Calendula suffruticosa, Vahl (1791: 94)

Gonçalves, Ana Carla, Ouhammoud, Ahmed, Amirouche, Rachid, Santos, Conceição, Figueiredo, Estrela & Silveira, Paulo, 2023, A taxonomic revision of Calendula (Asteraceae) in Morocco, including some taxa from Algeria and Tunisia, Phytotaxa 605 (1), pp. 1-83 : 80-82

C. suffruticosa Vahl (1791: 94)

subsp. foliosa Batt. in Battandier & Trabut (1902: 479) A.C. Gonç & P. Silveira, comb. nov.

Additional specimens examined: ALGERIA. Bou Zecza, April 1880, Battandier, J . A . s.n. ( G!); Gorges de Palestro , 6 May 1884, Battandier, J . A . 189 ( BR!); Alger, Jardin ou il es naturalisé, 1885, Battandier, J . A . 4 572 ( G!); 1885,

Battandier, J. A. s.n. ( G!); Bou Zecza , 1887, Battandier , J. A. 263 ( G!); Gorges de Palestro , 20 May 1889, (Chabern) s.n. ( G!); 20 May 1889, (Chabern) s.n. ( G!); culta in horto meo seminibus a mi lectis in loco classico: faucibus Palestro, May 1895, unknown s.n. ( G!); Gorge de Palestro, 100 m [36°36’ N, 3°35’ E], 7 June 2013, Silveira , P.; Gonçalves , ACRS; Amirouche , R. 3320 ( AVE!);

subsp. fulgida (Rafiniesque) Guadagno (1922: 64)

