Enhypnon punctatum Carter

Turco, Federica, Ślipiński, Adam & Lambkin, Christine L., 2013, Enhypnon Carter: a taxonomic revision of an endemic Australian genus of ground-dwelling beetles (Coleoptera: Zopheridae), Zootaxa 3681 (4), pp. 371-394 : 385-391

Enhypnon punctatum Carter

( Figs 2 f, 4f, 5f, 6f, 7e)

Enhypnon punctatum Carter, 1927: 231 .

Types. Holotype. Tasmania, Mount Wellington, H.H.D. Griffith, coll. Carter [Ga: 42.883S 147.233E] (AMSA, K.67211).

Other specimens examined. Tasmania: Mt. Wellington, Lea [Ga: 42.883S 147.233E] (1, SAMA); Sandspit River, 42.09S 147.55E, rainforest, Tube 149, pyrethrin knockdown Atherospermum, 2.vi.1989, J. Diggle & P. Greenslade (1, ANIC); Scotts Peak Road, DDRFB 18C, Pitfall trapping, FT44134, 1.iii.03, D.Driscoll, 43.0163 S 146.2254 E (1, FTas); South Picton River area, South Picton River area ( Baker), Pitfall trapping, FT44088, 2.ii.08, S. Baker, 43.1797 S 146.654 E (1, FTas); Warra: Manuka Road, SSTBIG100, Pitfall trapping, FT9567, 18.ii.00, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG100, Pitfall trapping, FT28547, 17.xii.99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT8577, 16.ii.99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG254, Pitfall trapping, FT9519, 24.i.00, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT9460, 17.xii.99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT8667, 19.iii.99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT8674, 19.iii.99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT9460, 17.xii.99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT8308, 17.xi.98, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, Yee site WR, WRLET 2, Emergence trapping from log of Eucalyptus obliqua , FT29927, 9.x.01, M. Yee, 43.0935 S 146.713 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, WRLDOG3.3, Emergence trapping from log of Eucalyptus obliqua , FT27654, 7.v.03, R.Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG4.4, Emergence trapping from log of Eucalyptus obliqua , FT13954, 12.iii.02, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDRG1.3, Emergence trapping from log of Eucalyptus obliqua , FT13422, 18.x.01, R. Bashford, 43.07 S 146.67 E, Regrowth, Eucalyptus obliqua (1, FTas); same locality, Yee site WR, WRLET 2, Emergence trapping from log of Eucalyptus obliqua , FT29925, 28.ii.02, M. Yee, 43.0935 S 146.713 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, Yee site WR, WRLET 2, Emergence trapping from log of Eucalyptus obliqua , FT29920, 15.xi.00, M. Yee, 43.0935 S 146.713 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, Yee site M, MSET 3, Emergence trapping from log of Eucalyptus obliqua , FT29298, 28.ii.02, M. Yee, 43.0933 S 146.6442 E, 110, Small log, Eucalyptus obliqua (1, FTas); same locality, Yee site M, MLET 1, Emergence trapping from log of Eucalyptus obliqua , FT29228, 28.ii.02, M. Yee, 43.0933 S 146.6442 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, Yee site M, MLET 3, Emergence trapping from log of Eucalyptus obliqua , FT29247, 10.i.02, M. Yee, 43.0933 S 146.6442 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, WRLDRG5.1, Emergence trapping from log of Eucalyptus obliqua , FT31496, 17.ii.06, R. Bashford, 43.07 S 146.67 E, Regrowth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG6.4, Emergence trapping from log of Eucalyptus obliqua , FT32602, 19.ii.07, R.

Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG1.2, Emergence trapping from log of Eucalyptus obliqua , FT30874, 25.i.05, R.Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG2.2, Emergence trapping from log of Eucalyptus obliqua , FT31304, 16.xi.05, R.Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDRG4.1, Emergence trapping from log of Eucalyptus obliqua , FT31419, 5.i.06, R. Bashford, 43.07 S 146.67 E, Regrowth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG4.3, Emergence trapping from log of Eucalyptus obliqua , FT31318, 16.xi.05, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG6.4, Emergence trapping from log of Eucalyptus obliqua , FT32514, 22.i.07, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG2.1, Emergence trapping from log of Eucalyptus obliqua , FT30852, 21.xii.04, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG1.4, Emergence trapping from log of Eucalyptus obliqua , FT31750, 22.v.06, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WR011B ( Baker), Pitfall trapping, FT44082, 1.vii.01, S. Baker, 43.05 S 146.70 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT9579, 18.ii.00, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG170, Pitfall trapping, FT9506, 24.i.00, R. Bashford, 43.07 S 146.67 E (1, FTas); Warra-Weld alt transect, A03, WR0300PMT, Pitfall trapping, FT5752, 18.xii.01, N. Doran & R. Bashford, 43.0347 S 146.6945 E, 315 m (1, FTas); same locality, A02, WR0200PLO, Pitfall trapping, FT5927, 27.ii.02, N. Doran & R. Bashford, 43.0321 S 146.7006 E, 190 m (1, FTas); same locality, A05, WR0500PLO, Pitfall trapping, FT5857, 22.i.02, N. Doran & R. Bashford, 43.0374 S 146.6883 E, 535 m (1, FTas); same locality, A05, WR0500PLO, Pitfall trapping, FT133, 30.iii.01, N. Doran & R. Bashford, 43.0374 S 146.6883 E, 535 m (1, FTas); same locality, D07, WD0700P45U, Pitfall trapping, FT64, 26.ii.01, N. Doran & R. Bashford, 42.9996 S 146.6154 E, 739 m (1, FTas); Warra: Warra Rd, WR011B ( Baker), Pitfall trapping, FT44073, 1.ii.01, S. Baker, 43.05 S 146.70 E (1, FTas); same locality, Pitfall trapping, FT44083, 1.xi.01, S. Baker, 43.05 S 146.70 E (1, FTas); Tarraleah: Hornes Dam area, HornesPF8.4, Pitfall trapping, FT22747, 16.xi.92, R. Brereton, 42.274 S 146.3641 E, 740, Logged, Eucalyptus delegatensis forest (1, FTas).

Diagnosis. Among the species with raised tubercles on elytra, Enhypnon punctatum is distinct in having a strongly compact and convex body and distinctly raised tubercles on elytra and pronotum

Redescription. Body length: 3.45 – 3.95 mm; body maximum width (elytra): 1.7 – 2.1 mm.

Body: compact, distinctly convex and comparatively stout; colour dark brown with lighter spots on elytra ( Figs 2 f, 4f); vestiture consisting of recumbent yellowish setae, very dense and thick, longer and filiform around antennomeres III – XI.

Head: surface coarsely sub-rugose; maximum width at eyes (0.98 – 1.13 mm), abruptly constricted behind eyes; frons widely concave between eyes; frontal ridge relatively sinuate in frontal view, thicker in front of eyes above antennal insertion; clypeus anteriorly slightly curved, not distinctly separated from frons, relatively wide (ratio clypeus width/head max. width: 0.49 – 0.51) and with obtusely rounded lateral angles; eyes small, very strongly protruding and coarsely faceted; antenna 11-segmented with a loose and narrow 2-segmented club; antennomere I thick and elongate, almost twice as long as wide; II distinctly elongate (about three times as long as wide), thinner than I and longer than III; II – VII elongate and progressively shorter; VIII – X cupuliform and increasing in size; X about 1.5 times as wide as IX; XI about as wide as X and slightly pointed. Last maxillary palpomere flat, short and wide, less than 1.5 times as long as wide (see Fig. 6 f).

Pronotum: widest at anterior third, only weakly explanate and gently constricted posteriorly; surface coarsely sub-rugose with large tubercles in anterior half of disc ( Figs 2 f, 4f); margins obsolete and not visible from above, barely traceable in anterior half in lateral view.

Leg: protibia flattened, distinctly larger than meso- and metatibia and strongly concave on the internal side; tarsomeres not transverse.

