PSAMMOBIIDAE J. Fleming, 1828

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 771-775

Family PSAMMOBIIDAE J. Fleming, 1828

Capsella – Deshayes, 1855c: 347-350, non J. E. Gray, 1851. — Type species (SD herein): Capsa (Capsella) crassula Deshayes, 1855 . — Current status: this is the same type species as that of Crassulobia Willan, 1993, so Deshayes genus falls into its synonymy. It is possible that Deshayes merely misused Gray’s genus, but that seems unlikely in that Gray’s genus belongs in the Donacidae and is now regarded as a synonym of Donax Linnaeus, 1758 .

REMARK

Capsa Bruguière, 1797 , was misused for some of the following species, but it is a tellinid genus. Recently, the long-used genus Soletellina Blainville, 1824 , was synonymized with Hiatula Modeer, 1793 ( Matsubara, 2013).

abrupta, Psammobia – Deshayes, 1855c: 324-325. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984300, holotype. — Current status: synonym of Gari pulcherrima (Deshayes, 1855) ( Willan 1993: 35, fig. 141 – holotype).

amoena , Psammobia – Deshayes, 1855c: 323. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1964010, holotype. — Current status: synonym of the Asian Gari amethystus (W. Wood, 1815) [ Solen ] ( Willan 1993: 38- 39, fig. 152 – holotype).

angusta , Psammobia – Deshayes, 1855c: 320. — Type locality: Senegal; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984288.1, lectotype ( Willan 1993); 1984288/2, paralectotype. — Current status: synonym of Gari modesta (Deshayes, 1855) [ Psammobia ] (E. A. Smith 1885: 95; Willan 1993: 23, fig. 84 – lectotype).

anomala , Psammobia – Deshayes, 1855c: 320. — Type locality: Zebu, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984290.1 lectotype ( Willan 1993); 1984290/2- 5 paralectotypes. — Current status: Gari anomala (Deshayes, 1855) ( Willan 1993: 22-23, figs 7, 8, 76 [lectotype]-81; Huber 2010a: 324-325; Willan & Tagaro, in Poppe 2011: 356, pl. 1173, figs 1-2; Matsukuma, in Okutani 2017: 1260, pl. 554, fig. 5).

basteroti , Tellina – Deshayes, 1857-DescrCoqFoss: 348. Based on Tellina biangularis Deshayes var. “β” of Basterot (1825: 86). — Type locality: Bordeaux. Miocene (Aquitanian). Synonym of Solecurtus affinis ( Dujardin, 1837) (d’Orbigny 1852: 99, no. 1853). Gari affinis ( Dujardin, 1837) ( Glibert & Van de Poel 1967: 122) . — Current status: Gari (Psammobia) affinis ( Dujardin, 1837) ( Lesport et al. 2019: 44, pl. 17, figs 6-9).

baudoni , Psammobia –Deshayes, 1857-DescrCoqFoss: 378-379, 23, pl. 25, figs 15-18 [in text as pl. 25]. — Type locality: Hadancourtle-Haut-Clocher (Oise). — Type age: Eocene (Lutetian). — Current status: Gari (Psammotaena) baudoni (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93). bicarinata , Psammobia – Deshayes, 1855c: 322. — Type localities: Madagascar & Zanzibar; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984297, 2 syntypes. — Current status: Gari bicarinata (Deshayes, 1855) ( Oliver 1992: 163, pl. 36), or synonym of Psammobia insignis Deshayes, 1855 ( Willan 1993: 37; Rusmore-Villaume 2008: 260-261).

caillati , Psammobia – Deshayes, 1857-DescrCoqFoss: 381, 23, pl. 23, figs 19-20. — Type localities: Parnes, Thiverval-Grignon, Chaumont-en-Vexin & Gomerfontaine. — Type age: Eocene (Lutetian). — Type species (OD) of Psammodonax Cossmann, 1886 . Psammodonax caillati (Deshayes, 1857) ( Glibert & Van de Poel 1967: 120) . — Current status: Gari (Psammodonax) caillati (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

callosa , Sportella – Deshayes, 1857-DescrCoqFoss: 48, pl. 48, figs 18-20; 1858-DescrCoqFoss: 595-596. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). Gobraeus callosus (Deshayes, 1857) ( Glibert & Van de Poel 1967: 124) . — Current status: Gari (Psammotaena) callosa (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

candidula , Psammobia – Deshayes, 1855c: 319. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984286, 3 syntypes. — Current status: Gari candidula (Deshayes, 1855) ( Huber 2015: chapter 5 on CD).

