MYTILIDAE Rafinesque, 1815

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 699-701

Family MYTILIDAE Rafinesque, 1815

Callitrichus – Deshayes, 1830a-EncyMeth: 164, ex Poli ms. Incorrect subsequent spelling of Callitriche Poli, 1791 , now regarded as a taxon inquirendum and listed under Gibbomodiola Sacco, 1898 . acutangulus , Mytilus –Deshayes, 1830-DescrCoqFoss: 274-275, 20, pl. 40, figs 1-2; 1836-HistNatAnim2: 53; 1843-HistNatAnim3: 23; 1861-DescrCoqFoss: 27. — Type localities: Valmondois & Senlis. — Type age: Eocene (Bartonian). Perna acutangula (Deshayes, 1830) ( Glibert & Van de Poel 1965b: 71; Le Renard & Pacaud 1995: 82). — Type species (OD) of Admytilus Berezovsky, 2015 . — Current status: Admytilus acutangulus (Deshayes, 1830) ( Berezovsky 2015b: 578-579, fig. 1).

acuminata , Modiola –Deshayes, 1830-DescrCoqFoss: 262-263, 20, pl. 40, figs 9-11, non Sedgwick, 1829;1832-EncyMeth: 562-563, as Mytilus ; 1836-HistNatAnim2: 31; 1843-HistNatAnim3: 16, both as Modiola ; 1861-DescrCoqFoss: 22. — Type locality: Paris (old district of Vaugirard). — Type age: Eocene (Lutetian). — Current status: Brachidontes (Brachidontes) jansseni Le Renard, 1994 ( Le Renard 1994: 39), replacement name ( Pacaud 2008: 82).

ambigua , Modiola – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 10-12; 1861-DescrCoqFoss: 21. — Type localities: Le Fayel, Bouconvilliers & Valmondois. — Type age: Eocene (Bartonian).— Current status: Brachidontes (Brachidontes) ambiguus (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 74; Le Renard & Pacaud 1995: 82).

analoga , Modiola – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 27-30;1861-DescrCoqFoss: 16, as Modiola (Modiolaris) . — Type localities: Étréchy & Jeurre. — Type age: Oligocene (Rupelian). — Current status: Musculus analogus (Deshayes, 1858) ( Lozouet & Maestrati 2012a: 446-447, fig. 156: 3-4).

angularis , Modiola – Deshayes, 1830-DescrCoqFoss: 260-261, 21, pl. 41, figs 4-5;1832-EncyMeth: 563, as Mytilus ; 1836-HistNatAnim2: 31; 1843-HistNatAnim3: 16, both as Modiola ; 1861-DescrCoqFoss: 13, as Modiola (Modiolaria) . — Type locality: Noailles. — Type age: Paleocene (Thanetian). Musculus (Semimodiola) angularis (Deshayes, 1830) ( Glibert & Van de Poel 1965b: 82) . — Current status: Semimodiola angularis (Deshayes, 1830) ( Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

argentina , Modiola –Deshayes, 1824a: 256-257, 258, pl. 15, fig. 15; 1830-DescrCoqFoss: 269-270, 21, pl. 42, figs 1-3; 1832-EncyMeth: 571, as Mytilus ; 1836-HistNatAnim2: 32; 1843-HistNatAnim3: 16, both as Modiola ; 1861-DescrCoqFoss: 19, as Modiola (Lithodomus) . — Type localities: Valmondois & near Dax – material from the latter locality more probably Botula aturensis ( Cossmann & Peyrot, 1914) or Botula subcordata (d’Orbigny, 1852) . — Type age: Miocene (Pacaud herein). The Valmondois material Eocene (Bartonian). — Current status: Botula argentina (Deshayes, 1824) ( Glibert & Van de Poel 1965b: 83; Le Renard & Pacaud 1995: 83).

attenuata , Modiola – Deshayes,1836-HistNatAnim2: 28;1843-HistNatAnim3: 14-15. — Type locality: Peru & Chile. — Type age: Recent. — Type species (OD) of Labis Dall, 1916. — Current status: Lithophaga (Labis) attenuata (Deshayes, 1836) ( Huber 2010a: 120, as Leiosolenus ; Coan & Valentich-Scott 2012: 138-139).