Elytra: very distinct tubercles on intervals 3 (four), 5 (four) and 7 (three), one small tubercle on interval 9; second tubercle from base very low on intervals 3 and 5 ( Figs 2 f, 4f).

Distribution. Tasmania ( Fig. 7 e).

Enhypnon simplex Turco and Ś lipi ń ski sp. n. ( Figs 3 a, 4g, 5g, 6g, 7f)

Types. Holotype. Tasmania, Hartz Mts. NP, Hartz Rd, 740 m, 8 – 10.ii.80, Eucalyptus - Nothofagus berlesed from leaf litter, A. Newton & M. Thayer [Ga: 43.233S 146.767E] (ANIC).

Paratypes. same data as Holotype (2, ANIC).

Other specimens examined. Tasmania: Scotts Peak Road, 42.5738S 146.223E, 1.iii.03, DDRFB 3A, Pitfall trapping, FT44095, D. Driscoll (1, FTas).

Diagnosis. Among the species with low tubercles on elytra, E. simplex is similar to E. laticeps in having lateral angles of clypeus acute and pronotal lateral margins obsolete. It can be distinguished by the frons weakly concave, frontal ridge only slightly sinuate and pronotum convex, without a medial depressed area.

Description. Body length: 2.15 – 2.3 mm; body maximum width (elytra): 1.1 – 1.15 mm. Body: small, compact and distinctly convex; colour uniformly light brown; vestiture consiting of sparse, fine and short recumbent setae, longer around antennomeres III – XI.

Head: surface coarsely sub-rugose; maximum width at eyes (0.68 – 0.70 mm), abruptly constricted behind eyes; frons weakly concave between eyes; frontal ridge slightly sinuate in frontal view, thicker in front of eyes above antennal insertion; clypeus narrow (ratio clypeus width/head max. width: 0.43 – 0.44), distinctly separated and with acute lateral angles; eyes small, protruding and coarsely faceted; antenna 11-segmented with a loose and weakly developed 2-segmented club; antennomere I thick and about twice as long as wide and about 1.5 times as long as II; I – III progressively thinner; III as long as II; IV – VIII shorter; VIII – X cupuliform; X about 1.5 times as wide as IX; XI weakly wider than X, rounded and only slightly pointed. Last maxillary palpomere flat and distinctly wide and securiform (see Fig. 6 g).

Pronotum: widest at anterior third, very weakly explanate anteriorly and constricted posteriorly; surface uniformly coarsely sub-rugose, with no medial depressed area or tubercles; margins obsolete and not visible from above.

Leg: simple; tarsomeres weakly transverse.

Elytra: not costate, with very wide and distinct punctures ( Figs 3 a, 4g) and two almost obsolete subapical tubercles on interval 3.

Etymology. The name simplex is a Latin neuter adjective ( simplex = simple) and it refers to the simple dorsum that characterises this species compared to other congeners.

Distribution. Tasmania ( Fig. 7 f).

Enhypnon squamosum Turco and Ś lipi ń ski sp. n. ( Figs 3 b, 4h, 5h, 6h, 7g)

Types. Holotype. Australia, Tasmania: track off Mt. Barrow Rd, 780 m, 15 – 17.ii.1980, Nothofagus , pyrethrin-fogging tree ferns, A. Newton & M. Thayer [Ga: 41.367S 147.417E] (ANIC).

