Wroughtonia Cameron

Yan, Cheng-Jin, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2017, Review of the tribe Helconini Foerster s. s. from China, with the description of 18 new species, Zootaxa 4291 (3), pp. 401-457 : 415-417

Key to the Chinese species of the genus Wroughtonia Cameron

1 Frontal protuberance near antennal sockets with distinct ledge halfway and blunt dorsally ( Figs 15 H, 20J, 24I, J); posterior half of frontal depression shallow and sublateral carina remain far from level of anterior ocellus ( Figs 15 H, 20J, 24J); [scapus about as long as wide ( Fig. 20 G)]............................................................................. 2

- Frontal protuberance near antennal sockets lamelliform and acute dorsally, without ledge ( Figs 10 H, J, 11H, 13H, 16J, 18J, 19H, 22H); posterior half of frontal depression and sublateral carina variable ( Figs 7 H, 8I, 16I, 18J)................... 4

2 Antenna of ♀ entirely dark brown ( Fig. 20 G); ovipositor sheath about 1.2 times as long as fore wing; setae of ovipositor sheath pale yellowish and slender ( Fig. 20 F); first and second metasomal tergite mainly smooth (except for dorsal carinae; Fig. 20 C). China (Yunnan)......................................................................... W. obtusa sp. nov.

- Antenna of ♀ with distinct white or ivory band ( Figs 15 F, 24G); ovipositor sheath 0.9–1.0 times as long as fore wing; setae of ovipositor sheath dark brown and either rather long and rather erect or short and rather adpressed; first and more or less second tergite coarsely sculptured ( Figs 15 C, 24D).................................................................3

3 Frontal protuberance dorsally convex (widened, tuberculate; Fig. 24 H J); scapus densely setose ventrally ( Fig. 24 G); subposterior crest of scutellum distinct and upcurved ( Fig. 24 C); ventral protuberance of hind femur narrow and partly parallel-sided ( Fig. 24 E); first tergite yellow or pale yellow, at most medio-posteriorly reddish-brown or black; length of eye 1.8–2.0 times temple in dorsal view ( Fig. 24 I); hind coxa brownish yellow; [frontal depression only anteriorly bordered by distinct carina and with minute tooth pointed to median protuberance; basal half of second tergite of female with coarse punctures; male antenna with white band]. China (Guangxi, Fujian, Yunnan, Tibet) ; Vietnam............................ W. unicornis (Turner)

- Frontal protuberance dorsally narrow and truncate ( Fig. 15 H); scapus with few medium-sized setae ( Fig 15 J); subposterior crest of scutellum absent; ventral protuberance of hind femur tooth-shaped and stout ( Fig. 15 D); first tergite brown or dark brown; length of eye about 1.2 times temple in dorsal view ( Fig. 15 H); hind coxa dark brown. China (Shaanxi)............................................................................................... W. eurygenys sp. nov.

4 Frontal protuberance bifurcate dorsally and posterior branch spatulate ( Figs 10 H, J); basal third of antenna yellowish (except largely dark brown scapus); first metasomal tergite very coarsely reticulate ( Fig. 10 D); hind femur with 5 costate transverse ridges ventrally and no single protuberance ( Fig. 10 E). China (Yunnan).......................... W. bifurcata sp. nov.

- Frontal protuberance simple dorsally ( Figs 11 H, 13H, 16J, 18J, 19H, 22H); basal third of antenna black; first metasomal tergite usually less sculptured ( Figs 7 C, 8D, 9C, 12C, 13D, 18C), but sometimes coarsely sculptured ( Figs 21 C, 22D, 27C); hind femur with a single tooth or a blunt protuberance ventrally ( Figs 7 D, 8E, 9D, 12D, 13E, 17D, 20D)....................5

5 First tergite of female subparallel-sided posteriorly, 1.7–1.8 times longer than its apical width; length of eye about 1.5 times temple in dorsal view; [ovipositor sheath about 1.2 times as long as fore wing; hind femur with acute triangular ventral tooth; basal half of second tergite of ♀ smooth]; China (Taiwan) .................................... W. petila Chou et Hsu

- First tergite of female distinctly widened posteriorly, 1.1–1.5 times longer than its apical width ( Figs 8 D, 9C, 11C, 22D, 25C), if rarely 1.6–2.3 times ( Fig. 7 C, 23D) and/or subparallel-sided then length of eye 0.9–1.0 times temple in dorsal view...... 6