bernayi , Modiola (Modiolaris) – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 13-16; 1861-DescrCoqFoss: 13-14. — Type locality: Valmondois. — Type age: Eocene (Bartonian). — Current status: Musculus (Planimodiola) bernayi (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 83; Le Renard & Pacaud 1995: 83).

biochii , Mytilus – Deshayes, 1866b: 335, 338, pl. 7, fig. 6. — Type locality: Argenteuil. — Type age: Eocene (Bartonian). — Current status: Brachidontes (Hormomya) biochii (Deshayes, 1866) ( Le Renard & Pacaud 1995: 82) .

brevis , Modiola – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 17- 19; 1861-DescrCoqFoss: 24. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Current status: Arcoperna brevis (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 77; Le Renard & Pacaud 1995: 83; Pacaud 2008: 81). capillaris , Modiola – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 22-24; 1861-DescrCoqFoss: 23. — Type localities: Ver-sur-Launette & Ermenonville. — Type age: Eocene (Bartonian). — Current status: Arcoperna capillaris (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 77; Le Renard & Pacaud 1995: 83).

crenella , Modiola – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 1-3; 1861-DescrCoqFoss: 20-21. — Type localities: Mouy & Parnes. — Type age: Eocene (Lutetian). — Current status: Brachidontes (Brachidontes) crenella (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 74; Le Renard & Pacaud 1995: 82; Pacaud 2008: 82).

cucullata , Crenella – Deshayes, 1858-DescrCoqFoss: 76, pl. 76, figs 10-12; 1861-DescrCoqFoss: 7. — Type localities: Thiverval-Grignon & Montmirail. — Type age: Eocene (Lutetian). — Current status: Crenella (Hippagus) cucullata Deshayes, 1858 ( Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

cultellus , Modiola – Deshayes, 1839d: 359; 1840: 2, pl. 13. — Type locality: Kamchatka. — Type age: Recent. Musculus cultellus (Deshayes, 1839) ( Kantor & Sysoev 2005: 317) . — Current status: synonym of Musculus svecicus (Fabricius, 1788) [ Modiola ] ( Huber 2010a: 110, 543; 2015: chapter C on CD).

delicatula , Modiola – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 25-26; 1861-DescrCoqFoss: 18-19, as Modiola (Lithodomus) . — Type locality: Morigny-Champigny. — Type age: Oligocene (Rupelian). — Current status: Modiolus delicatulus Deshayes, 1858 ( Lozouet & Maestrati 2012b: 22).

depressus , Mytilus (Septifer) –Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 25-27; 1861-DescrCoqFoss: 31-32, non G. Fischer, 1830. — Type localities: Parnes, Chaumont-en-Vexin & Chaussy. — Type age: Eocene (Lutetian). Septifer serratus [Melleville, 1843] depressus (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 77) . — Current status: Septifer pissarroi Le Renard, 1994 ( Le Renard 1994: 39), replacement name ( Pacaud 2008: 81, as “1861”).

difficilis , Modiola – Deshayes, 1863: 23-24, pl. 3, figs 22-24. — Type locality: La Réunion. — Type age: Recent. Modiolus difficilis (Deshayes, 1863) ( Poppe 2010: 510, pl. 946, figs 10-11, erroneously attributed to “Kuroda & Habe, 1950”). Senior secondary homonym of Volsella difficilis Kuroda & Habe, 1950 , which was replaced by Modiolus kurilensis F. R. Bernard, 1983 . — Current status: Gregariella difficilis (Deshayes, 1863) ( Huber 2010a: 545; Poppe 2018: 137).

dolabrata , Modiola – Deshayes, 1858-DescrCoqFoss: 71, pl. 71, figs 20-21; 1861-DescrCoqFoss: 26-27. — Type localities: Brimont, Châlons-sur-Vesle & Jonchery-sur-Vesle. — Type age: Paleocene (Thanetian). — Current status: Modiolus dolabratus Deshayes, 1858 ( Glibert & Van de Poel 1965b: 73; Le Renard & Pacaud 1995: 82; Leroy et al. 2014: 27, pl. 8, figs 1-4).

dutemplei , Mytilus – Deshayes, 1858-DescrCoqFoss: 71, pl. 71, fig. 22; 1861-DescrCoqFoss: 30-31. — Type localities: Mont-Bernon (Epernay) & Cramant. — Type age: Eocene (Ypresian). — Current status: Brachidontes (Hormomya) dutemplei (Deshayes, 1858) ( Le Renard & Pacaud 1995: 82) .

