Thesium scandens, Sond., DLN Thesium schaijesii, Lawalree, DLN Thesium schliebenii, Pilg., DLN Thesium schmitzii, Robyns & Lawalree, DLN Thesium schweinfurthii, Engl., DLN The scirpioides, A.W.Hill Thesium selagineum, A.DC. Thesium sertulariastrum, A.W. Hill Thesium setulosum, Robyns & Lawalree, DLN Thesium simplex, Velen., DLN Thesium sonderianum, Schltr., DLN Thesium sp. Thesium sp. Thesium spicatum, L. Thesium spinosum, L. Thesium spinulosum, A.DC Thesium stelleroides, Jaub. & Spach, DLN Thesium strictum, P. J. Bergius, DLN Thesium stuhlmannii, Engl., DLN Thesium subaphyllum, Engl., DLN Thesium subsimile, N. E. Br., DLN Thesium subsucculentum, (Kammer) J. C. Manning & F. Forest, DLN Thesium symoensii, Lawalree, DLN Thesium szowitsii, A.DC. Thesium tamariscinum, A.W.Hill Thesium tenuifolium, Saut. ex W. D. J. Koch, DLN Thesium tenuissimum, Hook. f., DLN Thesium tepuiense, Steyerm., DLN Thesium thamnus, Robyns & Lawalree, DLN Thesium translucens, A.W.Hill Thesium triflorum, L Thesium tuvense, I. M. Krasnoborov, DLN Thesium unyikense, Engl., DLN Thesium ussanguense, Engl., DLN Thesium utile, A.W.Hill Thesium vahrmeijeri, Brenan, DLN Thesium vimineum, Robyns & Lawalree, DLN Thesium virens, E. Mey. Thesium virgatum, Lam., DLN Thesium viride, A.W.Hill Thesium viridifolium, Levyns, DLN Thesium whyteanum, Rendle, DLN Thesium wilczekianum, Lawalree, DLN Thesium xerophyticum, A.W.Hill

García, Miguel A., Mucina, Ladislav & Nickrent, Daniel L., 2024, A tough nutlet to crack: Resolving the phylogeny of Thesium (Thesiaceae), the largest genus in Santalales, TAXON 73 (1), pp. 190-236 : 235-236

T. scandens Sond.,

DLN 4857, South Africa, R. Bayliss 1423 (NY), OP184353, –, – ; DLN 5399, South Africa, M. A. García & al. 5399 (MA, BH, NBG), OP184354, OP172424, OP172130 ; T. schaijesii Lawalrée,

DLN 4755, Democratic Republic of the Congo, M. Scheijes 2735 (BR), OP184355, OP172425, OP172131 ; T. schliebenii Pilg.,

DLN 4756, Zambia, H. Richards 5214 (BR), OP184356, OP172426, – ; DLN 4805, Democratic Republic of the Congo, M. Schaijes 1806 (BR), OP184357, OP172427, OP172132 ; T. schmitzii Robyns & Lawalrée,

DLN 4757, Democratic Republic of the Congo, F. Malaisse 13645 (BR), OP184358, –, – ; DLN 4758, Burundi, P. Auquier 4337 (BR), OP184359, OP172428, OP172133 ; T. schweinfurthii Engl.,

DLN 4759, Tanzania, R. Gereau & al. 4976 (BR), OP184360, OP172429, OP172134 ; DLN 4797, Uganda, P. Rwaburindore 3639 (BR), OP184361, –, – ; DLN 4824, Ethiopia, M. Gilbert & M. Thulin 619 (UPS), OP184362, OP172430, OP172135 ; T. scirpioides A. W.Hill ,

DLN 5233, Mozambique, D. Zunguze & C. Boane 96 (MO), OP184363, OP172431, – ; DLN 5292, South Africa, O. Hilliard & B. Burtt 18535 (MO), OP184364, OP172432, –; T. selagineum A.DC. ,

DLN 5378, South Africa, M. A. García & al. 5378 (MA, BH, NBG), OP184365, OP172433, OP172136 ; T. sertulariastrum A. W.Hill,

DLN 5425, South Africa, M. A. García & al. 5425 (MA, BH, NBG), OP184366, OP172434, OP172137 ; DLN 5511, South Africa, L. Mucina 100906/16 (MA), OP184367, OP172435, OP172138 ; T. setulosum Robyns & Lawalrée,

DLN 4760, Zambia, A. Angus 398 (BR), OP184368, –, – ; T. simplex Velen.,

DLN 5141, Moldova, A. Arvat s.n. (LE), OP184369, OP172436, – ; T. sonderianum Schltr.,