6 Ventral protuberance of hind femur wide and blunt, not distinctly tooth-shaped and area behind it distinctly concave ( Figs 8 E, 12D, 13E, 17D, 18F, 19D, 27D); precoxal sulcus usually vermiculate-rugose medially ( Figs 12 B, 17B, 18C)............. 7

- Ventral protuberance of hind femur tooth-shaped, narrow and acute, and area behind it straight or slightly concave ( Figs 7 D, 9D, 11D, 22E); precoxal sulcus variable, often medially mainly punctate ( Figs 11 B, 17B, 19B).......................15

7 Hind coxa black and most of basal half of hind femur dark brown or brown ( Figs 18 F, 19D); [second metasomal tergite extensively and coarsely punctate ( Figs 18 E, 19C)].............................................................. 8

- Hind coxa brownish yellow and at most basally infuscate ( Figs 8 E, 12D, 13F, 17E, 27F); basal half of hind femur brownish yellow ( Figs 8 E, 12D, 13E, 27D), but largely dark brown in W. indica ( Fig. 17 D)...................................9

8. Medial third of antenna of ♀ with 6–9 white or ivory antennal segments ( Fig. 19 F); hind femur with rather wide ventral protu- berance ( Fig. 19 D); palpi dark brown basally ( Fig. 19 G); fourth metasomal tergite of ♀ more or less exposed ( Fig. 19 F). China (Shaanxi, Henan, Beijing)........................................................... W. nigrifemoralis sp. nov.

- Antenna of ♀ entirely dark brown ( Fig. 18 H); hind femur slender and only rugose ventrally ( Fig. 18 F); palpi brownish yellow basally ( Fig. 18 H); fourth metasomal tergite of ♀ retracted under third tergite ( Fig. 18 E). China (Shaanxi) W. jiangliae sp. nov.

9. Hind coxa reticulate dorsally; vein 1r-m of hind wing slightly longer than vein 1-M; frons smooth posteriorly; [precoxal sulcus shallow and punctate to foveolate; palpi brownish yellow; second tergite blackish brown and foveolate-reticulate]. China (Taiwan) ............................................................................... W. alba (Chou et Hsu)

- Hind coxa largely smooth and sparsely punctulate to partly coarsely punctate dorsally ( Fig. 8 E); vein 1r-m of hind wing 1.7– 2.8 times longer than vein 1-M ( Figs 8 A, 12A, 13B, 17A, 26B, 27A); frons with sublateral carina nearly up to level of anterior ocellus ( Figs 8 F, J, 12H, 17G, 26H), but weakly developed in W. brevicarinata and W. zhejiangensis ( Figs 13 H, 27H); [frontal protuberance pointing posteriorly].......................................................................10

- Frontal protuberance wide triangular, posteriorly mainly flat and finely carinate dorso-laterally ( Figs 8 F, 13H, 17G, 26H); medial third of antenna of ♀ with 1–6 white or ivory antennal segments ( Figs 12 F, 13F, 26F) or basal half of antenna ivory ( Figs 8 G, 17E); middle lobe punctate or coarsely rugose posteriorly ( Fig. 13 C); sublateral carina of frons present anteriorly or border of depression crest-shaped ( Figs 8 F, 13H, 17G, 26H)................................................... 11

11 Temple very coarsely reticulate-punctate ( Fig. 8 H); palpi pale yellow basally ( Fig. 8 G); basal half of ♀ antenna ivory ( Fig. 8 G); [second metasomal tergite (except medially) largely densely punctate ( Fig. 8 D)]. China (Hubei)...................................................................................................... W. albobasalis sp. nov.

- Temple with rather sparse medium-sized punctures ( Figs 12 G, 13G, 17F, 26G); palpi dark brown basally ( Figs 12 F, 13G, 17F, 26G); medial third of ♀ antenna with 1–6 white or ivory antennal segments and remainder dark brown ( Figs 12 F, 13F, 26F), but basal half pale in W. indica ( Fig. 17 E)................................................................. 12

12 First tergite of ♀ with parallel carinae besides dorsal carinae ( Fig. 12 C, but rugose in ♂); antenna of ♀ largely dark brown, with one white or ivory antennal segment, subbasally mainly dark brown ( Fig. 12 F); basal half of ovipositor sheath with short, rather adpressed setae; antenna of ♀ with 26–27 segments; [inner side of hind coxa of male with parallel rugae basally]. China (Zhejiang)...................................................................... W. breviantennata sp. nov.

