GASTROCHAENIDAE J. E. Gray, 1840

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 795-796

Family GASTROCHAENIDAE J. E. Gray, 1840

Choena – Deshayes, 1830a-TraitMeth: 236. Incorrect subsequent spelling of Chaena Philipsson, 1788 .

agglutinans , Chaena – Deshayes, 1855c: 330. — Type locality: “Zebu” [Cebu] Island, Philippine Islands; Cuming coll. — Type age: Recent. Eufistulana agglutinans (Deshayes, 1855) ( Huber 2010a: 279) . — Current status: shell and tube morphometrics would be needed to know if this is distinct from Eufistulana mumia (Spengler, 1783) (J. G. Carter, pers. comm., August 2020).

angusta , Fistulana – Deshayes, 1824a: 251, 258, pl. 15, fig. 3. 1824-DescrCoqFoss: 16, 1, pl. 1, figs 11-15; 1824-DictClass6: 523; 1830b-EncyMeth: 143;1856-DescrCoqFoss: 2, pl. 2, fig. 11. — Type locality: Valmondois. — Type age: Eocene (Bartonian). — Current status: Carterochaena angusta (Deshayes, 1824) (J. G. Carter, pers. comm., August 2020).

apertissima , Gastrochaena – Deshayes, 1855c: 326. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978086, holotype. — Type species (OD) of Spenglerichaena Carter, 2011 . — Current status: Spenglerichaena apertissima (Deshayes, 1855) ( Huber 2010a: 279, as Spengleria ; Carter, in Carter et al. 2011: 23-24, fig. 2, holotype).

coarctata , Gastrochaena – Deshayes, 1856-DescrCoqFoss: 2, pl. 2, figs 12-14; 1857-DescrCoqFoss: 100. — Type locality: Damery. — Type age: Eocene (Lutetian). Gastrochaena coarctata Deshayes, 1856 ( Glibert & Van de Poel 1966a: 22). — Current status: Gastrochaena (Rocellaria) coarctata Deshayes, 1856 ( Le Renard & Pacaud 1995: 67; Pacaud 2008: 89). Either Rocellaria or Dufoichaena (J. G. Carter, pers. comm., August 2020).

contorta , Fistulana – Deshayes, 1824a: 251-252, 258, pl. 15, fig. 4; 1824-DescrCoqFoss: 16-17, 1, pl. 1, figs 24-25, 27.1824-DictClass6: 523; 1830b-EncyMeth: 143. — Type locality: Valmondois. — Type age: Eocene (Bartonian). — Current status: synonym of Rocellaria ampullaria ( Lamarck, 1806) ( Cossmann 1886a: 22; Le Renard & Pacaud 1995: 67).

cucullata , Gastrochaena – Deshayes, 1855c: 329. — Type locality: West Indies; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978100, 4 syntypes. — Current status: synonym of Lamychaena hians (Gmelin, 1791) [ Pholas ] ( Huber 2010a: 672).

defrancii , Gastrochaena – Deshayes, 1856-DescrCoqFoss: 2, pl. 2, figs 15-18; 1857-DescrCoqFoss: 102. — Type localities: Brasles, Les Groux, Lévemont. — Type age: Eocene (Lutetian). Gastrochaena defrancii Deshayes, 1856 ( Glibert & Van de Poel 1966a: 22). — Current status: Gastrochaena (Rocellaria) defrancii Deshayes, 1856 ( Le Renard & Pacaud 1995: 67; Pacaud 2008: 89; J. G. Carter, pers. comm., August 2020).

denticulata , Gastrochaena – Deshayes, 1855c: 327-328. — Type locality: Colombia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978092, holotype. — Current status: the tropical eastern Pacific Gastrochaena denticulata Deshayes, 1855 ( Huber 2010a: 671; Coan & Valentich-Scott 2012: 948).

difficilis , Gastrochaena – Deshayes, 1855c: 328. — Type locality: “Western India ” [ Saint Thomas, Virgin Islands]; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978093, 3 syntypes. — Current status: Gastrochaena difficilis Deshayes, 1855 ( Carter et al. 2008: 107, 110, figs 3.6, 6.6-9, the latter syntypes from MNHN [uncatalogued]; Huber 2010a: 671).

elongata , Fistulana – Deshayes, 1824-DescrCoqFoss: 15, 2, pl. 4, figs 17-19; 1824-DictClass6: 523; 1830b-EncyMeth: 141. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). — Type material: MNHN.F.A26030, holotype. — Current status: Eufistulana elongata (Deshayes, 1824) ( Glibert & Van de Poel 1966a: 23; Glibert, 1980: 43; Le Renard & Pacaud 1995: 67; Pacaud 2008: 89).

folini, Gastrochaena – Deshayes in Folin, 1867: 49-50 [11], pl. 1, figs 6-11. — Type locality: Pearl Islands, Panama. — Type age: Recent. — Type material: MNHN-IM-2000-38633, 4 syntypes (according to J. Carter, lot also includes Lamychaena truncata , largest specimen; NHMUK unnumbered, 2 syntypes, MacAndrew coll. are L. truncata ). — Current status: synonym of Lamychaena truncata (G. B. Sowerby I, 1834) [ Gastrochaena ] ( Coan & Valentich-Scott 2012: 950).

