Crassignatha quadriventris (Lin & Li, 2009) Li & Lin & Li, 2020

Li, Ya, Lin, Yucheng & Li, Shuqiang, 2020, A review of Crassignatha (Araneae, Symphytognathidae), ZooKeys 988, pp. 63-128 : 63

publication ID	https://dx.doi.org/10.3897/zookeys.988.56188
publication LSID	lsid:zoobank.org:pub:6E64D69B-DD73-4A7E-AE2B-3CD21247A5E3
persistent identifier	https://treatment.plazi.org/id/EAF96456-8E8A-5479-9AD3-4A848E56144A
treatment provided by	ZooKeys by Pensoft
scientific name	Crassignatha quadriventris (Lin & Li, 2009)
status	comb. nov.

Treatment

Crassignatha quadriventris (Lin & Li, 2009) View in CoL comb. nov. Figs 20 View Figure 20 , 21 View Figure 21 , 38 View Figure 38

Patu quadriventris Lin & Li, 2009: 55, figs 7A, B, 8A, B, 9A-E, 10A, B (♂♀).

Type material.

Holotype ♂ and paratypes 2♂ 2♀ (IZCAS), China: Hainan Province, Wuzhishan City, Mt. Wuzhishan Nature Reserve (18.90000°N, 109.65000°E), 9.VIII.2007, S. Li and C. Wang leg.; paratype 1♀ (IZCAS), China: Hainan Province, Qiongzhong County, Limushan Nature Reserve (19.18333°N, 109.73333°E; 655 m), 12.VIII.2007, S. Li and C. Wang leg. Examined.

Other material examined.

1♂ 4♀ (IZCAS-Ar 41030-41034), China: Hainan Province, Dongfang City, Donghe Town, Nanlang Village, at the foot of Exianling Mountain (19.00633°N, 109.08383°E; 214 m), 16.XII.2014, Q. Zhao and L. Shao leg.; 1♀ (NHMSU-HA025) used for sequencing, GenBank: MT991990, same data as for preceding.

Diagnosis.

This species differs from other congeneric species except C. baihua sp. nov. by the short, stiff embolus (Fig. 21A, B View Figure 21 ). It is most similar to C. baihua sp. nov. in the form of the palp and the vulva configuration but can be easily distinguished by the sloped embolic apex, rather than flat as in the latter, and by having six twists of the copulatory ducts, rather than four in the latter (Figs 21A, B, F, G View Figure 21 , 2A, B, F, G View Figure 2 ).

Description.

Male (IZCAS-Ar 41030). Total length 0.80. Carapace 0.36 long, 0.36 wide, 0.40 high. Clypeus 0.16 high. Sternum 0.28 long, 0.24 wide. Abdomen 0.52 long, 0.48 wide, 0.52 high. Length of legs: I 1.34 (0.42, 0.14, 0.34, 0.20, 0.24); II 1.10 (0.30, 0.14, 0.26, 0.16, 0.24); III 0.90 (0.24, 0.12, 0.18, 0.14, 0.22); IV 1.02 (0.30, 0.12, 0.22, 0.18, 0.20).

Somatic characters (Fig. 20A-C View Figure 20 ). Coloration: prosoma brown. Legs pale brown. Abdomen dorsally dark, with pale stripes and speckles, ventrally pale brown. Prosoma: carapace sub-rounded, granular, sculptured, with two cephalic setae. Cephalic area elevated. Clypeus concave. ALE protruded. PER strongly recurved. Chelicerae covered with few setae anteriorly. Labium tongue shaped, wider than long. Sternum heart shaped, surface rough, slightly plump, truncated posteriorly. Legs: tibia II with two clasping spurs. Abdomen: anteriorly round, nearly square posteriorly, with sparse, long setae, lateral scutum distinct. Spinnerets slightly sclerotized, surrounded by a circular plate.

Palp (Fig. 21A-C View Figure 21 ): tibia with four distal, dorsal, short setae. Cymbium bears a few distal setae. Apical cymbial tooth hook shaped. Tegulum broad, smooth, globular. Plate-like median apophysis with a sclerotized, longitudinal central ridge line and a prolaterally odontoid process. Embolic membrane arises near anterior part of median apophysis. Embolus short, stiff, basally constricted, mesally wide, distally coracoid.

Female (IZCAS-Ar 41031). Total length 1.04. Carapace 0.44 long, 0.40 wide, 0.40 high. Clypeus 0.14 high. Sternum 0.28 long, 0.28 wide. Abdomen 0.72 long, 0.68 wide, 0.80 high. Length of legs: I 1.72 (0.60, 0.18, 0.40, 0.26, 0.28); II 1.42 (0.44, 0.16, 0.34, 0.20, 0.28); III 0.98 (0.26, 0.10, 0.22, 0.16, 0.24); IV 1.18 (0.34, 0.12, 0.28, 0.18, 0.26).

Somatic characters (Fig. 20D-F View Figure 20 ). Coloration: prosoma and legs as in male. Abdomen dorsally darker than in male, ventrally lighter. Prosoma: modification and arrangement of eyes as in male, cephalic area lower than in male. Clypeus slightly concave. Mouthparts and sternum as in male. Abdomen: anteriorly round and posteriorly square, surface bears sparse, long setae, lateral scutum and circular plate absent. Spinnerets slightly sclerotized.

Epigyne (Fig. 21D-F View Figure 21 ): epigynal area weakly sclerotized, bears a few setae. Scape developed, protruded, longer than wide. Copulatory openings located at terminus. Internal structures faintly visible via translucent tegument. Paired spermathecae globose, widely separated by at least 1.5 × their diameter. Fertilization ducts originating from the lower inside margin of spermathecae, bent downwards and laterally. Copulatory ducts long, connected to dorsal surface of spermathecae, from below spermathecae to the center of vulva, making six bends, then reaching copulatory openings.

Taxonomic justification.

The shape of the male palps, the configuration of the epigyne, the modified carapace, and the male abdominal scutum and clasping spurs on tibia II leave no doubt that this species is a member of Crassignatha and not Patu . Therefore, we propose a new combination, C. quadriventris (Lin & Li, 2009) comb. nov., transferring it from Patu .

Distribution.

China (Hainan) (Fig. 38 View Figure 38 ).