Paratypes. Tasmania: Same data as holotype (1, ANIC); Lottah [Ga: 41.21S 148.017E] (1, SAMA); 4 km SE Weldborough, 450 m, 12 – 14.ii.80, Nothofagus , pyrethrin fogging Nothofagus cunninghami bark, A. Newton & M. Thayer [GE: 41.209S 147.948E] (2, ANIC); W side L. St. Clair, 750 m, 25 – 29.i.80, Nothofagus , pyrethrin-fogging tree ferns, A. Newton & M. Thayer [GE: 42.055S 146.140E] (5, ANIC); Lake St. Clair, 42.06S 146.10E, 750 m, 25 – 27.i.1980, Pyrethrum spray tree ferns, J. Lawrence & T. Weir (3, ANIC); Mt. Barrow Rd, 890 m, 15 – 17.ii.1980, Nothofagus berlesed from leaf litter, A. Newton & M. Thayer [Ga: 41.367S 147.417E] (1, ANIC); 4,4 km SE Weldborough, 460 m, 12 – 14.ii.1980, Nothofagus , forest, A. Newton & M. Thayer [GE: 41.210S 147.954E] (2, ANIC); Savage R. Pipeline Rd, 41.30S 145.20E, 19.iv.1989, Tube 72, pyrethrin knockdown Nothofagus, J. Diggle (1, ANIC); same locality, 20.iv.1989, Tube 17, pyrethrin knockdown. Nothofagus, P. Greenslade (1, ANIC); Mt. Victoria, 41.20S 147.50E, 900 m, 23.xi.1989, pyrethrin knockdown tree 1, Tube 297, R. Coy (1, ANIC); same locality, 25.xi.1989, pyrethrin knockdown tree 2, Tube 239, H. Mitchell (2, ANIC); same locality, 23.xi.1989, pyrethrin knockdown tree 2, Tube 296, R. Coy (1, ANIC); same locality, 23.xi.1989, pyrethrin knockdown tree 3, Tube 298, R. Coy (2, ANIC); Mt. Michael, 41.11S 148.00E, 740 m, 26.xi.1989, pyrethrin knockdown tree 1, Tube 187, R. Coy (1, ANIC); same locality, 27.xi.1989, pyrethrin knockdown tree 2, Tube 182, R. Coy (1, ANIC); same locality, 28.xi.1989, pyrethrin knockdown tree 1, Tube 176, R. Coy (1, ANIC); St. Columba Falls, Berlesate Anic 912, 16.xii.1981, G. Bornemissza [Ga: 41.317S 147.917E] (1, ANIC).

Notes on types. The specimen from Lottah (SAMA) bears an old cotype label and an identification label by Carter as Enhypnon horridum . Nonetheless, this specimen certainly belongs to the new species E. squamosum and it is not part of the original type series of E. horridum , which consists of three specimens collected on Cradle Mountain by Lea.

Other specimens examined. Tasmania: Ben Ridge Rd., 5.2 km E Telopea Rd., 41.21S 147.40E, 870 m, 16.i – 1.ii.1993, A. Newton and M. Thayer, ex window-trap Nothofagus cunninghamii rainforest (1, ANIC); Mt Michael, 41.1833S 148.0000E, FT38038, Knockdown fogging of canopy of Nothofagus cunninghamii , 21/02/90, R.Coy, P. Lillywhite & A.Yen (6, FTas); same locality, FT38037, Knockdown fogging of canopy of Nothofagus cunninghamii , 20/02/90, R.Coy, P. Lillywhite & A.Yen (9, FTas); same locality, 41.11S 148.00E, 740 m, 27.xi.1989, pyrethrin knockdown tree 1 – 2, Tube (ranging from 182 to 187), R. Coy (6, ANIC); same locality, 28.xi.1989, suction, Tube 245, R. Coy (1, ANIC); same locality, 11.vi.1990, moss on tree trunks, Tube 320, A.T. W. (1, ANIC); Mt. Victoria, 41.20S 147.50E, 900 m, 25.xi.1989, pyrethrin knockdown tree 1 – 3, Tube (ranging from 237 to 305), H. Mitchell (or R. Coy) (46, ANIC); same locality, 22 – 28.xi.1989, yellow pans (x5), Tube 169, H. Mitchell & R. Coy (1, ANIC); same locality, 29.xi.1989, litter, Tube 289, R. Coy (1, ANIC); 4,4 km SE Weldborough, 460 m, 12 – 14.ii.1980, Nothofagus , forest, A. Newton & M. Thayer [GE: 41.210S 147.954E] (1, ANIC); Savage R. Pipeline Rd, 41.30S 145.20E, 19.iv.1989, Tube 72, pyrethrin knockdown Nothofagus, J. Diggle (1, ANIC); Schulhofs Rd., 8.7 km NE Upper Blessington, 41.26S 147.36E, 870 m, 16.i – 1.ii.1993, A. Newton & M. Thayer, ex window-trap, Euc. delegatensis and Euc. obliqua forest (1, ANIC); Simons Rd., non-wilt area, 41.21S 147.31E, 8.iii.1989, leaf litter, Tube 273, P. Greenslade (4, ANIC); Simons Rd., wilt area, 41.21S 147.31E, 8.iii.1989, leaf litter, Tube 272, J. Diggle (1, ANIC); Near Weldborough, 41°10’S 147°08’E, 20 – 25.ii.1993, Nothofagus forest, litter, J. Trueman et al., site A (1, ANIC); 40 km SW Smithton, 4.iii.1977, FMHD #77 – 189, litter in Crown of Dicksonia antarctica, J. Kethley [GE: 41.047S 144.734E] (1, FMNH); Cradle Mtn. Nat. Pk, Waldheim For. (3500ft), 3.ii.1977, FMHD #77 – 109, litter of unburned Nothofagus gunnei, J. Kethley [Ga: 41.633S 145.933E] (1, FMNH); 2 km S. Revision Bell, 9.xii.1986, D. Burckhardt (1, ANIC).