chinensis , Capsa (Capsella) – Deshayes, 1855c: 348. — Type locality: China; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984261, 2 syntypes. — Current status: Gari (Psammotaena) chinensis (Deshayes, 1855) ( Willan 1993: 60; Huber 2010a: 329, 695).

compta , Psammobia – Deshayes, 1855c: 321. — Type locality: Tasmania; Cuming coll. — Type age: Recent. — Type material: NHMUK 1841.2.6.423, lectotype ( Willan 1993); 1837.7.15.310, paralectotype; 1837.7.15.144, paralectotype. — Current status: synonym of Gari livida ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 41-42, fig. 162 – lectotype).

consobrina , Psammobia – Deshayes, 1857-DescrCoqFoss: 373-374, 23, pl. 23, figs 3-4. — Type localities: Brimont & Châlons-sur-Vesle. — Type age: Paleocene (Thanetian). 1858: 552. Garum consobrina (Deshayes, 1857) ( Glibert & Van de Poel 1967: 119) . — Current status: Gari (Garum) consobrina (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Leroy et al. 2014: 28, pl. 12, fig. 1a-b).

contraria , Psammobia – Deshayes, 1863: 11, pl. 1, figs 20-21. — Type locality: La Réunion. — Type age: Recent. — Type material: missing ( Willan 1993: 9). — Type species (SD Oliver, 1992) of Heteroglypta Martens, 1880 . — Current status: Heteroglypta contraria (Deshayes, 1863) ( Oliver 1992: 162, pl. 36, as Gari (Heteroglypta) ; Willan 1993: 9-11, figs 27-37, 382; Huber, 2010a: 332; Willan & Tagaro, in Poppe 2011: 362, pl. 1176, figs 4-5; Matsukuma, in Okutani 2017: 1262, pl. 555, fig. 5; Lutaenko et al. 2021: 220).

corrugata , Psammobia – Deshayes, 1855c: 324. — Type locality: Zebu Island, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196413, holotype. — Current status: synonym of Gari maculosa ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 26-27, fig. 102 – holotype).

crassula , Capsa (Capsella) – Deshayes, 1855c: 349. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984268.1, lectotype ( Willan 1993); 1984268/2-3, paralectotypes. — Type species (OD) of Crassulobia Willan, 1993. — Current status: Gari (Crassulobia) crassula (Deshayes, 1855) ( Willan 1993: 68-70, figs 289 [lectotype]-297, 404-405; Huber 2010a: 328). debilis , Psammobia – Deshayes, 1857-DescrCoqFoss: 377-378, 22, as P. fragilis , pl. 22, figs 15-17; 1865-DescrCoqFoss: 666 [error corrected]. — Type locality: Châlons-sur-Vesle. — Type age: Paleocene (Thanetian). 1858: 552. Gobraeus debilis (Deshayes, 1857) ( Glibert & Van de Poel 1967: 124) . — Current status: Gari (Psammotaena) debilis (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Leroy et al. 2014: 28, pl. 12, figs 2-4).

difficilis , Capsa (Capsella) – Deshayes, 1855c: 347. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984262.1, lectotype ( Willan 1993); 1984268/2-3, paralectotypes; 1984262, 3 syntypes. — Current status: synonym of Gari (Psammotaena) elongata ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 61-62, fig. 251 – lectotype).

dispar , Psammobia – Deshayes, 1855c: 325. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985170.1, lectotype ( Willan 1993); 1985170/2, paralectotype. Gari (Heteroglypta) dispar (Deshayes, 1855) ( Oliver 1992: 164, pl. 36). — Current status: synonym of Gari pennata (Deshayes, 1855) ( Willan 1993: 31, fig. 120 – lectotype; Rusmore-Villaume 2008: 262-263).

donacilla , Psammobia – Deshayes, 1857-DescrCoqFoss: 382, 23, as Psammobia donacina , pl. 23, figs 21-22; 1865-DescrCoqFoss: 666 [error in pl. expl. corrected]. — Type localities: Thiverval-Grignon, Parnes, Damery, Chaumont-en-Vexin, Fontenay-Saint-Père, Boursault & Gomerfontaine. — Type age: Eocene (Lutetian). — Current status: Gari (Psammodonax) donacilla (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

dutemplei , Psammobia – Deshayes, 1857-DescrCoqFoss: 374, 23, pl. 23, figs 10-12. — Type locality: Damery. — Type age: Eocene (Lutetian). — Type species (OD) of Garum Dall, 1900 . Garum placed on Official List by ICZN Opinion, 910 (1970), but listed there with the wrong type species [ Psammobia filosa Conrad, 1833 ]. Garum dutemplei (Deshayes, 1857) ( Glibert & Van de Poel 1967: 119) . Gari (Psammodonax) dutemplei (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70) , – Current status: Gari (Garum) dutemplei (Deshayes, 1857) ( Pacaud 2008: 93) .