- First tergite of ♀ coarsely irregularly rugose laterally ( Figs 13 D, 26D) or partly smooth ( Fig. 17 C); antenna of ♀ with 3–6 white or ivory antennal segments ( Figs 13 F, 26F) or basal half pale yellowish ( Fig. 17 E); basal half of ovipositor sheath with longer and less adpressed setae; antenna of ♀ with 32–35 segments............................................13

13 Second metasomal tergite smooth ( Fig. 26 D); first tergite dark brown and less widened posteriorly ( Fig. 26 D); fourth tergite more or less visible beyond third tergite ( Fig. 26 F); lateral carina of frontal depression lamelliform, regular and nearly up to level of ocelli ( Fig. 26 H); [inner side of hind coxa of ♂ with irregular rugae basally]. China (Sichuan, Shaanxi)................................................................................................ W. yaanensis sp. nov.

- Second metasomal tergite coarsely punctate ( Figs 13 D, 17C); first tergite at least partly brownish yellow and strongly widened posteriorly ( Figs 13 D, 17C); fourth tergite more or less retracted below third tergite ( Figs 13 F, 17C); lateral carina of frontal depression less developed ( Fig. 13 H) and irregular if nearly up to level of ocelli ( Fig. 17 G)..........................14

14 First metasomal tergite at least partly smooth and laterally mainly coarsely punctate ( Fig. 17 C); vein r of fore wing 0.6–0.7 times as long as vein 3-SR ( Fig. 17 H); third tergite distinctly punctate medially and posteriorly ( Fig. 17 C); hind femur without distinct ventral serrations in lateral view ( Fig. 17 D); scutellum about twice as long as scutellar sulcus and sulcus with 1 coarse crenula and some rugae; borders of frontal depression weakly protruding and frons rugose-punctate posteriorly ( Fig. 17 G); antenna of ♀ without white submedial band ( Fig. 17 E), but subbasally yellowish. China (Yunnan) , India.................................................................................. W. indica (Singh, Belokobylskij et Chauhan)

- Entire first metasomal tergite coarsely and irregularly reticulate-rugose ( Fig. 13 D); vein r of fore wing 1.0–1.3 times as long as vein 3-SR ( Fig. 13 A); third tergite smooth medially and posteriorly ( Fig. 13 D); hind femur with distinct serrations ventrally in lateral view ( Fig. 13 E); scutellum 1.4–1.6 times as long as scutellar sulcus and sulcus with 3–4 coarse crenulae ( Figs 13 C); borders of frontal depression strongly protruding and frons smooth posteriorly ( Fig. 13 H); antenna of ♀ with white submedial band and subbasally dark brown ( Fig. 13 F). China (Guizhou, Yunnan).................... W. brevicarinata (Yan et Chen)

15 Basal half of antenna yellowish white ( Fig. 25 G); fore and middle femora with a flange-like protuberance medio-ventrally and with a fringe of long setae ventrally ( Figs 25 E, F); apical third of hind tibia pale yellowish ( Fig. 25 D); second tergite largely concentric punctate-striate, except medially ( Fig. 25 C). China (Guangxi)........................ W. varifemora sp. nov.

- Basal half of antenna black or dark brown ( Figs 7 F, 9F, 11F, 14F, 16G, 21F, 23G); fore and middle femora medially without a ventral protuberance and normally setose, at most with a ventral fringe of short setae; apical third of hind tibia dark brown ( Figs 7 D, 9D, 11D, 14E, 16D, 22E, 23E); second tergite with some oblique striae and punctures, rugose punctate or smooth ( Figs 7 C, 9C, 11B, 14D, 16C, 21C, 22D, 23D)..............................................................16

16 In dorsal view length of eye 1.3–1.8 times temple and eyes distinctly protruding ( Figs 11 H, 14H, 21H, 22H); length of first tergite of ♀ 1.2–1.5 times its apical width ( Figs 11 B, 14D, 21C, 22D); ventral tooth of hind femur often rather small or medium-sized ( Figs 14 E, 21D, 22E), except in W. brachygena ( Fig. 11 D); second tergite coarsely punctate or punctate-rugose ( Figs 14 D, 21C, 22D), except in W. brachygena ( Fig. 11 B); basal part of first metasomal sternite at most as long as wide and connected to tergite; [ovipositor sheath 0.6–0.8 times as long as fore wing; scapus about 1.5 times as long as wide]...... 1 7