elegans , Crenella – Deshayes, 1858-DescrCoqFoss: 76, pl. 76, figs 6-9; 1861-DescrCoqFoss: 6. — Type localities: Thiverval-Grignon, Parnes, Damery, Mouchy-le-Châtel & Chaussy. — Type age: Eocene (Lutetian). — Current status: Crenella (Crenella) elegans Deshayes, 1858 ( Glibert & Van de Poel 1965b: 78; Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

hastata , Modiola – Deshayes, 1830-DescrCoqFoss: 261-262, 19, pl. 38, figs 13-14; 1832-EncyMeth: 563, as Mytilus ; 1836-HistNatAnim2: 31; 1843-HistNatAnim3: 16, both as Modiola ; 1861-DescrCoqFoss: 13, as Modiola (Modiolaria) . — Type localities: Abbecourt. — Type age: Paleocene (Thanetian); Chaumont-en-Vexin, Eocene (Lu - tetian). — Type species (OD) of Semimodiola Cossmann, 1887 . Musculus (Semimodiola) hastatus (Deshayes, 1830) ( Glibert & Van de Poel 1965b: 82) . — Current status: Semimodiola hastata (Deshayes, 1830) ( Le Renard & Pacaud 1995: 83; Pacaud 2008: 81; Nevesskaja et al., 2013: 164, fig. 47-13).

heberti , Mytilus – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 20- 22; 1861-DescrCoqFoss: 32-33; 1865-DescrCoqFoss: 666 [last two references as Mytilus denticulatus Lamarck, 1805 ] and thus regarded as a synonym thereof. — Type localities: Longjumeau, Jeurre, Etréchy. — Type age: Oligocene (Rupelian). Cossmann & Peyrot (1914: 35) listed it as Septifer heberti (Deshayes, 1858) . — Current status: synonym of Septifer denticulatus ( Lamarck, 1805) ( Lozouet & Maestrati 2012a: 246, 247, fig. 155: 11-14.

incrassatus, Mytilus – Deshayes,1832-EncyMeth: 558, non Sheppard, 1821, nec three post-1832 uses of this name. Based on Deshayes, 1830-DescrCoqFoss: 20, pl. 40, figs 1-2, which he had described as Mytilus acutangulus ; 1861-DescrCoqFoss: 27, as synonym of M. acutangulus Deshayes, 1830 . — Type localities: Valmondois & Senlis. — Type age: Eocene (Bartonian). — Current status: synonym of Admytilus acutangulus (Deshayes, 1830) ( Berezovsky 2015b: 578-579, fig. 1, without synonymy).

interjecta, Modiola (Modiolaris) – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 16-18; 1861-DescrCoqFoss: 17. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). — Current status: Musculus (Planimodiola) interjectus (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 83; Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

laevigatus , Mytilus – Deshayes, 1858-DescrCoqFoss: 71, pl. 71, figs 17-19; 1861-DescrCoqFoss: 28-29, as Mytilus levigatus , non Mytilus laevigatus W. Wood, 1828 . — Type localities: Bazoches, Vauxbuin & Sinceny. — Type age: Eocene (Ypresian). Cowell Bay, Isle of Wight, England. — Current status: Perna ferussaci Le Renard, 1994 ( Le Renard 1994: 39), replacement name.

levesquei , Mytilus – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 4-5; 1861-DescrCoqFoss: 30. — Type localities: Retheuil & Cuise-la-Motte. — Type age: Eocene (Ypresian).— Current status: Brachidontes (Hormomya) levesquei (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 75; Le Renard & Pacaud 1995: 82).

microptera , Modiola – Deshayes, 1836-HistNatAnim2: 27-28; 1843-HistNatAnim3: 14. — Type locality: “Coromandel” [ New Zealand]. — Type age: Recent. — Current status: synonym of the Asian Modiolus modulaides (Röding, 1798) [ Musculus ] ( Huber 2010a: 554).

papyracea , Modiola – Deshayes, 1824a: 257, 258, pl. 15, fig. 16; 1830-DescrCoqFoss: 270-271, 21, pl. 41, figs 9-11; 1832-EncyMeth: 572, as Mytilus . 1836-HistNatAnim2: 33; 1843-HistNatAnim3: 16, both as Modiola ; 1861-DescrCoqFoss: 20, as Modiola (Lithodomus) . — Type locality: Valmondois. — Type age: Eocene (Bartonian). — Current status: Lithophaga papyracea (Deshayes, 1824) ( Le Renard & Pacaud 1995: 83) .