elegans , Psammobia – Deshayes, 1855c: 322. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984298, 6 syntypes. — Current status: synonym of the western Indo-Pacific Psammobia insignis Deshayes, 1855 ( Willan 1993: 37).

grata , Psammobia – Deshayes, 1855c: 318-319. — Type locality: “ Amboina ”; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984283, holotype. — Current status: synonym of the northeastern Atlantic Gari (Gobraeus) depressa (Pennant, 1777) [ Tellina ] ( Willan 1993: 49).

insignis , Psammobia – Deshayes, 1855c: 322. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984296, holotype. Gari insignis (Deshayes, 1855) ( Willan 1993: 37) . — Current status: the western Indo-Pacific Gari (Psammobia) insignis (Deshayes, 1855) ( Huber 2010a: 326) .

intermedia , Psammobia – Deshayes, 1855c: 319. — Type locality: Portugal; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984284, 2 syntypes. Synonym of Gari (Gobraeus) virgata ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 49). — Current status: Gari intermedia (Deshayes, 1855) ( Cossignani & Ardovini 2011: 57, 471).

lamarckii , Psammobia – Deshayes, 1857-DescrCoqFoss: 376; first edition, pl. 2, figs 24, 25 [in text as 24, 26]; 1865-DescrCoqFoss: 665 [error noted]. — Type localities: Parnes, Mouchy-le-Châtel, Montmirail & Vaudancourt. — Type age: Eocene (Lutetian). — Type species (M) of Amphipsammus Cossmann, 1913. Gobraeus lamarckii (Deshayes, 1857) , as G. “ lamarcki ”) ( Glibert & Van de Poel 1967: 125). — Current status: Gari (Amphipsammus) lamarckii (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 94).

lata , Psammobia – Deshayes, 1855c: 318. — Type localities: Central America & Santa Elena, Ecuador; Cuming coll. — Type age: Recent. — Type material: NHMUK 1966549, lectotype, 2 paralectotypes; 1984279, probable paralectotype. Gari (Gobraeus) lata (Deshayes, 1855) ( Huber 2010a: 327) . — Current status: Gari lata (Deshayes, 1855) ( Coan & Valentich-Scott 2012: 685) .

layardi , Capsa (Capsella) – Deshayes, 1855c: 348. — Type locality: Ceylon; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984276, 3 syntypes. — Current status: Gari (Psammotaena) layardi (Deshayes, 1855) ( Huber 2010a: 329) .

layardi , Psammobia –Deshayes, 1855c: 323.— Type locality: Philippine Islands;Cuming coll.— Type age:Recent. —Type material: NHMUK 196422, holotype. — Current status: synonym of Gari maculosa ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 26-27, fig. 99 – holotype).

lunulata , Capsa (Capsella) – Deshayes, 1855c: 349. — Type localities: Philippine Islands; Ceylon (var.); Cuming coll. — Type age: Recent. — Type material: NHMUK 1984272, 5 syntypes. — Current status: Soletellina lunulata (Deshayes, 1855) ( Willan 1993: 75; Huber 2010a: 330), now Hiatula lunulata (Deshayes, 1855) .

marmorea, Psammobia – Deshayes, 1855c: 324. — Type localities: Darnley Island, Sydney, Australia & Moluccas (latter two “var.); Cuming coll. — Type age: Recent. — Type material: NHMUK 1846.9.16.112, holotype; 1846.9.14.111, syntype of variety Alpha from Sydney; 196447-196448, syntypes of variety Beta from Moluccas. — Current status: synonym of Gari maculosa ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 26-27, fig. 103 – holotype).

maxima , Psammobia – Deshayes, 1855c: 317. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1966552.1-2, lectotype & paralectotype. — Current status: the Panamic Gari (Gobraeus) maxima (Deshayes, 1855) ( Huber 2010a: 326; Coan & Valentich-Scott 2012: 685).