- In dorsal view length of eye 0.7–1.1 times temple and eyes less protuberant ( Figs 7 H, 9H, 16I, 23I); length of first tergite of ♀ 1.5–2.3 times its apical width ( Figs 7 C, 9C, 16C, 23D); ventral tooth of hind femur robust, equilaterally triangular ( Figs 9 D, 16D, 23E), but slenderer in W. acuta ( Fig. 7 D); second tergite smooth or only partly rather superficially sculptured ( Figs 7 C, 9C, 16C, 23D); basal part of first metasomal sternite distinctly longer than wide apically and well separated from tergite; W. sibirica -group...................................................................................... 21

17 Antenna of ♀ entirely dark brown ( Figs 11 F, 21F); temple coarsely punctate medially ( Figs 11 G, 21G); hind femur robust ( Figs 11 D, 21D)..........................................................................................18

- Antenna of ♀ with white or ivory submedial band ( Fig. 14 F); temple finely punctate medially ( Figs 14 G, 22G); hind femur slender ( Figs 14 E, 22E)................................................................................ 19

18. Middle lobe of mesoscutum distinctly convex and normally protuberant ( Fig. 11 B); mesoscutum punctate-reticulate medioposteriorly; ventral half of temple with distinct smooth interspaces between coarse punctures ( Fig. 11 G); precoxal sulcus densely reticulate-punctate ( Fig. 11 B); second tergite largely smooth, only punctate baso-laterally ( Fig. 11 C); setae of ovipositor sheath dark brown and normal. China (Zhejiang)........................................ W. brachygena sp. nov.

- Middle lobe of mesoscutum rather flat and slightly protuberant ( Figs 21 B, I); mesoscutum medio-posteriorly ( Fig. 21 I) and temple ventrally ( Fig. 21 G) densely vermiculate-rugose, without smooth interspaces; precoxal sulcus coarsely costate-rugose ( Fig. 21 B); second tergite costate-rugose and coarsely punctate, only basally and medially narrowly smooth ( Fig. 21 C); setae of ovipositor sheath pale yellowish and adpressed. Japan, Russia ; China (Zhejiang)............ W. planidorsum (Watanabe)

19 Hind coxa black and basal half of hind femur largely dark brown ( Figs 14 E, F); vein cu-a of fore wing vertical, not parallel with vein 3-CU1 ( Fig. 14 A); ventral tooth of hind femur slender triangular and rather large ( Fig. 14 E); dorsal carinae of first tergite complete ( Fig. 14 D); [lamelliform frontal protuberance wide triangular in lateral view ( Fig. 14 G)]. China (Zhejiang)......................................................................................... W. chui sp. nov.

- Hind coxa and mainly basal half of hind femur brownish yellow ( Figs 22 E, F); vein cu-a of fore wing oblique, subparallel to vein 3-CU1 ( Fig. 22 A); ventral tooth of hind femur more robust (fig. 82 in Chou et Hsu, 1998) or small ( Fig. 22 E); dorsal carinae of first tergite absent posteriorly ( Fig. 22 D)............................................................20

20 Second metasomal tergite about 1.5 times wider than long; fourth and fifth hind tarsal segments brown, darker than third segment; area behind ventral tooth of hind femur rather concave (fig. 86 in Chou et Hsu, 1998); [first tergite largely smooth to moderately foveolate-punctate]. China (Taiwan) ..................................... W. pterolophiae (Chou et Hsu)

- Second metasomal tergite about twice as wide as long ( Fig. 14 D); fourth and fifth hind tarsal segments ivory as third segment ( Fig. 14 E); area behind ventral tooth of hind femur straight ( Fig. 14 E). China (Yunnan)................ W. rugosa sp. nov.

21 Antenna of ♀ with white or ivory submedial band; apical half of hind femur and hind tibia yellowish brown, except apically; [dorsal carinae of first tergite strongly developed, lamelliform basally and converging to each other (fig. 44 in van Achterberg, 1987)]. Palaearctic and Oriental regions; China (Taiwan) .................................... W. spinator (Lepeletier)

- Antenna of ♀ entirely dark brown ( Figs 7 F, 9F, 16G, 23G); apical half of hind femur and hind tibia black or dark brown ( Figs 7 D, 9D, 16D, 23E), at most base of hind tibia brown.........................................................22

22 Hind tarsus dark brown or black ( Figs 7 D, F); hind femur slender and its ventral tooth narrow and acute ( Fig. 7 D); [ovipositor sheath 1.0–1.2 times (Tibet: 1.4 times) as long as fore wing and 0.9 times (Tibet: 1.2 times) as long as body; first metasomal tergite slender ( Fig. 7 D]. China (Sichuan, Tibet)................................................ W. acuta s p. nov.