DLN 5392, South Africa, M. A. García & al. 5392 (MA, BH, NBG), OP184469, OP172517, OP172213; DLN 5394, South Africa, M. A. García & al. 5394 (MA, BH, NBG), OP184370, OP172437, OP172139 ; Thesium sp. ,

DLN 4393, South Africa, E. Cloete s.n. (MO), OP184371, OP172438, – ; Thesium sp. , DLN 4686, South Africa, P. Goetghebeur 4332 (BR), OP184372, OP172439, OP172140 ; Thesium sp. , DLN 4709, South Africa, G. Germishuizen 3389 (BR), OP184373, OP172440, OP172141 ; Thesium sp. , DLN 4710, Zimbabwe, C. Vanden Berghen 91 (BR), OP184374, OP172441, OP172142 ; Thesium sp. , DLN 4796, Burkina Faso, J. Lejoly 82/382 (BR), OP184375, OP172442, OP172143 ; Thesium sp. , DLN 4807, Zimbabwe, T. Baudesson & al. 471 (BR), OP184376, OP172443, OP172144 ; Thesium sp. , DLN 4818, Leshoto, O. Hedberg 82120 (UPS), OP184377, OP172444, OP172145 ; Thesium sp. , DLN 4819, Ethiopia, M. Thulin & A. Hunde 3975 (UPS), OP184378, OP172445, OP172146 ; Thesium sp. , DLN 4825, Ethiopia, M. Gilbert & al. 8287 (UPS), OP184379, OP172446, OP172147 ; Thesium sp. , DLN 5237, South Africa, K. Balkwill 7868 (MO), OP184380, OP172447, OP172148 ; Thesium sp. , DLN 5263, China, Y. Shu-Xiu 160 (MO), OP184381, OP172448, –; Thesium sp. , DLN 5300, South Africa, C. Bredenkamp 717 (MO), OP184382, OP172449, OP172149 ; Thesium sp. , DLN 5308, South Africa, K. Balkwill & Balkwill 4555 (MO), OP184383 (ITS clone A), OP184384 (ITS clone B), OP172450, – ; Thesium sp. , DLN 5309, South Africa, E. Oliver 10324 (MO), OP184385, OP172451, OP172150 ; Thesium sp. , DLN 5377, South Africa, M. A. García & al. 5377 (MA, BH, NBG), OP184386, OP172452, OP172151 ; Thesium sp. , DLN 5393, South Africa, M. A. García & al. 5393 (MA, BH, NBG), OP184387 (ITS clone A), OP184388 (ITS clone B), OP184389 (ITS clone C), OP172453, OP172152 ; Thesium sp. , DLN 5566, South Africa, E. Pienaar M336 (MA), OP184393, OP172457, OP172156 ; Thesium sp. , DLN 5567, South Africa, E.Pienaar T 65 (MA), OP184394, OP172458, – ; Thesium sp. , DLN 5574, South Africa, E. Pienaar M1184 (MA), OP184395, OP172459, OP172157 ; Thesium sp. sect. Frisea,

DLN 5384, South Africa, M. A. García & al. 5384 (MA, BH, NBG), OP184396, OP172460, OP172158 ; Thesium sp. sect. Frisea, DLN 5416, South Africa, M. A. García & al. 5416 (MA, BH, NBG), OP184397, OP172461, OP172159 ; Thesium sp. sect. Frisea, DLN 5422, South Africa, M. A. García & al. 5422 (MA, BH, NBG), OP184398, –, – ; Thesium sp. sect. Frisea, DLN 5426, South Africa, M. A. García & al. 5426 (MA, BH, NBG), OP184399, OP172462, OP172160 ; Thesium sp. sect. Frisea, DLN 5431, South Africa, M. A. García & al. 5431 (MA, BH, NBG), OP184400, OP172463, OP172161 ; Thesium sp. sect. Frisea, DLN 5437, South Africa, M. A. García & al. 5437 (MA, BH, NBG), OP184401, OP172464, OP172162 ; Thesium sp. sect. Frisea, DLN 5453, South Africa, M. A. García & al. 5453 (MA, BH, NBG), OP184402, OP172465, OP172163 ; Thesium sp. sect. Frisea, DLN 5460, South Africa, M. A. García & al. 5460 (MA, BH, NBG), OP184403, OP172466, OP172164 ; Thesium sp. sect. Frisea, DLN 5482, South Africa, M. A. García & al. 5482 (MA, BH, NBG), OP184404, OP172467, OP172165 ; Thesium sp. sect. Frisea, DLN 5494, South Africa, M. A. García & al. 5494 (MA, BH, NBG), OP184405, –, –; Thesium sp. sect. Frisea, DLN 5523, South Africa, L. Mucina 190206/3 (MA), OP184406, OP172468, OP172166 ; Thesium sp. sect. Frisea, DLN 5577, South Africa, E. Pienaar M87 (MA), OP184407, –, – ; T. spicatum L. ,