menkeana, Psammobia – Deshayes, 1855c: 319-320. — Type locality: east coast of Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984287.1, lectotype ( Willan 1993); 1984287/2-3, paralectotypes. — Current status: synonym of Gari modesta (Deshayes, 1855) [ Psammobia ] (E. A. Smith 1885: 95; Willan 1993: 23, fig. 83 – lectotype).

minima , Capsa –Deshayes, 1857-DescrCoqFoss: 386, 11 bis, pl. 11 bis, figs 29-31 [in text as pl. 16 bis, figs 81-33]; 1865-DescrCoqFoss: 665 [errors noted]. — Type localities: Montjavoult, Thiverval-Grignon, Houdan & Coincourt. — Type age: Eocene (Lutetian). — Type species (OD) of Asaphinella Cossmann, 1886 . — Current status: Asaphinella minima (Deshayes, 1857) ( Glibert & Van de Poel 1967: 126; Le Renard & Pacaud 1995: 71; Pacaud 2008: 93; Nevesskaja et al. 2013: 377, fig. 138-5).

minor , Capsa (Capsella) – Deshayes, 1855c: 347. — Type locality: Manilla, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984269.1, lectotype ( Willan 1993); 1984269/2-4, 3 paralectotypes. Synonym of Gari (Psammotaena) elongata ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 61-62, fig. 250 – lectotype). — Current status: Psammotaena minor (Deshayes, 1854) ( Qi 2004: 292, pl. 159, fig. A).

modesta , Psammobia – Deshayes, 1855c: 319. — Type locality: Moreton Bay, Queensland, Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984285.1, lectotype ( Willan 1993); 1984285/2-3, paralectotypes. — Current status: Gari modesta (Deshayes, 1855) ( Willan 1993: 23-26, figs 82 [lectotype]-93, 383). neglecta , Psammobia – Deshayes, 1857-DescrCoqFoss: 375-376, 23, pl. 23, figs 5-7. — Type locality: Ludes. — Type age: Eocene (Priabonian). Gobraeus neglectus (Deshayes, 1857) ( Glibert & Van de Poel 1967: 125) . — Current status: Gari (Psammotaena) neglecta (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Pacaud & Ledon 2007: 11).

nitens , Psammobia – Deshayes, 1857-DescrCoqFoss: 380, 21, pl. 21, figs 27-28. — Type locality: Ormoy-la-Rivière. — Type age: Oligocene (Rupelian). — Current status: Gari nitens (Deshayes, 1857) ( Lozouet & Maestrati 2012a: 258, 261, fig. 166: 5-8).

nitida , Psammobia –Deshayes, 1857-DescrCoqFoss: 370, 24, pl. 24, figs 1-2. — Type localities: Parnes, Thiverval-Grignon, Houdan, Hermonville & Coincourt. — Type age: Eocene (Lutetian). Synonym of Soletellina appendiculata ( Lamarck, 1806) [ Solen ] ( Cossmann 1886a: 89-90). — Current status: Gari (Psammoica) appendiculata ( Lamarck, 1806) ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

nivosa, Psammobia – Deshayes, 1855c: 317-318. — Type locality: Negros, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984277.1, lectotype ( Willan 1993); 1984277/2, paralectotype. — Current status: synonym of Gari (Dysmea) occidens (Gmelin, 1791) [ Solen ] ( Willan 1993: 56-57, fig. 228 – lectotype).

oblonga , Psammobia – Deshayes, 1855c: 321. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984295, holotype. — Current status: synonym of the Red Sea Soletellina rosea (Gmelin, 1791) [ Solen ] ( Huber 2010a: 331, 695), now Hiatula rosea (Gmelin, 1791) .

obtusalis , Donax – Deshayes, 1825-DescrCoqFoss: 109-110, 9, pl. 18, figs 7-8; 1830b-EncyMeth: 101; 1835-HistNatAnim2: 251- 252; 1839-HistNatAnim3: 583; 1857-DescrCoqFoss: 383-384, as Psammobia obtusalis . — Type localities: Pierrelaye & Beauchamp. — Type age: Eocene (Bartonian). Psammodonax obtusalis (Deshayes, 1825) ( Glibert & Van de Poel 1967 120). — Current status: Gari (Psammodonax) obtusalis (Deshayes, 1825) ( Le Renard & Pacaud 1995: 70) .

