Sciasmomyia Hendel

Shi, Li, Gaimari, Stephen D. & Yang, Ding, 2013, Revision of Sciasmomyia Hendel (Diptera: Lauxaniidae), with eight new species, Zootaxa 3691 (4), pp. 401-435 : 402-403

Sciasmomyia Hendel

Sciasmomyia Hendel 1907: 233 . Type species: Sciasmomyia meijerei Hendel (subsequent designation by Hendel 1908: 17). Hendel 1908: 11 (in key), 17 (diagnosis, type species designation). Shewell 1971: 2 (differentiation from new genus Sciasminettia ). Stuckenberg 1971: 541 (in key). Shewell 1977: 193 (catalog entry). Shatalkin 2000: 22 (in key). Schacht et al. 2004: 49 (in key).

Shatalkinia Papp 1984: 172 (as subgenus of Lyciella ). Type species: Lyciella supraorientalis Papp (original designation). Papp & Shatalkin 1998: 397 (in key). Shatalkin 1999: 539 (in key). Shatalkin 2000: 22 (in key, as subgenus of Sciasmomyia ), 41 (synonymy, subgenus moved to Sciasmomyia ). Schacht et al. 2004: 49 (in key, as subgenus of Sciasmomyia ).

Diagnosis. MALE. Body length 4.4–7.2 mm, wing length 4.9–6.8 mm. FEMALE. Body length 4.2–7.3 mm, wing length 4.8–7.1 mm.

Head: vertex of head acute; frons wider than long and parallel-sided, with pair of brown stripes usually extending to vertex; 2 or, anterior or shorter than posterior or; a brownish spot present at base of each or; oc equal to or longer than posterior or. Fronto-facial angle obtuse; face with brown markings. Antennal 1st flagellomere short, round apically; arista pubescent or only with microscopic setulae, with longest setula shorter than 1/3 height of 1st flagellomere; a blackish brown rounded to subtriangular orbito-antennal spot present. Subgena moderately large, about 1/2 height of gena.

Thorax: mesonotum with a pair of brown spots or short stripes anteriorly, a pair of brownish or grayish median stripes; all setae and setulae with a brown basal spot; postpronotum, anepisternum and katepisternum with brown markings. Chaetotaxy: 1+3 dc; 1–2+3 acr; 1 prsc, equal to or longer than posteriormost dc; 1 anepst; 2 kepst (anterior one weak and hair-like, posterior one much stronger). Scutellum brown to blackish brown except for lateral and posterior margins yellow, with two black apical spots in posterior view. Legs: femora with brown spot or ring on apical 1/3; tibiae with a brown ventral spot on basal 1/3 and a black ring on apical 1/3; fore tibia with 1 long preapical ad and 1 short apv; mid tibia with 1 strong preapical ad and 1 strong apv; hind tibia with 1 long preapical ad, 1 strong curved apv and 1 short apv. Wing pale brown with hyaline spots (except for Sciasmomyia supraorientalis ); costal margin sapromyziform (with a row of small black spinules extending to or slightly beyond tip of R2+3); r-m distal to middle of discal cell.

Abdomen: yellow, brownish yellow to blackish brown; tergites with or without grayish pruinosity, yellow spots or black posterior margins. Male genitalia: syntergosternite 7+8 narrow, forming a complete ring or not, or degenerated to a small structure fused with epandrium; epandrium broad or slender in lateral view; surstylus small to large, articulated with epandrium; hypandrium Y-shaped (except V-shaped in S. tubata ), lateral arms of hypandrium long and apically bent to near tip of phallus; phallapodeme fused to and continuous with base of phallus (except narrowly separated in S. quadricuspis and S. tubata ); pregonite and postgonite absent; ejaculatory apodeme connected to phallus by wide membrane dorsolaterally. Female: eversible vescicle extending out from brown pleural membrane at level between tergites 5 and 6, or absent (i.e., not everted); tergite and sternite 8 confluent, forming a complete ring; tergite 9 (epiproct) and sternite 9 (hypoproct) triangular or rectangular; hypogynial valve present (shape of apical section and whether bare or setulose are regarded as important diagnostic characters to separate species) or absent; three spermathecae (2+1) blackish brown and spherical.