Additional specimens examined: MOROCCO. Oriental: Beni Snassen Mountains , near Jbel Tizi, 1200 m, [34°48’ N, 2°20’ W], 20 February 1995, Jury, S. L .; Lafkih, M. A .; Abu Sbaih, H .; Ziri, R . 15795 ( SEV!); Beni Snassen, gargantas Zegzel hacia Aroujene , 460 m, [34°48’ N, 2°20’ W], 13 April 1994, Romo, A. M .; Bouhmadi, M .; Peris, J. B .; Stubing, G . 6637 ( BM!); Beni Snassen , [34°48’ N, 2°20’ W], 28 May 1935, Sennen, F .; Mauricio, Hno. s.n. ( MA!); Beni Snassen, pista a 6.5 km de Zegzel y hacia Ain- Almou , 1050 m, [34°48’ N, 2°20’ W], 3 May 2000, Vargas, P .; Fiz, O .; Martínez, J .; Valcárcel, V . 201 PV ( MA!); Berkane, marges Beni Snassen , [34°48’ N, 2°20’ W], 2 April 1938, Sennen, F .; Mauricio, Hno. s.n. ( BM!); Beni Snassen, Zegzel, ad ruper supra Laonia Moulay Achmed , [34°48’ N, 2°20’ W], 19 April 1928, Wilczek, E .; Briquet, J .; Dutoit, D. 1020 ( K!); Beni Snassen , lieux herbeux de la vallée de l’Oued Zegzel, 300 m, [34°48’ N, 2°20’ W], 16 April 1928, Emberger, L . s.n. ( RAB!); Beni Snassen, eboulis calcaires dans la vallée d l’Oued Zegzel , 300 m, [34°48’ N, 2°20’ W], 16 April 1928, Emberger, L . s.n. ( RAB!); 10 km S of Berkane Grottte du Chameau , 360 m, [34°48’ N, 2°20’ W], 5 March 1994, Upson, T. M .; Lafkih, M. A .; Hassan, M .; Walters, G. S . 13943 ( RNG!); Beni Snassen , rocailles calc. herbeux de la vallée de l’Oued Zegzel, 300 m, [34°48’ N, 2°20’ W], 16 April 1928, Emberger, L . s.n. ( RAB!); environ de Berkane, Massif des Beni Snassen. Montagnes de Iasil à l’ouest du refuge du Zegzel , 400- 900 m, [34°48’ N, 2°20’ W], 20 April 1928, Wilczek, E .; Briquet, J .; Dutoit, D. 1103 ( G!); environs de Berkane, Massif des Beni Snassen, Gorges inf. de Zegzel, 150- 300 m, [34°48’ N, 2°20’ W], 16 April 1928, Wilczek, E .; Briquet, J .; Dutoit, D. 867 ( G!); Plaine subaride entre Oujda (22 km W) et Sidi Bouhria , versant S de Beni Snassen , 550 m, [34°48’ N, 2°20’ W], 9 April 2007, Dobignard, A. AD 14237 ( G!); Cabo de Agua , [35°02’59” N, 2°33’52” W], April 1912, Caballero, A . s.n. ( MA!); Kebdana , [35°02’59” N, 2°33’52” W], 12 June 1934, Sennen, F .; Mauricio, Hno. s.n. ( MAF!); Nador, Monts des Kebdana , road P 27 between Nador and Berkane , c. 6 km W Zaio road embankments, 240 m, [35°02’59” N, 2°33’52” W], 4 May 1995, Vogt, R .; Oberprieler, Ch. 14141 ( RAB!); Oujda, between Oujda and Ahfir , Col de Guerbouss , 530 m, [34°40’53” N, 1°54’31” W], 30 May 1993, Lafkih, M. A .; Mateos, M. A .; Valdés, B. BV 461 93 ( RNG, SEV!); Monts des Beni Snassen, Taforalt, N-facing slopes of Jbel Achaoun c. 1 km E Taforalf, stony slopes and limestone rocks, 850- 900 m, [34°48’ N, 2°20’ W], 4 May 1993, Vogt, R .; Oberprieler, Ch. 10756 ( RAB!); Berkane, Montes de Beni Snassen, estrada Zegzel entre Tazarhine e Takerboust , 54 m, [34°50’01” N, 2°22’17” W], 11 May 2014, Silveira, P .; Gonçalves, A. C. R. S . 3335 ( AVE!); Berkane, Montes de Beni Snassen, estrada Zegzel proximo do Oued Zegzel , 260 m, [34°52’40” N, 2°21’20” W], 11 May 2014, Silveira, P .; Gonçalves, A. C. R. S . 3336 ( AVE!); Beni Snassen, Zegzel , [34°48’ N, 2°20’ W], Emberger, L . s.n. ( RAB!); Beni Snassen, vallée du Zegzel , [34°48’ N, 2°20’ W], 16 April 1928, Emberger, L . s.n. ( RAB!); Beni Snassen , [...] au NW du Zegzel, 600 m, [34°48’ N, 2°20’ W], 20 April 1928, Emberger, L . s.n. ( RAB!);

subsp. lusitanica (Boissier) Ohle (1974: 270)

Additional specimens examined: SPAIN. Ceuta: Benzu, base del Jbel Musa , [35°55’ N, 5°24’ W], 2 January 1990, Cabezudo, B .; Nieto , J. M .; Perez-Latorre , AV s.n. ( MGC!); Benzu , 5–20 m, [35°54’06” N, 5°20’44” W], 14 June 2012, Silveira, P .; Gonçalves, ACRS 3 269 ( AVE!);

subsp. monardii (Boissier & Reuter 1859: 108) Ohle (1975b: 532)

