Betaburmesebuthus fleissneri Lourenço & Velten, 2016

XUAN, QIANG, CAI, CHEN-YANG & HUANG, DI-YING, 2023, Revision of palaeoburmesebuthid scorpions in mid-Cretaceous amber from northern Myanmar (Scorpiones: Buthoidea), Palaeoentomology 6 (1), pp. 64-101 : 79-83

publication ID	https://doi.org/ 10.11646/palaeoentomology.6.1.10
publication LSID	lsid:zoobank.org:pub:B822ED53-D559-4DD8-8803-3F2850A00445
DOI	https://doi.org/10.5281/zenodo.7757593
persistent identifier	https://treatment.plazi.org/id/F308E17F-435F-FFF0-FF6F-486DFD5C1E56
treatment provided by	Plazi
scientific name	Betaburmesebuthus fleissneri Lourenço & Velten, 2016
status

Treatment

( Figs 12–14 View FIGURE 12 View FIGURE 13 View FIGURE 14 )

Material. NIGP 200653, an adult male, part of right pedipalp missing.

Diagnosis (emended). This species can be clearly distinguished from other Betaburmesebuthus species by the following set of characters: 1) anterior margin of carapace with a single moderately marked median concavity ( Figs 13A View FIGURE 13 , 14A View FIGURE 14 ; Lourenço & Velten, 2016a: fig. 5); 2) spiracles oval to round ( Lourenço & Velten, 2016a: fig. 6); 3) pectines with 19–21 teeth ( Fig. 13B View FIGURE 13 ; Lourenço & Velten, 2016a); 4) vesicle pear-shaped and very long, and aculeus very long and strongly curved ( Figs 13B View FIGURE 13 , 14B View FIGURE 14 ); 5) dorsal patellar spur carina with four strong tubercles and femur with three to four internal strong tubercles ( Fig. 14C, D View FIGURE 14 ; Lourenço & Velten, 2016a: fig. 2); 6) Chela manus with two small tubercles on internal surface ( Lourenço & Velten, 2016a: fig. 10); 7) telotarsus covered by numerous ventrosubmedian setae ( Fig. 14E– G View FIGURE 14 , Lourenço & Velten, 2016a: fig. 8).

Description. Carapace. Covered by coarse granules; a pair of superciliary carinae well developed; median, posterior median and posterior furrows evident ( Figs 13A View FIGURE 13 , 14A View FIGURE 14 ); median eyes oval, large sized and separated by one ocular diameter ( Fig. 14A View FIGURE 14 ); PDMi posterodorsal to PLMa, ADMi mixed with coarse granules, not clear ( Fig. 14A View FIGURE 14 ).

Coxosternal region. Surface covered by impurity, not clear but obscure outline visible; genital operculum longitudinally divided, two valves rounded and full ( Fig. 13B View FIGURE 13 ).

Chelicerae. With numerous setae on internal surface ( Fig. 14A View FIGURE 14 ); cheliceral dentition not visible ( Fig. 14A View FIGURE 14 ).

Pedipalps. Femur with five carinae ( Figs 13E View FIGURE 13 , 14D View FIGURE 14 ): internomedian carina well-marked with several strong tubercles; dorsointernal carina crenulate; dorsoexternal, ventroexternal and ventrointernal carinae costate. Patella with seven carinae ( Figs 13E View FIGURE 13 , 14C View FIGURE 14 ): Dorsal Patellar Spur carina (DPSc) and Ventral Patellar Spur carina (VPSc) well-developed; dorsointernal, dorsomedian and ventrointernal carinae costate with several granules; ventroexternal and dorsoexternal carinae smooth and costate. Chela very slender (Cl/Cw = 7.67, Table 1 View TABLE 1 ).

Trichobothrial pattern ( Figs 13E View FIGURE 13 , 14C, D View FIGURE 14 ). Femur with 7 trichobothria observed, 4 dorsal, 1 internal and 2 external trichobothria, trichobothrium e 1 proximal to trichobothrium d 5. Patella with 8 trichobothria observed, including 1 dorsal, 1internal and 6 external trichobothria. Chela fixed finger with 2 dorsal trichobothria observed.

Legs. Prolateral and retrolateral pedal spurs present on all legs ( Fig. 14E–G View FIGURE 14 ). Ungues long and moderately curved, dactyl pointed ( Fig. 14E–G View FIGURE 14 ).

Pectines ( Fig. 13B View FIGURE 13 ). Lamellae not clear, peg sensillae very short and sensory area fully occupied distal end of teeth.

Mesosoma. Tergites covered by coarse granules ( Figs 12A View FIGURE 12 , 13A, D View FIGURE 13 ), median carina evident on posterior half of all tergites ( Fig. 12A View FIGURE 12 ); dorsolateral carinae present on tergites IV–VI ( Fig. 13D View FIGURE 13 ); tergite VII with five costate carinae ( Fig. 13D View FIGURE 13 ): paired dorsolateral and lateral carinae, and one axial carina on anterior half. Sternites covered by impurity, not clear.

Metasoma ( Fig. 13C, J View FIGURE 13 ). All segments longer than width and covered by macrosetae ventrally.Segments I–III with distinct depression dorsally; segments I and II with ten well marked carinae (paired ventromedian, ventrolateral, lateral, dorsolateral, and dorsal carinae); segments III and IV with eight well marked carinae (paired ventromedian, ventrolateral, dorsolateral, and dorsal carinae) and lateral carina reduced on segment III; segment V with five well marked carinae (single ventromedian carina, paired ventrolateral and dorsolateral carinae); dorsal carinae on segments I–IV obviously serrated.

Telson ( Figs13C View FIGURE 13 , 14B View FIGURE 14 ).Vesicle with a well-developed ventromedian carina, ventral surface covered by several macrosetae and subaculear setal pair significantly evident; one or two small subaculear tuberance present on terminal of ventromedian carina; base of aculeus moderately wide and shorter than half of vesicle; aculeus with several microsetae proximally and progressively darker towards distal.

Remarks. Lourenço & Velten (2016) regarded the serrated dorsal carinae of metasoma I–IV as a diagnostic character of Betaburmesebuthus fleissneri . However, we noticed that this character is shared by all members of Betaburmesebuthus and especially evident in mature individuals ( Figs 13C, J View FIGURE 13 , 24H View FIGURE 24 , 29C View FIGURE 29 ). Our adult specimen provides more reliable features than the immature holotype.

We remeasured the ratio of chela length and width from figure 2 in Lourenço & Velten (2016) and found it is about 7.50 in holotype rather than 6.82 in the original paper, close to the value in our specimen (7.67) .