oriens , Psammobia – Deshayes, 1855c: 318. — Type locality: Japan; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984277.1, lectotype ( Willan 1993); 1984277/2, paralectotype. Gari (Gobraeus) oriens (Deshayes, 1855) ( Willan 1993: 52-55, figs 214-216 [lectotype]-222, 395; Willan & Tagaro, in Poppe 2011: 350, pl. 1170, figs 4-5, who treated it as a subspecies of the West African Gari castrensis Spengler, 1794 ). Gari (Gobraeus) oriens (Deshayes, 1855) ( Huber 2010a: 327) . — Current status: Gobraeus oriens (Deshayes, 1855) (Matsukuma, in Okutani 2017: 1261-1262, pl. 555, fig. 2). Poppe (2018: 173) concluded that Gari oriens (Deshayes, 1855) should be used for the Indo-Pacific species, with G. castrensis (Spengler, 1794) restricted to the tropical eastern Atlantic.

ornata , Psammobia – Deshayes, 1855c: 323. — Type locality:Ticao Island, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196423, lectotype ( Willan 1993); 196424/1- 2, paralectotypes; 196425, 1 paralectotype; 196426, 2 paralectotypes. — Current status: synonym of Gari maculosa ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 26-27, fig. 100 – lectotype).

pallida , Psammobia – Deshayes, 1855c: 323-324. — Type locality: Red Sea; Cuming coll. — Type age: Recent. — Type material: NHMUK 1964046.1, lectotype ( Willan 1993); 1964046/2, paralectotype; 1844.6.3.594, paralectotype. — Current status: Gari pallida (Deshayes, 1855) ( Willan 1993: 19-22, figs 6, 7, 57 [lectotype]-75; Huber 2010a: 325; Willan & Tagaro, in Poppe 2011: 356, pl. 1173, figs 3-4; Matsukuma, in Okutani 2017: 1260-1261, pl. 554, fig. 6).

palmula , Psammobia – Deshayes, 1855c: 325-326. — Type locality: Sydney, Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985171, holotype. — Current status: synonym of Gari squamosa ( Lamarck, 1818) [ Psammobia ] ( Hedley 1913: 274; Willan 1993: 33, fig. 133 – lectotype), or separable Grammatomya palmula (Deshayes, 1855) (Matsukuma, in Okutani 2017: 1261, pl. 554, fig. 9, as G. “ palmura ”).

papyracea , Psammobia – Deshayes, 1857-DescrCoqFoss: 372. Replacement name for Psammobia tellinella (Deshayes, 1824) [ Solen ] (see below), non P. tellinella Lamarck, 1818 .

pennata , Psammobia – Deshayes, 1855c: 325. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material:NHMUK 1985169, holotype. — Current status: the Indo-Pacific Gari pennata (Deshayes, 1855) ( Willan 1993: 31-33, figs 119 [holotype]-130, 386; Huber 2010a: 325; Severns 2011: 480, pl. 220, fig.3; Willan &Tagaro, in Poppe 2011: 362, pl. 1176, figs 1-3; Boutet et al. 2020: 661).

petalina , Psammobia – Deshayes, 1855c: 320-321. — Type locality: China; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984292.1, lectotype ( Willan 1993); 1984292/2-3, paralectotypes. — Current status: Soletellina petalina (Deshayes, 1855) ( Willan 1993: 82-83, figs 331 [lectotype]-335, 411; Huber 2010a: 331; Matsukuma, in Okutani 2017: 1262, pl. 555, fig. 8), now Hiatula petalina (Deshayes, 1855) .

praestans , Psammobia – Deshayes, 1855c: 322-323. — Type locality: Moluccas Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196416, lectotype ( Willan 1993); 196417, paralectotype. — Current status: synonym of Gari maculosa ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 26-27, fig. 98 – lectotype).

puella, Psammobia – Deshayes, 1855c: 320. — Type locality: Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984289, holotype.— Current status:synonym of Gari livida ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 41-42, fig. 161 – holotype).

pulcherrima , Psammobia – Deshayes, 1855c: 325. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985168, holotype. — Current status: the Indo-Pacific Gari pulcherrima (Deshayes, 1855) ( Willan 1993: 35-37, figs 140 [holotype]-148, 388; Huber 2010a: 325; Willan & Tagaro, in Poppe 2011: 358, pl. 1174, figs 5-8; Matsukuma, in Okutani 2017: 1261, pl. 554, fig. 10).

purpurea , Sanguinolaria – Deshayes, 1855c: 346. — Type locality: none given [clarified as Guaymas, Mexico, byCoan 2002: 4]; Cuming coll. — Type age: Recent. — Type material: NHMUK 1966539.1- 3, lectotype & 2 paralectotypes. — Current status: synonym of the Panamic Sanguinolaria tellinoides A. Adams, 1850 ( Coan & Valentich-Scott 2012: 693).