Diagnosis. Among the species with raised tubercles on elytra, Enhypnon squamosum is distinct in having deeply denticulate pronotal margins and strongly squamiform dorsal vestiture. This species is quite variable in size and colour, from uniformly yellowish-brown to distinctly bicoloured with patterned elytra.

Description. Body length: 1.9 – 3.0 mm; body maximum width (elytra): 1.0 – 1.7 mm.

Body: small, compact and distinctly convex; colour from uniformly light brown to distinctly bicoloured with patterned elytra and lateral part of pronotum; vestiture consiting of dense, short and strongly squamiform setae ( Figs 3 b, 4h); long and filiform around antennomeres III – XI.

Head: surface coarsely sub-rugose; maximum width at eyes (0.60 – 0.88 mm), abruptly constricted behind eyes; frons broadly concave between eyes; frontal ridge weakly sinuate in frontal view, thicker in front of eyes above antennal insertion; clypeus anteriorly curved, relatively wide (ratio clypeus width/head max. width: 0.49 – 0.58) and with obtuse lateral angles; eyes strongly protruding and coarsely faceted; antenna 11-segmented with a loose and weakly developed 2-segmented club; antennomere I relatively thin and elongate (about twice as long as wide); II more than twice as long as wide; II – VIII elongate and progressively shorter; IX more elongate and wider than VIII; X cupuliform and relatively narrow (less than 1.5 times as wide as IX); XI only slightly wider than X, sub-oval, weakly elongate and pointed. Last maxillary palpomere flat, distinctly wide and sub-triangular (see Fig. 6 h).

Pronotum: explanate and abruptly constricted posteriorly; surface sub-rugose with two admedian shallow depressions on pronotal disc; margin denticulate, usually with four/five teeth or spines.

Leg: simple; tarsomeres weakly transverse.

Elytra: distinctly tuberculate along intervals 3, 5, 7, and 9; two larger sub-apical tubercles on interval 3, clearly visible in lateral view ( Figs 3 b, 4h).

Etymology. The name squamosum is a Latin neuter adjective, squamosum (= that bears scales) and it refers to the distinctive vestiture, consisting of dense and strongly squamiform setae.

Distribution. Tasmania ( Fig. 7 g).

Enhypnon tuberculatum Turco and Ś lipi ń ski sp. n. ( Figs 3 d, 4j, 5j, 6j, 7h)

Types. Holotype. Australia, Tasmania: Hartz Mts. NP, Hartz Rd, 740 m, 8 – 10.ii.80, Eucalyptus - Nothofagus , berlesed from leaf litter, Newton & Thayer [Ga: 43.233S 146.767E] (ANIC).