parasitica, Modiola – Deshayes, 1863: 24-25, pl. 4, figs 1-3, non Eudes-Deslongchamps, 1838. — Type locality: La Réunion. — Type age: Recent. NHMUK 20080540, 3 syntypes. — Current status: Adula kleemanni M. Huber, 2010a ( Huber 2010a: 553), replacement name.

pectiniformis , Modiola – Deshayes, 1830 DescrCoqFoss: 263, 20, pl. 39, figs 14-16; 1832-EncyMeth: 564, as Mytilus ; 1836-HistNatAnim2: 31-32; 1843-HistNatAnim3: 16, both as Modiola ; 1861-DescrCoqFoss: 14, as Modiola (Modiolaris) . — Type locality: Houdan. — Type age: Eocene (Lutetian). — Current status: Musculus (Gregariella) pectiniformis (Deshayes, 1830) ( Glibert & Van de Poel 1965b: 80; Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

piethei , Modiola – Deshayes, 1858-DescrCoqFoss: 75, as Modiola pihetei , pl. 75, figs 14-15; 1861-DescrCoqFoss: 14-15, as Modiola (Modiolaris) piethei . — It is obvious from the 1861 text that the plate explanation had a misspelling, in that the species was from the collection of Mr. Piethe of Crouy; however, this does not meet the test of ICZN Code Art. 32.5. — Type localities: Crouy, Acy-en-Multien, Caumont & La Ferté-sous-Jouarre. — Type age: Eocene (Bartonian). — Current status: Musculus (Gregariella) pihetei (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 80; Le Renard & Pacaud 1995: 83; Pacaud 2008: 80).

profunda , Modiola –Deshayes, 1830-DescrCoqFoss: 264, 21, pl. 41, figs 12-14. 1832 -EncyMeth: 564, as Mytilus ; 1836-HistNatAnim2: 32; 1843-HistNatAnim3: 16, both as Modiola ; 1861-DescrCoqFoss: 23. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Current status: Arcoperna profunda (Deshayes, 1830) ( Glibert & Van de Poel 1965b: 78, as “1832”; Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

radiolata , Modiola – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 10-13; 1861-DescrCoqFoss: 22. — Type localities: Coeuvres-et-Valsery, Aizy-Jouy & Laon. — Type age: Eocene (Ypresian). — Current status: Arcoperna radiolata (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 78, as A. “ pradiolata ”; Le Renard & Pacaud 1995: 83).

rigaultii , Mytilus –Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 23- 24; 1861-DescrCoqFoss: 29. — Type localities: Caumont, Jaignes, Attainville, Montagny-en-Vexin & Le Guépelle. — Type age: Eocene (Bartonian). — Current status: Brachidontes (Hormomya) rigaultii (Deshayes, 1858) ( Glibert & Van de Poel 1965b: 76; Le Renard & Pacaud 1995: 82; Pacaud 2008: 82).

semilaevigata , Modiola – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 19-20; 1861-DescrCoqFoss: 26, as Modiola “ semilevigata ”. — Type locality: Damery. — Type age: Eocene (Lutetian). — Current status: Modiolus semilaevigata Deshayes, 1858 ( Le Renard & Pacaud 1995: 82; Pacaud 2008: 81).

seminuda , Modiola – Deshayes, 1830-DescrCoqFoss: 264-265, 20, pl. 39, figs 20-22; 1832-EncyMeth: 569-570, as Mytilus ; 1836-HistNatAnim2: 32; 1843-HistNatAnim3: 16, both as as Modiola ; 1861-DescrCoqFoss: 12-13, as Modiola (Modiolaria) . — Type locality: La Chapelle-en-Serval. — Type age: Eocene (Bartonian). — Current status: Musculus (Skarlatella) seminudus (Deshayes, 1830) ( Glibert & Van de Poel 1965b: 82; Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

