Dyrana flexuosa (NEWBERRY) GOLOVNEVA

Manchester, Steven R., 2014, Revisions To Roland Brown’S North American Paleocene Flora, Acta Musei Nationalis Pragae Series B 70 (3 - 4), pp. 153-210 : 166-167

Dyrana flexuosa (NEWBERRY) GOLOVNEVA

This species is based on leaves found initially in the late Paleocene or early Eocene Chuckanut Formation of coastal Washington, and subsequently recognized to have been distributed in the Rocky Mountain region, treated as Dicotylophyllum flexuosa (NEWBERRY) J. WOLFE (1966) . Golovneva (1994b) established the genus Dyrana based on specimens from the Paleogene of the Koryak Highlands of NE Russia, and subsequently transferred the North American species to the same genus (Golovneva, L. B. 2000, Budantsev and Golovneva 2009). Budantsev and Golovneva also illustrated numerous examples of D. flexuosa from the early Paleocene Barensburg flora of Spitsbergen. The leaves are elliptical to ovate, with pinnate craspedodromous secondary veins and teeth with rounded sinuses. As mentioned above, these leaves were treated as Quercus sullyi NEWBERRY by Brown (1962) and Meliosma longifolia (HEER) HICKEY 1977 . The attribution to Platanaceae is supported by glandular teeth with rounded sinuses, petioles with expanded bases, and the orthogonal pattern of higher order venation, but epidermal characters remain unknown and associated reproductive structures have not been clearly demonstrated.

Trochodendraceae