 1:1.4; ultimate section of CuA1 about 1/7 of penultimate. Halter yellow.

Abdomen yellow to brownish yellow, tergites 2–6 (and 7 in female) each with pale brown to black posterior margin. Female with eversible vescicle extending out from brown pleural membrane at level between tergites 5 and 6. Male genitalia ( Figs 64–67 ): syntergosternite 7+8 degenerate, spiracle present on membrane; epandrium blackish brown and slender, but broad and strongly clavate (ball-like) apically with setulae in lateral view; surstylus consisting of a pair of yellow elliptical sclerites with setulae in posterior view, and top much broader than bottom in lateral view; hypandrium with lateral arms very narrow, hypandrial apodeme much shorter than lateral arms in ventral view; phallus broad with four pairs of lateral processes, distance between last two processes wide in lateral view, and a weak quadrate sclerite with a pair of small mastoid apical processes in ventral view; phallapodeme (fused to phallus) short, narrow. Female terminalia ( Figs 61 , 68–70 ): tergite and sternite 8 yellow; epiproct triangular and hypoproct quadrate in lateral view, hypogynial valve slender with setulae except for a bare triangular area at tip in ventral view, curved upwards apically in lateral view.

Type material. Lectotype 3 (NHMW): Tonkin / Montes Mauson / April, Mai, 2–3000' / H. Fruhstorfer; Sciasmomyia / meijerei Hendel ; Syntype / Sciasmomyia / meijerei / Hendel / det. S.D. Gaimari, 2002; Lectotype 3 / Sciasmomyia / meijerei / Hendel / det. Li, Gaimari, / Yang 2012. Paralectotype: VIETNAM, Tonkin, Montes Mauson, April, Mai, 2000–3000 feet, H. Fruhstorfer (1Ƥ, NHMW). This species was described from two syntypes, one male and one female (although Hendel (1907) indicated two females), represented above. Due to the importance of male genitalia in differentiating species of Lauxaniidae , and in this genus in particular, we here select the male, labeled as above (semi-colon (;) separates labels, and slash (/) separates lines on each label) as the lectotype, to fix the identity of this species.

Other materials. CHINA, Fujian Province: Fukien SE., Gang-keu, 26. VII. 1936, A. L. Melander (13, USNM); Jiangxi Province: Kiangsi SE., Hong San, 25. VI. 1936, A. L. Melander (13, USNM); Zhejiang Province: Longquan, Fengyangshan, Fengyangmiao, 1470 m, 12. XII. 2007, Shenglong Liu (13 CAUC); Longquan, Fengyangshan, Miaopudi, 25. VIII. 2008, Shenglong Liu (13, CAUC); Tianmushan, 31. V. 1937, E. Suenson (1Ƥ, USNM).

Distribution. Vietnam ( Tonkin). New records for China (Fujian, Jiangxi, Zhejiang).

Remarks. This species is very similar to Sciasmomyia leishanensis sp. nov., S. longissima sp. nov., and S. thaii sp. nov., but can be differentiated from them by characteristics of the male and female genitalia.

Figures

FIGURES 58 – 63. Sciasmomyia meijerei Hendel, 1907. Lectotype male and paralectotype female. 58. male habitus, lateral view; 59. male head, anterior view; 60. female thorax, dorsal view; 61. female hypogynial valve, lateral view; 62, male wing; 63. female wing.

FIGURES 64 – 70. Sciasmomyia meijerei Hendel, 1907. Non-type male. 64. epandrial complex, lateral view; 65. epandrial complex, posterior view; 66. phallus complex, ventral view; 67. phallus complex, lateral view. Non-type female. 68. abdomen tergites and sternites 5 – 9, lateral view; 69. hypogynial valve, ventral view; 70. spermathecae. Scale 0.1 mm.