Porricondyla pallidigenae, Jaschhof & Jaschhof, 2020

Jaschhof, Mathias & Jaschhof, Catrin, 2020, The Porricondylini (Diptera: Cecidomyiidae: Porricondylinae) of Sweden revisited: descriptions of nineteen new species of ten genera, including a new genus, Zootaxa 4728 (2), pp. 151-182 : 173

Porricondyla pallidigenae sp. nov.

Figs 42–43

Diagnosis. This small-size Porricondyla is another member of species group C. The gonostylar tooth of P. pallidigenae is serrate ( Fig. 42 , ↓ 2), a peculiarity shared only with P. serrulata Jaschhof , also in group C, and no other Porricondyla . Unlike in P. serrulata ( Jaschhof & Jaschhof 2013: fig. 113), the gonocoxal emargination of P. pallidigenae is strongly and extensively sclerotized basally (↓ 3); the medial gonocoxal bridges are unmodified (i.e. lack protuberances with thick setae); and the parameres are thin and entire apically (↓ 4) (not broadened and serrate). The extensively unpigmented genae giving P. pallidigenae the name provide an eye-catching feature, which is similarly found in several other Porricondyla , though. Females and larvae of this new species are unknown.

Other male characters. Body size 1.2–1.4 mm. Head. Eye bridge 1–2 ommatidia long at vertex. Antenna longer than body. Scape and pedicel concolorous with flagellum. Circumfila on flagellomeres 1–7. Neck of fourth flagellomere 1.2 times as long as node ( Fig. 43 ). Palpus slightly longer than head height, 4 setose, subcylindrical segments, apical segment longest of all. Thorax. No scutal windows. Pronotum with 1–4 setae; anepisternum nonsetose; anepimeron with 1–3 setae. Wing about as long as body. Length / width ratio 3.2. Setae on proximal half of membrane sparse. M 1+2 absent. M 4 extremely weak. Hind fork incomplete. Legs. Foreleg: femur 0.9 times the length of tibia, tibia 0.6 times the length of T 2. Claws toothless. Empodia broad, slightly longer than claws. Genitalia ( Fig. 42 ). Gonocoxal synsclerite: setae largely confined to ventrolateral and dorsal portions; ventral emargination small, shallow, bordered by weak, membranous processes; dorsal apodemes short. Gonostylus dorsoventrally flattened, outline subtriangular seen from ventrally; 1–2 long bristles subbasally. Parameres crossing each other apically; a narrow transverse bridge subbasally. Aedeagal apodeme nearly as long as gonocoxae; rodlike portion moderately sclerotized; apical portion weakly sclerotized, broadened, with indistinct contours. Ducts of accessory glands barely discernible.

Etymology. The species epithet, meaning ̒with pale cheeks’, refers to the partially unpigmented head capsule found in this species.

Type material. Holotype. Male , Sweden, Småland, Alsterbro, backyard, 15 July–19 August 2016, Malaise trap, M. & C. Jaschhof & S.-O. Ulefors (spn. CEC 2281 in NHRS) . Paratypes. 1 male, same data as the holotype (spn. CEC 2282 in SDEI) ; 1 male, Sweden, Östergötland, Odeshög, Omberg, Stocklycke äng, meadow on limestone, 25 May–8 June 2003, MT, SMTP (trap 13, collecting event 909) (spn. CEC 2283 in NHRS) ; 1 male, Sweden, Öland, Mörbylånga, Skogsby , Station Linné , swampy meadow next to willow scrub, 5 July–4 August 2016, MT, MCJ & E. Gustavsson (spn. CEC 2284 in SDEI) .

Figures

FIGURES 40–43. Male morphology of Porricondyla ottenbyensis (40–41) and Porricondyla pallidigenae (42–43), holotypes. 40: Fourth flagellomere, lateral. 41: Genitalia, ventral. 42: Genitalia, ventral. 43: Fourth flagellomere, lateral. Scales 0.05 mm. Arrows refer to characters described in the diagnoses.