- Hind tarsus brownish yellow, ivory or white ( Figs 9 D, F, 16D, 23G); hind femur stout and its ventral tooth wide and less acute ( Figs 9 D, 16D, 23E)..................................................................................23

23 Ovipositor sheath 1.2–1.4 times as long as body; length of first tergite 1.8–2.0 times its apical width; scutellum only finely punctate; [vein cu-a of fore wing nearly straight; first metasomal tergite with strong dorsal carinae; hind tarsus white]. Japan, Russia; China (Taiwan) ................................................................ W. uchidai (Watanabe)

- Ovipositor sheath 0.7–0.9 times as long as body ( Figs 9 F, 16G, 23G); length of first tergite 1.5–1.7 times its apical width ( Figs 9 C, 16C, 23D); scutellum at least partly coarsely rugose.....................................................24

24 Propodeum with lamelliform elongate areola ( Fig. 9 E); frontal depression ( Figs 9 H, J), basal half of first (except dorsal carinae) and second metasomal tergites ( Fig. 9 C) smooth; [posterior half of frontal depression shallow]. China (Shanxi)............................................................................................. W. areolata sp. nov.

- Propodeum without areola ( Figs 16 E, 23F), at most with a pair of carinae posteriorly; frontal depression coarsely rugose ( Figs 16 I, 23I); basal half of first tergite coarsely sculptured and second tergite with some rugae or with punctures ( Figs 16 C, 23D)....................................................................................................25

25 Posterior half of frontal depression deep ( Fig. 16 I) and in dorso-lateral view with coarse and emarginated lateral crest ( Fig. 16 H); basal half of hind femur slightly less widened towards level of ventral tooth and length of femur 3.2–3.3 times as long as wide (without ventral tooth; Fig. 16 D); hind femur densely rugose ventrally ( Fig. 16 D); mesoscutum in front of tegulum without lateral carina and largely smooth; occipital carina meeting hypostomal carina near base of mandible ( Fig. 16 H). China (Jilin).................................................................................... W. hei sp. nov.

- Posterior half of frontal depression shallow and in dorso-lateral view without distinct lateral crest ( Fig. 23 I); basal half of hind femur distinctly widened towards level of ventral tooth and length of femur 2.8–3.0 times as long as wide (without ventral tooth; Fig. 23 E); hind femur sparsely rugose ventrally ( Fig. 23 E); mesoscutum in front of tegulum with lateral carina and crenulate or rugose; occipital carina meeting hypostomal carina above level of base of mandible ( Fig. 23 H). China (Shaanxi, Jilin)..................................................................................... W. sibirica (Tobias)

Figures

FIGURE 7. Wroughtonia acuta sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, first and second metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, propodeum, medio-dorsal aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect.

FIGURE 8. Wroughtonia albobasalis sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, first-fourth metasomal tergites, dorsal aspect; E, hind leg, lateral aspect; F, frontal protuberance, dorso-lateral aspect; G, habitus, lateral aspect; H, head, lateral aspect; I, head, dorsal aspect; J, head, anterior aspect; K, apex of ovipositor sheath, lateral aspect.

FIGURE 9. Wroughtonia areolata sp. nov. ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, first-third metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, propodeum, medio-dorsal aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, apex of ovipositor sheath, lateral aspect. J, detail of frontal protuberance, laterodorsal aspect.

FIGURE 10. Wroughtonia bifurcata sp. nov., ♂, holotype. A, fore wing; B, hind wing; C, mesosoma, lateral aspect; D, first-fourth metasomal tergites, dorsal aspect; E, hind leg, lateral aspect; F, propodeum, dorsal aspect; G, habitus, lateral aspect; H, head, lateral aspect; I, head, dorsal aspect; J, frontal bifurcate protuberance, dorso-lateral aspect.

FIGURE 11. Wroughtonia brachygena sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, propodeum, first and second metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, propodeum, medio-dorsal aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect.

FIGURE 12. Wroughtonia breviantennata sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, first and second metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, face and clypeus, anterior aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, propodeum, medio-dorsal aspect.