DLN 4095, South Africa, D. Nickrent & al. 4095 (BH), OP184408, –, – ; DLN 4761, South Africa, R. Bayliss 7748 (BR), OP184409, –, – ; T. spinosum L. f.,

DLN 4762, South Africa, E. Coppejans 1476 (BR), OP184410, OP172469, OP172167 ; DLN 5513, South Africa, L. Mucina 090306/07 (MA), OP184411, OP172470, OP172168 ; T. spinulosum A.DC .,

DLN 5295, South Africa, E. Esterhuysen 34460 (MO), OP184412, –, –; DLN 5424, South Africa, M. A. García & al. 5424 (MA, BH, NBG), OP184413, KP318978, OP172169 ; DLN 5498, South Africa, M. A. García & al. 5498 (MA, BH, NBG), OP184414, OP172471, OP172170 ; T. stelleroides Jaub. & Spach,

DLN 4848, Türkiye, F. Muñoz Garmedia & al. 4525 (MA), KP318963, KP318973, OP172171 ; T. strictum P.J.Bergius,

DLN 4092, South Africa, D. Nickrent & al. 4092 (BH), OP184415, –, – ; DLN 4763, South Africa, P. Bohnen 9003 (BR), OP184416, OP172472, OP172172; DLN 4764, South Africa, E. Coppejans 459 (BR), OP184417, OP172473, OP172173 ; DLN 4811, South Africa, H. Breyne 5414 (BR), OP184418, OP172474, OP172174 ; DLN 5505, South Africa, L. Mucina 021106/7 (MA), OP184419, OP172475, OP172175 ; DLN 5508, South Africa, L. Mucina 171205/02 (MA), OP184420, OP172476, OP172176 ; DLN 5515, South Africa, L. Mucina 090206/27 (MA), OP184421, OP172477, OP172177 ; DLN 5516, South Africa, L. Mucina 090206/10 (MA), OP184422, OP172478, OP172178; DLN 5526, South Africa, L. Mucina 211006/51 (MA), OP184423, OP172479, OP172179 ; DLN 5568, South Africa, E. Pienaar M646 (MA), OP184424, OP172480, OP172180 ; T. stuhlmannii Engl.,

DLN 4765, Tanzania, R. Gereau & C. Kayombo 4664 (BR), OP184425, OP172481, OP172181 ; DLN 4766, Tanzania, J. Lovett & C. Kayombo 3402 (BR), OP184426, OP172482, OP172182 ; T. subaphyllum Engl.,

DLN 4767, Tanzania, R. Gereau & C. Kayombo 3899 (BR), OP184427, OP172483, OP172183 ; T. subsimile N.E.Br.,

DLN 5254, South Africa, K. Balkwill & M-J. Balkwill 11245 (MO), OP184428, OP172484, OP172184 ; T. subsucculentum (Kämmer) J.C.Manning & F.Forest,

DLN 4374, Spain: Canary Islands, A. Santos Guerra s.n. (MA), KP318962 (ITS clone A), OP184429 (ITS clone B), OP184430 (ITS clone C), KP318972, OP172185 ; T. symoensii Lawalrée,

DLN 4768, Democratic Republic of the Congo, S. Lisowski & al. 4276 (BR), OP184431, OP172485, – ; T. szowitsii A.DC. ,

DLN 4434, Iran, K. Rechinger 41333 (MO), OP184432, OP172486, OP172186 ; DLN 5191, Armenia, C. Navarro & al. CN-5602 (MA), OP184433, OP172487, OP172187 ; T. tamariscinum A. W.Hill ,

DLN 4769, Tanzania, J. Lovett & C. Kayombo 3401 (BR), OP184434, OP172488, OP172188 ; T. tenuifolium Saut. ex W.D.J.Koch,

DLN 5142, Russian Federation, N. N. Tzvelev 95 (LE), OP184435, OP172489, OP172189 ; T. tenuissimum Hook.f.,

