Bifidocoelotes, WANG, 2002

WANG, XIN-PING, 2002, A Generic-Level Revision Of The Spider Subfamily Coelotinae (Araneae, Amaurobiidae), Bulletin of the American Museum of Natural History 2002 (269), pp. 1-150 : 37-38

BIFIDOCOELOTES , NEW GENUS

TYPE SPECIES: Coelotes bifida Wang, Tso and Wu, 2001

ETYMOLOGY: The generic name derives from its similarity to Coelotes and from the bifurcated epigynal teeth of females, and is masculine in gender.

DIAGNOSIS: Females can be easily recognized by their bifurcated epigynal teeth (fig. 86); males have a broad, bifid conductor and a small, rounded spoonlike median apophysis (figs. 88, 89). Both sexes have three promarginal and two retromarginal cheliceral teeth (fig. 91).

DESCRIPTION: See description of type species (below).

DISTRIBUTION: China (map 3).

COMPOSITION: Two species are included. Both of them are new combinations.

1. Bifidocoelotes bifida ( Wang, Tso and Wu, 2001) : female holotype and male paratype from Nantou, Taiwan, in THU, examined. NEW COMBINATION (transferred here from Coelotes ).

2. Bifidocoelotes primus (Fox, 1937) : female holotype and female paratype from Hong Kong, China, in AMNH and USNM, examined. NEW COMBINATION (transferred here from Coelotes ).

Bifidocoelotes bifida

( Wang, Tso, and Wu, 2001)

Figures 86–100

Coelotes bifida Wang, Tso and Wu, 2001: 128 , figs. 1–4 (female holotype and male paratype from Nantou, Taiwan, in THU, examined).

DIAGNOSIS: Females differ from B. primus by the anteriorly situated copulatory openings, the different shape of bifurcated epigynal teeth, the large copulatory ducts, the short spermathecal heads, and the longitudinally elongated spermathecae (figs. 86, 87), males have the broad, bifid conductor and small, rounded spoonlike median apophysis (figs. 88, 89).

DESCRIPTION: Total length 4.00 –9.50. From front, anterior eye row slightly procurved, posterior row procurved; eye sizes and arrangements: AME smallest, about half ALE size or smaller, ALE largest, PLE slightly smaller than ALE, PME slightly smaller than PLE; AME­AME slightly short­ er than their diameter, AME­ALE and ALE­ PLE close together, PME­PLE almost subequal to PME­PME or slightly longer, about size of PLE radius. Clypeal height slightly more than twice AME diameter; chilum divided, elongated (fig. 90). Chelicerae with three promarginal and two retromarginal teeth (fig. 91). Labium slightly shorter than wide. Length of female 1st leg patella + tibia shorter than carapace length.

Tarsal organ situated close to distal end of tarsus, anterior of distalmost trichobothrium (figs. 98–100). Apex of ALS with 2 major ampullate gland spigots (MAP), 8–13 piriform gland spigots (PI) in both sexes; PMS with 7–10 aciniform gland spigots (AC) in both sexes, 2 cylindrical gland spigots (CY) in female, both sexes PMS with minor ampullate gland spigots (mAP) not apparent; PLE with 7–11 aciniform gland spigots in both sexes and 2 cylindrical gland spigots in female (figs. 92–97).

Female epigynum with single bifurcated epigynal teeth; atrium small, situated anteriorly; copulatory ducts large; spermathecal heads situated mesad of spermathecae; spermathecae elongated, convoluted, widely separated, with indistinct stalks and bases (figs. 86, 87). Male palp with large patellar apophysis; RTA long, with distal end broad, not extended beyond tibia; lateral tibial apophysis small; cymbial furrow long, with dorsal edge strongly concave and distal end slightly extend beyond cymbium; conductor strong, more or less spiral, with bifurcated apex; conductor dorsal edge with broad membranous extension; conductor dorsal apophysis small; conductor lamella large; embolus posterior in origin, very long; median apophysis spoonlike, small, with distal end smooth (figs. 88, 89).

MATERIAL EXAMINED: CHINA: Taiwan: Nantou County, Huei­Sun Experimental Forest , elevation 1680 m, March 31, 1998, female holotype (Hai­Yin Wu; THU, THU­Ar­ 990017) ; Nantou County, Huei­Sun Experimental Forest , March 31, 1998, 1 male par­ atype (Hai­Yin Wu; THU, THU­Ar­990020) ; Nantou County, Huei­Sun Experimental Forest , elevation 1680 m, March 31, 1998, 1 male (Hai­Yin Wu, THU, THU­Ar­990019) ; Nantou County, Huei­Sun Experimental For­

Figures

Figs. 86, 87. Bifidocoelotes bifida (Wang, Tso and Wu), female. 86. Epigynum. 87. Vulva.

Figs. 88–91. Bifidocoelotes bifida (Wang, Tso and Wu). 88. Male palp, ventral view. 89. Male palp, retrolateral view. 90. Female, eyes, front view. 91. Female, chelicera, ventral view.

Figs. 92–94. Bifidocoelotes bifida (Wang, Tso and Wu), female spinnerets, ventral view, right. 92. ALS. 93. PMS. 94. PLS. Abbreviations are spelled out on p. 6.