FIGURE 13. Wroughtonia brevicarinata (Yan et Chen), ♀, holotype. A, fore wing; B, hind wing; C, mesosoma, medio-dorsal aspect; D, metasoma, dorsal aspect; E, hind femur and tibia, lateral aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, propodeum, medio-dorsal aspect.

FIGURE 14. Wroughtonia chui sp. nov., ♀, holotype. A, fore wing; B, hind wing; C, mesosoma, lateral aspect; D, metasoma, dorsal aspect; E, hind femur and tibia, lateral aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, propodeum, medio-dorsal aspect.

FIGURE 15. Wroughtonia eurygenys sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, first and second metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, face and clypeus, anterior aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, propodeum, medio-dorsal aspect.

FIGURE 16. Wroughtonia hei sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, first-third metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, propodeum, medio-dorsal aspect; F, apex of ovipositor sheath, lateral aspect; G, habitus, lateral aspect; H, head, lateral aspect; I, head, dorsal aspect; J, frontal protuberance, latero-dorsal aspect.

FIGURE 17. Wroughtonia indica (Singh et al.), ♀, China. A, hind wing; B, mesosoma, lateral aspect; C, metasoma, dorsal aspect; D, hind femur and tibia, lateral aspect; E, habitus, lateral aspect; F, head, lateral aspect; G, head, dorsal aspect; H, fore wing.

FIGURE 18. Wroughtonia jiangliae sp. nov., ♀, holotype. A, fore wing; B, hind wing; C, mesosoma, lateral aspect; D, mesosoma, dorsal aspect; E, metasoma, dorsal aspect; F, hind leg, lateral aspect; G, apical part of ovipositor sheath, lateral aspect; H, habitus, lateral aspect; I, head, lateral aspect; J, head, dorsal aspect; K, face and clypeus, anterior aspect.

FIGURE 19. Wroughtonia nigrifemoralis sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, first-third metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, face and clypeus, anterior aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, propodeum, medio-dorsal aspect.

FIGURE 20. Wroughtonia obtusa sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, metasoma, dorsal aspect; D, hind leg, lateral aspect; E, propodeum, dorsal aspect; F, apex of ovipositor sheath; G, habitus, lateral aspect; H, head, lateral aspect; I, head, dorsal aspect; J, detail of frontal protuberance, dorso-lateral aspect.

FIGURE 21. Wroughtonia planidorsum (Watanabe), ♀, China. A, wings; B, mesosoma, lateral aspect; C, first and second metasomal tergites, dorsal aspect; D, hind femur and tibia, lateral aspect; E, propodeum, medio-dorsal aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, mesosoma, dorsal aspect.

FIGURE 22. Wroughtonia rugosa sp. nov., ♀, holotype. A, fore wing; B, hind wing; C, mesosoma, lateral aspect; D, first-third metasomal tergites, dorsal aspect; E, hind femur and tibia, lateral aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect.

FIGURE 23. Wroughtonia sibirica (Tobias), ♀, China. A, fore wing; B, hind wing; C, mesosoma, lateral aspect; D, first and second metasomal tergites, dorsal aspect; E, hind femur and tibia, lateral aspect; F, propodeum, medio-dorsal aspect; G, habitus, lateral aspect; H, head, lateral aspect; I, head, dorsal aspect.

FIGURE 24. Wroughtonia unicornis (Turner), ♀, China. A, fore wing; B, hind wing; C, mesosoma, dorsal aspect; D, first-second metasomal tergites, dorsal aspect; E, hind femur and tibia, lateral aspect; F, face and clypeus, anterior aspect; G, habitus, lateral aspect; H, head, lateral aspect; I, head, dorsal aspect; J, frontal depression and protuberance, latero-dorsal aspect.

FIGURE 25. Wroughtonia varifemora sp. nov., ♀, holotype. A, wings; B, mesosoma, lateral aspect; C, metasoma, dorsal aspect; D, hind femur and tibia, lateral aspect; E, middle femur, lateral aspect; F, fore femur, lateral aspect; G, habitus, lateral aspect; H, head, lateral aspect; I, head, dorsal aspect; J, mesosoma, dorsal aspect.

FIGURE 26, Wroughtonia yaanensis sp. nov., ♀, holotype. A, fore wing; B, hind wing; C, mesosoma, lateral aspect; D, first-second metasomal tergites, dorsal aspect; E, hind femur and tibia, lateral aspect; F, habitus, lateral aspect; G, head, lateral aspect; H, head, dorsal aspect; I, propodeum, medio-dorsal aspect.