Paratypes. Tasmania: Same data as holotype (1, ANIC); 22 mi. SE Smithton, Savage Riv, 5.iii.1977, FMHD #77 – 193, Nothofagus cunninghami litter at base of Nothofagus cunninghami log, J. Kethley [GE: 41.103S 145.371E] (1, FMNH); 35 km SW Smithton, Christmas Hills, 27.ii.1977, FMHD #77 – 177 N. cunninghami litter around log, J. Kethley [GE: 40.901S 144.997E] (1, FMNH); Corinna W Darlingtons,. iii.57 [Ga: 41.65S 145.067E] (1, ANIC); Mt. Murchison, 41.50S 145.37E, 21.iv.1989, rainf. Tube 129 moss on dead log, J. Diggle & H. Mithell (2, ANIC); same locality, 21.iv.1989, rainf. Tube 132, leaf litter, J. Diggle (1, ANIC); Riveaux Creek, 43.11S 146.40E, 20.xii.1988, Tube 58, rainforest litter, P. Greenslade (1, ANIC); Strahan, 25.xii.1953, C. Oke (1, MVMA T-17401); Willie's Saddle, 9 km W Geeveston, 310 m, 8 – 10.ii.1980, berlese from leaf litter, Eucalyptus forest, A. Newton & M. Thayer [GE: 43.165S 146.810E] (1, ANIC).