gigantea , Fistulana – Deshayes, 1830b-EncyMeth: 142; 1839-TraitElem-1(2): 29, 32, 35-36, 2, pl. 2, figs 6-8, as Gastrochaena . — Type locality: “Indies”. — Type age: Recent. — Type material: NHMUK 1978079, syntypes. Gastrochaena gigantea (Deshayes, 1830) ( Oliver 1992: 199, pl. 45; Poppe 2011: 178, pl. 1084, fig. 5). — Current status: more likely synonym of Gastrochaena cuneiformis Spengler, 1783 ( Carter et al. 2008: 106; Huber 2010a: 278, 671; Lutaenko et al. 2021: 239).

gigas , Gastrochaena – Deshayes, 1840-RegAnim: pl. 116, fig. 3, 3a-b. — Type locality: none given. — Type age: Recent. Perhaps intended to be Gastrochaena gigantea Deshayes, 1830 , made available in the Encyclopédie Methodique, but he remembered the name incorrectly; the figures are very similar. — Current status: synonym of Gastrochaena cuneiformis Spengler, 1783 (J. G. Carter, pers. comm., August 2020).

grandis , Chaena – Deshayes, 1855c: 330. — Type locality: “Zebu” [Cebu] Island, Philippine Islands; Cuming coll. — Type age: Recent. — Current status: Eufistulana grandis (Deshayes, 1855) ( Qi 2004: 319-320, pl. 174, fig. B; Huber 2010a: 279, 671, 673; Poppe 2011: 178, pl. 1084, fig. 1; Okutani 2017: 1231, pl. 526, fig. 7). Population-level shell morphometrics would be needed to distinguish this from Eufistulana mumia (Spengler, 1783) (J. G. Carter, pers. comm., August 2020).

humilis , Gastrochaena – Deshayes, 1855c: 327. — Type locality: “Zebu” [Cebu] Island, Philippine Islands; Cuming coll. & West Indies. — Type age: Recent. — Type material: NHMUK 1978089, syntype from Philippine Islands; NHMUK 1840.10.8.41, syntype from St. Vincent, Caribbean. Gastrochaena humilis Deshayes, 1855 ( Huber 2010a: 671-672). — Current status:synonym of Gastrochaena difficilis Deshayes, 1855 (J. G. Carter, pers. comm., August 2020).

impressa, Gastrochaena – Deshayes, 1855c: 327. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978091, 1 syntype pair & 1 valve. — Current status: possible synonym of the Philippines Islands Spenglerichaena apertissima (Deshayes, 1855) [ Gastrochaena ], but population-level shell morphometric and anatomical analysis needed (J. G. Carter, pers. comm., August 2020).

indistincta , Gastrochaena – Deshayes, 1855c: 328. — Type locality: Singapore; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978096, 2 syntype pairs & one valve. Gastrochaena , but taxon inquirendum (J. G. Carter, pers. comm., August 2020). — Current status: synonym of Gastrochaena cuneiformis Spengler, 1783 ( Lutaenko et al. 2021: 239).

interrupta , Gastrochaena – Deshayes, 1855c: 329. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978098, 2 syntypes. — Type species (M) of Dufoichaena Jousseaume , in E. Lamy, 1925. — Current status: not synonym of Dufoichaena dentifera (Dufo, 1840) [ Gastrochaena ], which has much shorter, less cylindrical siphons and longer intersiphonal pallial septum (J. G. Carter, pers. comm., August 2020), thus Dufoichaena interrupta (Deshayes, 1855) .

intersecta , Gastrochaena – Deshayes, 1855c: 327. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978090, 3 syntypes. Nomen dubium ( Huber, 2010a: 672) . — Current status: the syntype illustrated by G. B. Sowerby, II, 1878, is a Lamychaena ; the non-figured types are Gastrochaena , but lack of locality data prohibits meaningful comparisons with other type specimens of Lamychaena (J. G. Carter, pers. comm., August 2020).

laevigata , Gastrochaena – Deshayes, 1855c: 326-327. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978088, 2 syntypes. — Current status: nomen dubium ( Huber, 2010a: 672) ; juvenile Lamychaena or Dufoichaena (J. G. Carter, pers. comm., August 2020).

lamellosa , Gastrochaena – Deshayes, 1855c: 328. — Type locality: “Zebu” [Cebu] Island, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978094, 3 syntypes & 1 broken valve. — Current status: synonym of Gastrochaena cuneiformis Spengler, 1783 ( Huber 2015: chapter 5 on CD; Lutaenko et al. 2021: 239).

macroschisma , Gastrochaena – Deshayes, 1855c: 326. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978087, 3 syntypes. — Current status: the Indo-Pacific Gastrochaena macroschisma Deshayes, 1855 ( Carter et al. 2008: 105, 107, 110, figs 2.4, 6, 3.4; Poppe 2011: 178, pl. 1084, fig. 3).