DLN 4772, Cameroon, W. Meijer 15407 (BR), OP184436, OP172490, OP172190 ; T. tepuiense Steyerm.,

DLN 4858, Guyana, B. Maguire & D. Fanshawe 32543 (NY), OP184437, OP172491, – ; T. thamnus Robyns & Lawalrée,

DLN 4773, Democratic Republic of the Congo, A. Schmitz 8124 (BR), OP184438, OP172492, OP172191 ; DLN 4792, Democratic Republic of the Congo, M. Schaijes 2025 (BR), OP184439, OP172493, OP172192 ; T. translucens A. W.Hill ,

DLN 5500, South Africa, M. A. García & al. 5500 (MA, BH, NBG), OP184440, OP172494, OP172193 ; T. triflorum L .f.,

DLN 4438, Tanzania, J. A. Mlangwa & W. Kindeketta 717 (MO), OP184441, OP172495, OP172194 ; DLN 4679, South Africa, R. Bayliss 7876 (BR), OP184442, –, – ; DLN 4774, Tanzania, D. Goyder & al. 3850 (BR), OP184443, OP172496, OP172195 ; DLN 5291, South Africa, P. Phillipson & T. Dold 4188 (MO), OP184444, –, – ; T. tuvense I.M.Krasnoborov,

DLN 5172, Mongolia, R.V. Kamelin & al. 1476 (LE), OP184445, OP172497, OP172196 ; T. unyikense Engl.,

DLN 4392, Malawi, I.F. LaCroix 4756 (MO), OP184446, OP172498, OP172197 ; DLN 4776, Tanzania, R. Brummitt & al. 18192 (BR), OP184447, OP172499, OP172198 ; T. ussanguense Engl.,

DLN 4777, Zimbabwe, P. Bamps & al. 691 (BR), OP184448, OP172500, – ; DLN 4778, Malawi, P. Tyrer 821 (BR), OP184449, OP172501, OP172199 ; T. utile A. W.Hill ,

DLN 4689, South Africa, G. Germishuizen 3380 (BR), OP184450, OP172502, OP172200 ; DLN 5298, South Africa, N. Kroon 11710 (MO), OP184451, OP172503, OP172201 ; T. vahrmeijeri Brenan,

DLN 4779, Mozambique, J. Koning 7869 (BR), OP184452, –, – ; T. vimineum Robyns & Lawalrée,

DLN 4783, Democratic Republic of the Congo, S. Lisowski & al. 7720 (BR), OP184453, OP172504, OP172202 ; T. virens E.Mey. ex A.DC .,

DLN 5299, South Africa, O. Hilliard & B. Burtt 9061 (MO), OP184454, OP172505, OP172203 ; T. virgatum Lam.,

DLN 5293, South Africa, P. Goldblatt 3737 (MO), OP184455, –, – ; DLN 5448, South Africa, M. A. García & al. 5448 (MA, BH, NBG), OP184456, OP172506, OP172204 ; DLN 5481, South Africa, M. A. García & al. 5481 (MA, BH, NBG), OP184457, OP172507, OP172205 ; DLN 5594, South Africa, L. Mucina 130808/07 (MA), OP184458, OP172508, – ; T. viride A. W.Hill ,

DLN 4770, Cameroon, A. Jacques-Georges 27654 (BR), OP184459, OP172509, OP172206 ; DLN 4784, Togo, J. Brunel 708 (BR), OP184460, OP172510, – ; DLN 4785, Nigeria, R. Meikle 1198 (BR), OP184461, OP172511, OP172207 ; T. viridifolium Levyns,

DLN 4388, South Africa, E.R. Orchard 571 (MO), OP184462, –, –; DLN 5442, South Africa, M. A.García & al. 5442 (MA, BH, NBG), OP184463, OP172512, OP172208 ; DLN 5451, South Africa, M. A. García & al.5451 (MA, BH, NBG), OP184464, –, – ; DLN 5456, South Africa, M. A. García & al. 5456 (MA, BH, NBG), OP184465, OP172513, OP172209 ; T. whyteanum Rendle,

DLN 4787, Malawi, J. Chapman & I. Patel 5779 (BR), OP184466, OP172514, OP172210 ; T. wilczekianum Lawalrée,

DLN 4789, Democratic Republic of the Congo, F. Malaisse 8407 (BR), OP184467, OP172515, OP172211 ; T. xerophyticum A. W.Hill ,

DLN 5302, Namibia, H. Merxmüller & J. Giess 32566 (MO), OP184468, OP172516, OP172212 .