Additional specimens examined: ALGERIA. Alger: Paroi N du Gran Rocher, Forêt de Bainem, près d’Alger , 19 April 1960, Dubuis, A .; Faurel, L . 4496 ( G!); Alger, 21 April 1861, Lefebre, J . 186 ( BM, BM813733 !); environ d’ Alger, 1851, Jamin, P . s.n. ( BM!); Alger, 1844, unknown s.n. ( K!); Fauché , M . s.n. ( G!); sur la montagne d’Aïn- Telazi, près Blidah, March 1849, Reuter, s.n. ( G!); Le Gouraya de Bougie , 600 m, June 1896, Reverchon, E . 62 ( G!); La Calle (El Kala), October 1924, Cuenod, A . s.n. ( G!); Guyoteville, près d’Alger, May 1919, Alleizette, Ch. d’ s.n. ( MA!); Dep. d’Alger , dunes littorales près de Guyotville , 11 May 1947, Dubuis, A .; Faurel, L . 230 ( RAB, K, G!); Guyotville , 5 December 1910, Brichi, M . s.n. ( G!); In arenosis litoralis prope Guyotville , 2 June 1935, Maire, R . s.n. ( G!); sables maritimes, La Madragne, 20 Km W Alger, 27 March 1960, Charpin, A . s.n. ( G!); Ligne de routes entre le Lalla Khedidja et le Col de Lirourda , 26 June 1875, Duhamel, M . s.n. ( G!); a Sidi Ferruch , aux environs d’ Alger, 24 March 1867, Durando 582 ( BM!); Sidi Ferruch, province d’Alger, May 1884, Allard s.n. ( MA!); Sidi Ferruch, near Algier, April 1856, unknown s.n. ( K!); Sidi Ferruch, July 1830, Monard, P .; Monard, C . 10 ( G!); February 1882, Meyer, A .; Allard, C . 3362 ( G!); March 1886, Battandier, J. A .; Trabut, L . 81 ( G!); In arenosis prope Sidi Ferruch , 10 March 1936, Maire, R . s.n. ( G!); Sidi Ferruch, July 1830, unclear 42 ( G!); Zeralda , 28 May 1939, Maire, R . s.n. ( K!); 28 May 1939, Maire, R . s.n. ( K!);

subsp. suffruticosa Vahl (1791: 94)

Additional specimens examined: TUNISIA. Tunis: Porto Farina, Vahl s.n. ( K!); Korbous , W side of Cap Bon Peninsula, 10–50 m, [37°05’18” N, 11°02’09” E], 27 April 1975, Davis ; Lamond D 56 824 ( BM!); Tunis, Hammam-Lif, base du Cap Bon, côte Nord, 15–20 m, [36°43’59” N, 10°20’10” E], 12 April 1999, Lambinon, J .; Margot , J . 99 Tu 176 ( MA, LG!); Ghar el Melh [Porto Farina], 10 m [37°10’ N, 10°11’ E], 11 April 2009, Silveira , P . 3 038 ( AVE!);

subsp. trialata P. Silveira & A.C. Gonç.

Additional specimens examined: MOROCCO. Rif: Tanger Peninsula, Cap Ras Ciress, surroundings of the lighthouse 9.7 km NE of Ksar-es-Seghir , coastal rocks, moist place in Pinus woodland, 5-60 m, 20 April 1993, Vogt, R .; Oberprielier, Ch. 9942 ( RAB!) ;

subsp. tunetana (Cuénod) A.C.Gonç. & P.Silveira

Additional specimens examined: TUNISIA. Nabeul: Oued el Kébir, May 1910, Pitard, C . J . 706 ( MA!); Nabel , October 1907, Gandonger, M . 147 ( COI, K, W, G!); entre Bir Bou Regba e Hammamet , [36°26’ N, 10°35’ E], 11 April 2009, Silveira, P . 3039 d ( AVE!); Berges de l’Oued entre Bir bou Rekha et Hamamet, May 1906, Cuenod, A . s.n. ( G!); Bir bou Rekba , April 1913, Cuenod, A . s.n. ( G!); Hammamet (Oued El Faouara), March 1936, Cuenod, A . s.n. ( G!); Bir bou Rekba , May 1909, Cuenod, A . s.n. ( G!); Nabeul , Oued el Kébir, January 1918, Pitard, C . J . s.n. ( G!); May 1910, Pitard, C . J . 706 ( MA, G!); sine loco in herb. Cuenod , Cuenod, A . s.n. ( G!); Bir bou Rekba, 1910, Cuenod, A . s.n. ( G!);