radiata , Capsa (Capsella) – Deshayes, 1855c: 348. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984267.1, lectotype ( Willan 1993); 1984267/2-3, paralectotypes. — Current status: synonym of Gari (Psammotaena) elongata ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 61-62, fig. 253 – lectotype).

rosacea , Capsa (Capsella) – Deshayes, 1855c: 348. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984275.1 lectotype ( Willan 1993); 1984275/2-3, paralectotypes. — Current status: synonym of Gari (Psammotaena) elongata ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 61-62, fig. 254 – lectotype).

rubicunda, Psammobia – Deshayes, 1855c: 324. — Type locality: Ticao Island , Philippine Islands ; Cuming coll. — Type age:Recent. — Type material: NHMUK 1984299.1, lectotype ( Willan 1993);

1984299/2, paralectotype. Synonym of Gari maculosa ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 26, fig. 101 – lectotype).

rufa , Capsa (Capsella) –Deshayes, 1855c: 347-348. — Type locality: Manilla, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984263.1, lectotype ( Willan 1993); 1964263/2, paralectotype. — Current status: synonym of Gari (Psammotaena) elongata ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 61-62, fig. 252 – lectotype).

solenella, Capsa (Capsella) –Deshayes, 1855c: 349-350.— Type locality: Manilla, Philippine Islands; Cuming coll. — Type age: Recent. —Type material:NHMUK 1984271.1, lectotype ( Willan 1993); 1984271/2-3, paralectotypes.— Current status: synonym of Gari elongata ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 61-62, fig. 255 – lectotype).

spathula , Psammobia –Deshayes, 1857-DescrCoqFoss: 382-383, 24, pl. 24, figs 3-5. — Type locality: Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Current status: Gari (Psammodonax) spathula (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

stampinensis , Psammobia – Deshayes, 1857-DescrCoqFoss: 378, 23, pl. 23, figs 13-14. — Type localities: Morigny-Champigny, Etréchy. — Type age: Oligocene (Rupelian). Haesslet, Belgium; Délémont, Switzerland. — Current status: Psammobia stampinensis Deshayes, 1857 ( Lozouet & Maestrati 2012a: 258, 261, figs 166: 1-4).

striata , Psammobia –Deshayes, 1855c: 321. — Type locality: Tasmania; Cuming coll. — Type age: Recent. — Type material: NHMUK 1842.11.2.8.9, 2 possible syntypes. — Current status: synonym of Gari livida ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 41-42, fig. 163 – possible syntype).

tellinella , Solen – Deshayes, 1824-DescrCoqFoss: 28, 2, pl. 4, figs 1-2, non Psammobia tellinella Lamarck, 1818 ; 1832-EncyMeth: 963; 1857-DescrCoqFoss: 372, Psammobia papyracea , replacement name, when he moved this Solen into Psammobia . — Type locality: Tancrou. — Type age: Eocene (Bartonian). Macropsammus tellinella (Deshayes, 1824) ( Glibert & Van de Poel 1967: 126) . Gari (Psammoica) tellinella (Deshayes, 1824) ( Le Renard & Pacaud 1995: 70) . — Current status: because Lamarck’s species is now placed in Gari , this should once again be Gari (Psammoica) papyracea (Deshayes, 1857) .

tenera , Psammobia – Deshayes, 1857-DescrCoqFoss: 372, 24, pl. 24, figs 6-8. — Type localities: Damery, Parnes & Hermonville. — Type age: Eocene (Lutetian). Synonym of Soletellina appendiculata ( Lamarck, 1806) [ Solen ] ( Cossmann 1886a: 89-90). — Current status: Gari (Psammoica) appendiculata ( Lamarck, 1806) ( Le Renard & Pacaud 1995: 70) .

tenuicula , Psammobia – Deshayes, 1857-DescrCoqFoss: 380-381, 24, as P. “ tenuis ”, pl. 24, figs 12-14; 1866: 666 [error corrected]. — Type localities: Gentilly, Hermonville & La Chapelle-en-Serval. — Type age: Eocene (Bartonian). Gobraeus tenuiculus (Deshayes, 1857) ( Glibert & Van de Poel 1967: 125) . — Current status: Gari (Psammotaena) tenuicula (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70) .

