Coelophthinia cirra Kerr, 2023

Kjaerandsen, Jostein, Kerr, Peter H., Lindemann, Jon Peder & Kurina, Olavi, 2023, When details matter: Integrative revision of Holarctic Coelophthinia Edwards (Diptera, Mycetophilidae), including mapping of its mitogenome, leads to the description of four new pseudocryptic species, Biodiversity Data Journal 11, pp. 98741-98741 : 98741

Coelophthinia cirra Kerr sp. n.

Materials

Type status: Holotype. Occurrence: catalogNumber: 12J162 ; recordedBy: Peter H. Kerr; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: 7D202C2D-CD98-5304-B629-4F12A82473A2; Taxon: scientificName: Coelophthinia cirra; order: Diptera ; family: Mycetophilidae ; genus: Coelophthinia ; specificEpithet: cirra; scientificNameAuthorship: Kerr ; Location : country: United States; countryCode: USA; stateProvince: California; county: Alpine ; locality: Grover Hot Springs State Park , nr. Hoffman house ; verbatimLocality: Grover Hot Springs State Park ; verbatimElevation: 1800m; verbatimCoordinateSystem: decimal degrees; decimalLatitude: 38.6952; decimalLongitude: -119.838; Identification: identificationID: 12J162; identifiedBy: Peter H. Kerr; dateIdentified: 6/9/2021; Event: eventID: CSCA06LOT476; samplingProtocol: Malaise Trap (2m); eventDate: 14.viii-3.ix.2006; Record Level: type: PreservedSpecimen; collectionID: urn:lsid:biocol.org:col:33156; institutionCode: CSCA; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: 11G757 ; recordedBy: Peter H. Kerr; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: 7F33526F-B6AA-5F25-BD93-54915F0936EB; Taxon: scientificName: Coelophthinia cirra; order: Diptera ; family: Mycetophilidae ; genus: Coelophthinia ; specificEpithet: cirra; scientificNameAuthorship: Kerr ; Location : country: United States; countryCode: USA; stateProvince: California; county: Alpine ; locality: Grover Hot Springs State Park , forest/meadow edge ; verbatimLocality: Grover Hot Springs State Park ; verbatimElevation: 1800m; verbatimCoordinateSystem: decimal degrees; decimalLatitude: 38.6996; decimalLongitude: -119.8457; Identification: identificationID: 11G757; identifiedBy: Peter H. Kerr; dateIdentified: 6/9/2021; Event: eventID: CSCA06LOT275; samplingProtocol: Malaise Trap (2m); eventDate: 11-25.v.2006; Record Level: type: PreservedSpecimen; collectionID: urn:lsid:biocol.org:col:33156; institutionCode: CSCA; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: 21K109 ; recordedBy: Peter H. Kerr; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: 46672366-0481-5138-BA92-938A553F2952; Taxon: scientificName: Coelophthinia cirra; order: Diptera ; family: Mycetophilidae ; genus: Coelophthinia ; specificEpithet: cirra; scientificNameAuthorship: Kerr ; Location : country: United States; countryCode: USA; stateProvince: California; county: Alpine ; locality: Grover Hot Springs State Park , nr. Hoffman house ; verbatimLocality: Grover Hot Springs State Park ; verbatimElevation: 1800m; verbatimCoordinateSystem: decimal degrees; decimalLatitude: 38.6952; decimalLongitude: -119.8380; Identification: identificationID: 21K10; identifiedBy: Peter H. Kerr; dateIdentified: 6/9/2021; Event: eventID: CSCA06LOT275; samplingProtocol: Malaise Trap (2m); eventDate: 11-25.v.2006; Record Level: type: PreservedSpecimen; collectionID: urn:lsid:biocol.org:col:33156; institutionCode: CSCA; basisOfRecord: PreservedSpecimen

Description

Male. (Fig. 13 Fig. 14)

Coloration and most body characteristics as in the genus description. Body length 2.8 mm. Wing length 2.5 mm; ratio of length to width 2.8. Sensory organ dorsally on the basal half of mid-tibia elongate oval, 3.5 times longer than wide, 0.1 times of tibial length.

Terminalia. Tergite 9 setose, except basal fifth medially, nearly quadrate, slightly longer than wide, medially slightly constricted, basal margin concave, curved like a hood in lateral view. Medial protrusion of tergite 10 a modest bulbous swelling, densely setose. Cerci medially fused except for apical third, setose. Gonocoxites in lateral view with a rounded outline, with straight ventral margin. Posterolateral lobe of gonocoxite long, slender and evenly tapering, about 3-4x longer than wide at apex, rounded apically, laterally bare for about 2x apical width. Gonocoxal lobe spathulate, long and slender, with slightly constricted base, basad of base lacking row of distinctive setae, evenly tapered along dorsoventral margins, external surface bare on apical length approximately 3x apical width, apically rounded. Dorsal branch of gonostylus small, simple, oblong and semicircular, dorsally setose. The broad v br lobe 1 fan-shaped, carrying 6-7 blunt-tipped setae along rim and 4 normal setae on inner side. The narrow, acute tipped v br lobe 2 with a tiny, practically imperceptible seta subapically; with thickened, sharp-tipped seta at approximately its length from the apex of the lobe, followed by one fine seta of similar length and one longer seta near the base of the lobe approximately 1.5x length of v br lobe 2. Aedeagus of same length or slightly longer than spathulate gonocoxal lobe in lateral view, slender throughout, ventral margin entire along length, without rounded cutout at base of hook; with long apical down-curved hook, hook approx. 0.4x length of rest of aedeagus; with a short, inconspicuous sessile projection at the outer side below the curving point of hook.

Diagnosis

Coelophthinia cirra can be distinguished from other species of the genus by the combination of the following features: mid-tibial organ only slightly longer than width of tibia, gonocoxal lobe situated along straight ventral margin, slender and evenly rounded apically, thinner than constricted length; aedeagus slender throughout, without cutout at base of apical hook.

Etymology

Adjective taken from the Latin word cirrus, meaning "tendrils" and descriptive of clouds. This species is thin and leggy and found at high elevations, reminiscent of the clouds.