philippinensis , Gastrochaena – Deshayes, 1855c: 328. — Type locality: “Zebu” [Cebu] Island, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978095, 3 syntypes. — Current status: Gastrochaena philippinensis Deshayes, 1855 ( Huber 2010a: 278).

plicatilis , Gastrochaena – Deshayes, 1855c: 329. — Type locality: “Zebu” [Cebu] Island, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978101, 3 syntypes. — Current status: synonym of Spengleria mytiloides ( Lamarck, 1818) [ Gastrochaena ], or Spengleria plicatilis (Deshayes, 1855) ( Poppe, 2011: 178, pl. 1084, figs 7-8). Poppe (2018: 104) concluded that this was a junior synonym of Spengleria mytiloides ( Lamarck, 1818) . Population-level shell morphometrics and genetic analyses needed to characterize intra- and interspecific variation in shell sculpture in the two nominal species (J. G. Carter, pers. comm., August 2020).

provigny , Fistulana – Deshayes, 1824a: 251, 257, pl. 15, fig. 2; 1824-DescrCoqFoss: 17, 1, as F. provignyi , pl. 1, figs 16, 19, 22; 1824-DictClass6: 523, as F. provigny ; 1830b-EncyMeth: 141- 142. — Type locality: Valmondois. — Type age: Eocene (Bartonian). Gastrochaena provignyi (Deshayes, 1824) ( Glibert & Van de Poel 1966a: 23) . — Current status: Gastrochaena (Rocellaria) provignyi (Deshayes, 1824) ( Glibert 1980: 43-44; Le Renard & Pacaud 1995: 67; Pacaud 2008: 89; Carter et al. 2008: 110, 112, fig. 8.1-5).

pupina , Gastrochaena – Deshayes, 1855c: 326. — Type locality: Moreton Bay, Queensland, Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978085, 2 syntypes. — Current status: synonym of Cucurbitula cymbium (Spengler, 1783) ( Taki & Habe 1955: 3-4; Huber 2010a: 279; Lutaenko et al. 2021: 240). rauliniana , Gastrochaena – Deshayes, 1856-DescrCoqFoss: 2, pl. 2, figs 22-24; 1857: 100-101. — Type localities: Jeurre & Étréchy. — Type age: Oligocene (Rupelian). Gastrochaena “ raulini ” Deshayes, 1856 (unjustified emendation by Cossmann & Peyot 1909a: 81). — Current status: Gastrochaena rauliniana Deshayes, 1856 ( Lozouet & Maestrati 2012a: 266, 268, fig. 171: 27-30). Possibly a Rocellaria (J. G. Carter, pers. comm., August 2020).

retzii , Gastrochoena – Deshayes, 1863: 7-9, pl. 1, figs 1-3. — Type locality: La Réunion. — Type age: Recent. Synonym of Spengleria mytiloides ( Lamarck, 1818) [ Gastrochaena ] ( Taki & Habe 1955: 5). — Current status: synonym of Spengleria plicatilis (Deshayes, 1855) [ Gastrochaena ] ( Nielsen 1986).

ruppellii , Gastrochaena – Deshayes, 1855c: 328-329. — Type locality: Red Sea, Ruppell; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978097, holotype. — Current status: synonym of Gastrochaena cuneiformis Spengler, 1783 ( Carter et al. 2008: 110), or Rocellaria ruppellii (Deshayes, 1855) ( Huber 2010a: 278, 672), perhaps unaware of Carter’s analysis. Shell atypical of G. cuneiformis in terms of some (but not all) posterior commarginal ribs more rounded, less distinct, and shell higher near umbo, but probably intraspecific variation (J. G. Carter, pers. comm., August 2020).

spathulata , Gastrochaena – Deshayes, 1855c: 329. Bohol. — Type locality: Philippine Islands ; Cuming coll. — Type age: Recent. — Type material: NHMUK 1978099, holotype. — Current status: Gastrochaena spathulata Deshayes, 1855 ( Huber 2010a: 671).

spengleri , Gastrochaena – Deshayes, 1857-DescrCoqFoss: 105-106, 17, pl. 17, figs 1-5 [fig. 5 not listed in text], not preoccupied by Blainville, 1824, nomen nudum. — Type locality: Valmondois. — Type age: Eocene (Bartonian). Spengleria spengleri (Deshayes, 1857) ( Glibert & Van de Poel 1966a: 23) . — Current status: Gastrochaena (Spengleria) spengleri Deshayes, 1857 ( Le Renard & Pacaud 1995: 67).

tenera , Gastrochaena – Deshayes, 1855c: 327. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: not found. — Current status: nomen dubium ( Huber, 2010a: 672) , or Gastrochaena tenera Deshayes, 1855 ( Poppe 2011: 178, pl. 1084, fig. 2).

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dilatata , Gastrochaena – Leymerie, 1841: 341, nomen nudum; 1842: 2-3, 24, pl. 3, fig. 1a-c, ex Deshayes ms. — Type localities: Chenay, Vendeuvre & Fouchères. — Type age: Cretaceous (Hauterivian). — Current status: possibly Stenochaena (J. G. Carter, pers. comm., August 2020).