tenuis , Capsa (Capsella) – Deshayes, 1855c: 349. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984274.1, lectotype ( Willan 1993); 1984274/2, paralectotype. Soletellina tenuis (Deshayes, 1855) ( Willan 1993: 76) , now genus Hiatula . — Current status: synonym of Gari virescens (Deshayes, 1855) ( Huber 2015: chapter 5 on CD).

tenuis , Psammobia – Deshayes, 1855c: 320. — Type localities: Philippine Islands & China (var.); Cuming coll. — Type age: Recent. — Type material: NHMUK 1984291.1/2, lectotype & paralectotype. — Current status: synonym of Gari anomala (Deshayes, 1855) ( Willan 1993: 22, fig. 77 – lectotype). togata , Psammobia – Deshayes, 1855c: 318. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984280.1, lectotype; 1984280/2, paralectotype. Gari (Psammotaena) togata (Deshayes, 1855) ( Willan 1993: 64-66, figs 269 [lectotype]-274, 401). — Current status: Psammotaena togata (Deshayes, 1854) ( Qi 2004: 292, pl. 159, fig. D; Huber 2010a: 329; Willan & Tagaro, in Poppe 2011: 346, pl. 1168, fig. 6; Matsukuma, in Okutani 2017: 1261, pl. 554, fig. 15).

tripartita , Psammobia – Deshayes, 1855c: 321-322. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196414, lectotype ( Willan 1993); 196415/1-2, paralectotypes; 196416, paralectotype. — Current status: synonym of Gari amethystus (W. Wood, 1815) [ Solen ] ( Willan 1993: 38-39, fig. 151 – lectotype).

tristis , Psammobia – Deshayes, 1855c: 318. — Type locality: Amboina ; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984282.1, lectotype ( Willan 1993); 1984272/2, paralectotype. — Current status: synonym of Gari (Gobraus) strangeri (J. E. Gray, 1843) [ Psammobia ] ( Willan 1993: 49-50, fig. 206 – lectotype).

vaudini , Psammobia – Deshayes, 1857-DescrCoqFoss: 383, 22, pl. 22, figs 12-14. — Type localities: Laon & Hérouval. — Type age: Eocene (Ypresian). Psammodonax vaudini (Deshayes, 1857) ( Glibert & Van de Poel 1967: 120) . — Current status: Gari (Psammodonax) vaudini (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70) .

virescens , Capsa (Capsella) –Deshayes, 1855c: 349. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984270, 3 syntypes. Soletellina virescens (Deshayes, 1855) ( Willan 1993: 76; Qi 2004: 292-293, pl. 159, fig. B), now genus Hiatula . — Current status: the southeast Asian Gari (Psammotaena) virescens (Deshayes, 1855) ( Huber 2010a: 329) .

vitrea , Sanguinolaria – Deshayes, 1855c: 326. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985172, 2 syntypes. — Current status: Sanguinolaria vitrea Deshayes, 1855 ( Huber 2010a: 332-333, 696; 2015: 749) from Madagascar .

zelandica , Psammobia – Deshayes, 1855c: 319. — Type locality: New Zealand; Cuming coll. — Type age: Recent. — Type material: MNHN-IM-2000-35993, lectotype ( Willan 1993); MNHN- IM-2000-35994, paralectotype. — Current status: synonym of Gari (Gobraus) strangeri (J. E. Gray, 1843) [ Psammobia ] ( Powell 1979: 418; Willan 1993: 49-50, fig. 207 – lectotype).

●

Psammotella – H. Adams & A. Adams, 1856 (Adams & Adams 1856: 393), ex Deshayes ms, non Herrmannsen, 1852. — Type species (M): Solen ruber Schroter, 1788 . — Current status: synonym of Hiatula Modeer, 1793 .

REMARK

Deshayes apparently added manuscript names on labels in Cuming’s material of this family, but he never published the species involved. Reeve then credited Deshayes with the following ms names, although Reeve must be considered their author. These Reeve species are listed here along with one other species credited as “ ex Deshayes ms”.

acuminata , Soletellina – Reeve, 1857a: pl. 3, fig. 12, ex Deshayes ms. Soletellina acuminata Reeve, 1857 ( Willan 1993: 75). — Current status: synonym of Hiatula diphos (Linnaeus, 1771) [ Solen ] ( Huber, 2015: chapter 5 on CD), the type species of Hiatula Modeer, 1793 .

adamsii , Soletellina – Reeve, 1857a: pl. 2, fig. 5, ex Deshayes ms. — Current status: Soletellina adamsii (Reeve, 1857) ( Willan 1993: 75) .