Other specimens examined. Tasmania: Mt. Field NP, Lake Dobson Rd, 710 m, 30.i – 5.ii.1980, berlesed from leaf litter, Nothofagus - Eucalyptus forest, A. Newton & M. Thayer [GE: 42.681S 146.656E] (1, ANIC); Waratah, A. Lea [Ga: 41.45S 145.533E] (1, SAMA); South Weld Road Spur 1 Yee site S, SSET 2, FT29820, Emergence trapping from log of Eucalyptus obliqua , 30.i.02, M.Yee, 43.0826S 146.7223E, 110 m (1, FTas); Riveaux Road Yee site R, RLET 2, FT29665, Emergence trapping from log of Eucalyptus obliqua , 13.xii.00, M.Yee, 43.1033S 146.728E, 110 m (1, FTas); same locality, RLET 2, FT29670, Emergence trapping from log of Eucalyptus obliqua , 17.i.02, M.Yee, 43.1033S 146.728E, 110 m (1, FTas); Edwards Road, Yee site A (E), ELET 2, FT29056, Emergence trapping from log of Eucalyptus obliqua , 13.xii.00, M.Yee, 43.0918S 146.7473E, 110 m (1, FTas); same locality, Yee site A (E), ELET 2, FT29057, Emergence trapping from log of Eucalyptus obliqua , 31.i.01, M.Yee, 43.0918S 146.7473E, 110 m (1, FTas); same locality, Yee site A (E), ELET 3, FT29066, Emergence trapping from log of Eucalyptus obliqua , 4.v.01, M.Yee, 43.0918S 146.7473E, 110 m (1, FTas); same locality, Yee site A (E), ESET 2, FT29095, Emergence trapping from log of Eucalyptus obliqua , 15.v.02, M.Yee, 43.0918S 146.7473E, 110 m (2, FTas); same locality, Yee site A (E), ELET 1, FT29052, Emergence trapping from log of Eucalyptus obliqua , 31.i.01, M.Yee, 43.0918S 146.7473E, 110 m (1, FTas); West Picton Road Yee site PO1, PO1LET2, FT29350, Emergence trapping from log of Eucalyptus obliqua , 13.xii.00, M.Yee, 43.1667S 146.6869E, 217 m (1, FTas); Warra: Manuka Road, Yee site M, MLET 1, FT29225, Emergence trapping from log of Eucalyptus obliqua , 29.v.01, M.Yee, 43.0933S 146.6442E, 110 m (1, FTas); same locality, Yee site M, MSET 3, FT29297, Emergence trapping from log of Eucalyptus obliqua , 1.v.02, M.Yee, 43.0933S 146.6442E, 110 m (1, FTas); same locality, SSTBIG170, FT8398, Pitfall trapping, 16.xii.98, R.Bashford, 43.07S 146.67E (1, FTas); same locality, WRLDOG4.5, FT13620, Emergence trapping from log of Eucalyptus obliqua , 18.xii.01, R.Bashford, 43.07S 146.67E (1, FTas); same locality, WRLDOG6.5, FT27321, Emergence trapping from log of Eucalyptus obliqua , 15.i.03, R.Bashford, 43.07S 146.67E (1, FTas); same locality, WRLDOG1.2, FT27074, Emergence trapping from log of Eucalyptus obliqua , 12.xi.02, R.Bashford, 43.07S 146.67E (1, FTas); same locality, WRLDRG1.4, FT31547, Emergence trapping from log of Eucalyptus obliqua , 7.iii.06, R.Bashford, 43.07S 146.67E (1, FTas); same locality, WRLDOG6.4, FT33308, Emergence trapping from log of Eucalyptus obliqua , 18.xii.07, R.Bashford, 43.07S 146.67E (1, FTas); same locality, KH Tree 21, KH5244, Emergence trapping from cut billets of Eucalyptus obliqua , 1.i.04, K.Harrison, 43.07S 146.67E (1, FTas); same locality, WRLDOG6.4, FT32688, Emergence trapping from log of Eucalyptus obliqua , 22.iii.07, R.Bashford, 43.07S 146.67E (1, FTas); same locality, SSTCON059, FT7811, Pitfall trapping, 17.vi.98, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTBIG106, FT8748, Pitfall trapping, 16.iv.99, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTBIG106, FT9377, Pitfall trapping, 19.xi.99, R. Bashford, 43.07S 146.67E (1, FTas); same locality, WRLDOG6.4, FT27414, Emergence trapping from log of Eucalyptus obliqua , 11.ii.03, R.Bashford, 43.07S 146.67E (1, FTas); same locality, SSTCON147, FT8960, Pitfall trapping, 17.vi.99, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTCON059, FT9397, Pitfall trapping, 19.xi.99, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTCON059, FT35220, Pitfall trapping, 8.ii.05, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTMID280, FT7674, Pitfall trapping, 22.iv.98, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTWES120, FT36439, Pitfall trapping, 7.vii.05, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTSMA254, FT8347, Pitfall trapping, 17.xi.98, R. Bashford, 43.07S 146.67E (1, FTas); same locality, SSTBIG170, FT9396, Pitfall trapping, 19.xi.99, R. Bashford, 43.07S 146.67E (1, FTas); same locality, Yee site WR, WRLET 2, FT29921, Emergence trapping from log of Eucalyptus obliqua , 31.i.01, M.Yee, 43.0935S 146.713E, 110 m (1, FTas); same locality, Yee site WR, WRLET 2, FT29924, Emergence trapping from log of Eucalyptus obliqua , 13.xii.00, M.Yee, 43.0935S 146.713E, 110 m (1, FTas); same locality, WRLDOG6.3, FT13522, Emergence trapping from log of Eucalyptus obliqua , 17.xi.01, R.Bashford, 43.07S 146.67E (1, FTas); same locality, SSTSMA254, FT7227, Pitfall trapping, 7.xi.97, R. Bashford, 43.07S 146.67E (1, FTas); Warra, end of Manuka Rd., 24 km WNW Geeveston, 43°5’27’’S 146°39’9’’E, 120 m, 5.iii.2008, T. Weir & C. Leeman, ANIC Berlesate 3748, Rainforest with Eucalyptus sp., sifted litter and mossy logs (3, QMBA ethanol coll.); Warra Road, Yee site W, WSET 1, FT30013, Emergence trapping from log of Eucalyptus obliqua , 31.i.01, M.Yee, 43.0872S 146.7186E, 110 m (1, FTas); same locality, Yee site W, WSET 1, FT30014, Emergence trapping from log of Eucalyptus obliqua , 29.v.01, M.Yee, 43.0872S 146.7186E, 110 m (1, FTas); Hartz Mts., 575 m, 12.xi.1983, Bornemissza [Ga: 43.233S 146.767E] (3, HNHM); 7 km NEbyE of Mt Wedge, 42.49S 146.22E, 550 m, 3.ii.1980, Berlesate ANIC 670 litter Nothofagus cunninghamii & Anodopetalum biglandulosum, J. Lawrence & T. Weir (1, ANIC); Hartz Mts., Huon river, Tahune bridge, 150 m, 7.xii.86, Burckhardt (1, ANIC); Strahan, 25.xii.53, C. Oke (1, ANIC); Waratah, Lea (1, ANIC). Victoria: Wilson's Prom N.P. Lilly Pilly Tr, 14.v.1978, log litter and fungi, S. & J. Peck (1, ANIC).