spathulata , Modiola – Deshayes, 1830-DescrCoqFoss: 259, 20, pl. 39, figs 11-13; 1832-EncyMeth: 566-567, as Mytilus ; 1836-HistNatAnim2: 30; 1843-HistNatAnim3: 15-16, both as Modiola ; 1861-DescrCoqFoss: 17, as Modiola (Modiolaris) . — Type localities: Parnes & Les Groux. — Type age: Eocene (Lutetian). — Current status: Musculus (Planimodiola) spathulatus (Deshayes, 1858) ( Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

striatina , Crenella – Deshayes, 1858-DescrCoqFoss: 76, pl. 76, figs 3-5;1861-DescrCoqFoss: 6-7. — Type localities: Chaumont-en-Vexin, Parnes & Chaussy. — Type age: Eocene (Lutetian). — Current status: Crenella (Crenella) striatina Deshayes, 1858 ( Glibert & Van de Poel 1965b: 79; Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

subangulata , Modiola – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 21; 1861-DescrCoqFoss: 25-26. — Type locality: Liancourt-Saint-Pierre. — Type age: Eocene (Lutetian). — Current status: Modiolula subangulata (Deshayes, 1858 ( Glibert & Van de Poel 1965b: 77; Le Renard & Pacaud 1995: 82; Pacaud 2008: 81).

subrostrata , Modiola – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 4-6; 1861-DescrCoqFoss: 15-16, as Modiola (Modiolaris) . — Type locality: Hermonville, Fleury-la-Rivière, Boursault & Damery. — Type age: Eocene (Lutetian). Synonym of Musculus (Planimodiola) sulcatus ( Lamarck, 1805) [ Modiola ] ( Glibert & Van de Poel 1965a: 83). — Current status: Musculus (Planimodiola) subrostratus (Deshayes, 1858) ( Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

tenera , Modiola – Deshayes, 1858-DescrCoqFoss: 74, pl. 74, figs 7-9; 1861-DescrCoqFoss: 24. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Current status: Arcoperna tenera (Deshayes, 1858) ( Le Renard & Pacaud 1995: 83; Pacaud 2008: 81).

tenuis , Mytilus – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 1-3; 1861-DescrCoqFoss: 28, non J. S. Schröter, 1803. — Type localities: Châlons-sur-Vesle & Jonchery-sur-Vesle. — Type age: Paleocene (Thanetian). — Current status: synonym of Perna antiqua (Melleville, 1843) [ Dreissena ] (Pacaud 2007: 55).

vaudini , Mytilus (Septifer) – Deshayes, 1858-DescrCoqFoss: 75, pl. 75, figs 6-9; 1861-DescrCoqFoss: 31. — Type localities: Laon & Mons-en-Laonnais. — Type age: Eocene (Ypresian). — Current status: synonym of Septifer serratus (Melleville, 1843) [ Dreissena ] ( Glibert & Van de Poel 1965b: 77).

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amygdaloides , Modiola – Leymerie, 1841: 342, nomen nudum; 1842: 8, 25, pl. 6, fig. 4a-b, ex Deshayes ms. — Type localities: Marolles-sous-Lignières & Chenay. — Type age: Cretaceous (Hauterivian). — Current status: Lithophaga amygdaloides (Leymerie, 1842) ( Masse et al. 2022: 5, as “Deshayes”).

rimosa , Perna – Glibert & Van de Poel (1965b: 72) listed this species as “ Perna rimosa (Deshayes, 1830) ”, but it was originally described as Mytilus rimosus Lamarck, 1805 .

ropan , Mytilus – Deshayes, 1836-HistNatAnim2: 27, ex Adanson ms. 1843-HistNatAnim3: 14. As indicated by Huber (2015: chapter 5 on CD), this synonymous species name was first made available as Fistulana ropan Blainville, 1820 , ex Adanson ms. — Current status: synonym of Leiosolenus aristatus (Dillwyn, 1817) [ Mytilus ] ( Coan & Valentich-Scott 2012: 140).

simplex , Modiola – Leymerie, 1841: 336, nomen nudum; 1842: 8, 26, pl. 7, fig. 8, ex Deshayes ms, non (Passy, 1832) [ Mytilus ]. — Type locality: Ville-sur-Terre. — Type age: early Cretaceous (Hauterivian). Mytilus subsimplex d’Orbigny, 1850b (d’Orbigny 1850b: 81, no. 346), replacement name. — Current status: Musculus subsimplex (d’Orbigny, 1850) ( Calzada & Urquiola 1999: 294) .