ambigua , Psammotella – Reeve, 1857b: pl. 1, fig. 5, ex Deshayes ms. — Current status: Soletellina ambigua Reeve, 1857 ( Willan 1993: 67), now Hiatula ambigua (Reeve, 1857) .

appendiculatus , Solen (Macropsammus) – Listed by Cossmann & Peyrot (1910: 295, fig. 44) as a Deshayes species, it was actually made available by Lamarck, 1806. Lamarck’s species is the type species (OD) of Psammoica Dall, 1900, non Solier, 1835 [ Coleoptera ], which was renamed Macropsammus Cossmann, 1902 . — Current status: Macropsammus appendiculatus ( Lamarck, 1806) .

atrata , Soletellina – Reeve, 1857a: pl. 3, fig. 14, ex Deshayes ms. Soletellina atrata Reeve, 1857 ( Willan 1993: 75; Huber 2010a: 330). Matsukuma, in Okutani (2017: 1262), attributed this species to “Deshayes in Reeve”. — Current status: Hiatula atrata (Reeve, 1857) .

cumingiana , Soletellina – Reeve, 1857a: pl. 1, fig. 4, ex Deshayes ms. Soletellina cumingiana Reeve, 1857 ( Willan 1993: 75). — Current status: synonym of Hiatula adamsii (Reeve, 1857) ( Huber 2015: chapter 5 on CD).

epidermia , Soletellina – Reeve, 1857a: pl. 1, fig. 3, ex Deshayes ms. — Current status: synonym of Soletellina biradiata (W. Wood, 1815) [ Solen ] ( Willan 1993: 77), now Hiatula biradiata (W. Wood, 1815) .

incerta , Soletellina – Reeve, 1857a: pl. 3, fig. 13, ex Deshayes ms. — Current status: synonym of Soletellina alba ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 79), now Hiatula alba ( Lamarck, 1818) .

japonica , Soletellina – Reeve, 1857a: pl. 4, fig. 16, ex Deshayes ms. — Current status: Nuttallia japonica (Reeve, 1857) ( Huber 2010a: 331) . Kantor & Sysoev (2005: 362), as “Deshayes in Reeve”.

malaccensis , Psammotella – Reeve, 1857b: pl. 1, fig. 2, ex Deshayes ms. — Current status: synonym of Hiatula ambigua (Reeve, 1857) ( Willan 1993: 67) .

nymphalis, Soletellina – Reeve, 1857a: pl. 1, fig. 2, ex Deshayes ms. — Current status: synonym of Soletellina biradiata (W. Wood, 1815) [ Solen ] ( Willan 1993: 77), now Hiatula biradiata (W. Wood, 1815) .

oblonga , Psammotella – Reeve, 1857b: pl. 1, fig. 7, ex Deshayes ms. — Current status: synonym of Hiatula rosea (Gmelin, 1791) [ Solen ] ( Huber 2015: chapter 5 on CD).

obscurata , Soletellina – Reeve, 1857a: pl. 4, fig. 21, ex Deshayes ms. — Current status: Nuttallia obscurata (Reeve, 1857) ( Huber 2010a: 331) . Kantor & Sysoev (2005: 362) cited this as “Deshayes in Reeve”.

ovalina , Donax – Reeve, 1854d: pl. 3, fig. 17, ex Deshayes ms. No locality indicated.Deshayes, 1855c: 352. Central America; Cuming coll. — Current status: Synonym of Heterodonax pacificus (Conrad, 1837) [ Psammobia ] ( Coan & Valentich-Scott 2012: 689).

philippinensis , Psammotella – Reeve, 1857b: pl. 1, fig. 1, ex Deshayes ms. — Current status: synonym of Hiatula ambigua (Reeve, 1857) ( Willan 1993: 67) .

subradiata , Psammotella – Reeve, 1857b: pl. 1, fig. 6, ex Deshayes ms. — Current status: synonym of Hiatula ambigua (Reeve, 1857) ( Willan 1993: 67) .

tellinaeformis, Psammobia – Reeve, 1856: pl. 5, fig. 31, ex Deshayes ms. — Current status: synonym of Gari livida ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 41).

tumens , Soletellina – Reeve, 1857a: pl. 4, fig. 20, ex Deshayes ms. — Current status: Soletellina tumens Reeve, 1857 ( Willan 1993: 89), now Hiatula tumens (Reeve, 1857) .