Diagnosis. Among the species with raised tubercles on elytra, Enhypnon tuberculatum is distinct in having the pronotum tuberculate ( Figs 3 d, 4j), pronotal margins not explanate or denticulate, tarsomeres not lobed below and protibia not flattened.

Description. Body length: 2.25 – 2.90 mm; body maximum width (elytra): 1.10 – 1.45 mm.

Body: small, compact and distinctly convex, usually covered with a thin layer of dirt; colour uniformly dark brown; vestiture consisting of fine, short and dense recumbent setae (denser on tubercles); long and filiform around antennomeres III – XI.

Head: surface coarsely sub-rugose; maximum width at eyes (0.60 – 0.73 mm), abruptly constricted behind eyes; frons broadly concave between eyes; frontal ridge weakly sinuate in frontal view, thicker in front of eyes above antennal insertion; clypeus anteriorly straight, very short and relatively narrow (ratio clypeus width/head max. width: 0.45 – 0.47) and with obtuse lateral angles; eyes small, weakly protruding and coarsely faceted; antenna 11- segmented with a loose and weakly developed 2-segmented club; antennomere I thick, about twice as long as wide; II about as long as I and rounded; III elongate and thin, about twice as long as wide; I – III progressively decreasing in thickness; IV – IX uniformly shorter and rounded; X cupuliform and almost twice as large as IX; XI slightly narrower than X, rounded and only very weakly pointed. Last maxillary palpomere flat, elongate, parallel-sided and slightly narrower at apex (see Fig. 6 j).

Pronotum: not distinctly explanate, maximum width at anterior third and abruptly constricted posteriorly; surface coarsely sub-rugose with two sets of admedian tubercles, one small pair on pronotal disc and one more prominent pair on anterior third reaching the anterior margin of pronotum ( Figs 3 d, 4j); lateral margin well defined in anterior half and visible from above, crenulated.

Leg: simple; tarsomeres not transverse.

Elytra: intervals 3, 5 and 7 with distinct small tubercles; basal tubercles on intervals 3 and 5 usually lower and longer ( Figs 3 d, 4j).

Etymology. The name tuberculatum is a Latin neuter adjective, tuberculatum (= that bears tubercles), and it refers to the presence of distinctive tubercles on both pronotum and elytra.

Distribution. Tasmania and Victoria ( Fig. 7 h).

Figures

FIGURE 2. Dorsal habitus: a. Enhypnon cordicollis; b. E. costatum; c. E. horridum; d. E. kosciuszko; e. E. laticeps; f. E. punctatum. Scale bars = 0.5 mm.

FIGURE 3. Dorsal habitus: a. Enhypnon simplex; b. E. squamosum; c. E. latitarsis; d. E. tuberculatum. Scale bars = 0.5 mm.

FIGURE 6. Maxillary palps: a. Enhypnon cordicollis; b. E. costatum; c. E. horridum; d. E. kosciuszko; e. E. laticeps; f. E. punctatum; g. E. simplex; h. E. squamosum; i. E. latitarsis; j. E. tuberculatum. Scale bars = 0.1 mm.

FIGURE 7. Distribution map of Enhypnon species endemic to Tasmania.