Toxophorinae, Schiner, 1868

Greathead, D., Lovell, S., Barraclough, D., Slotow, R., Hamer, M. & Herbert, D., 2006, An ecological and conservation assessment of the fauna of Bombyliidae (Diptera) occurring in the Mkhuze, Phinda and False Bay reserves, KwaZulu-Natal, South Africa, African Invertebrates 47, pp. 185-206 : 204-206

Toxophorinae : Systropodini Anthracinae : Anthracini

Systropus zuluensis Hesse, 1938 (xii.1911) Anthrax hemimelas Speiser, 1910 (xii.1911)

Anthrax pitheceus Fabricius, 1805 (x.1911)

Bombyliinae : Bombyliini Anthrax triatomus Hesse, 1956 (v.1916) Bombylella delicata (Wiedemann, 1830) (iii.1911)* Spogostylum robustale (Hesse, 1956) (iv–v.1934), Bombylella elegans (Wiedemann, 1828) (xii.1911) possible synonym of S. incisurale Bombylella kilimanjarica (Speiser, 1910) (iii.1911) Bombylella rufiventris (Macquart, 1846) (xii.1911) Anthracinae : Exoprosopini Bombylisoma cinereitinctum (Hesse, 1938) (iv– Atrichochira pediformis (Bezzi, 1921) (iii.1917) v.1934, xii[sic]– ii.1935) Exoprosopa heros (Wiedemann, 1819) (xii.1911) Eurycarenus laticeps (Loew 1852) (xii.1914) Heteralonia (Homolonia) neurospila (Bezzi, 1921) Systoechus bombycinus Hesse, 1938 (iii.1914, (ii.1914) iii.1916)* Systoechus deceptus Hesse, 1938 (n.d.) Anthracinae : Villini Systoechus inordinatus Hesse, 1938 (v.1917) Exhyalanthrax abruptus (Loew, 1860) (iii.1914) Systoechus neglectus Hesse, 1938 (xii.1914) Systoechus silvaticus Hesse, 1938 (n.d.)* Anthracinae : Xeramoebini

Petrorossia angustibasalis Hesse, 1956 (iii.1914,

Mariobezzinae vi.1916, vi– v.1931, xii.1916) Hyperusia minor Bezzi, 1921 (iv.1916) Petrorossia fulvipes (Loew, 1860) (n.d.)

Additional Bombyliidae recorded by Hesse and Greathead (DG) from nearby localities. Hesse often did not give detailed localities, but simply ‘Zululand’. Lowland species so designated are included.

Bombyliinae : Bombyliini

Toxophora coeruliventris Karsch, 1887 : ‘Zululand’ Bombylisoma rhodesianum Hesse, 1938 : Umfolozi Bombomyia discoidea (Fabricius, 1794) : Reserve (xi.1992, DG)

‘Zululand’* Gonarthrus leucomelas Bowden, 1975 : ‘Zululand’ Bombylella kilimandjarica (Speiser, 1910) :

Anthracinae : Anthracini

Hluhluwe G.R. (xi.1967), Charter’s Creek (Lake

St Lucia, xi.1967) (in BMNH, DG) Anthrax aygulus Fabricius, 1805 : ‘Zululand’ Anthracinae : Exoprosopini L. paris is doubtfully a distinct species Exoprosopa albata Bezzi, 1924 : Manguzi River Litorhina nyasae (Ricardo, 1901) : Hluhluwe (nr Maputa, xi–xii.1945), Imyalazi River Reserve (iv.1951)*

(xii.1928), Hluhluwe Reserve (i.1950) Litorhina obumbrata (Bezzi, 1924) : ‘Zululand’* Exoprosopa brachipleuralis Hesse, 1956 : Pon- Micomitra perlucida (Hesse, 1956) : Manguzi River gola River (x.1929), probably a synonym of (nr Maputa, xi–xii.1945), Hluhluwe Reserve E. luteicosta (i.1950)*

Exoprosopa dimidiata Macquart, 1846 : Anthracinae : Villini

‘Zululand’* Exhyalanthrax abruptus (Loew, 1860) : Hluhluwe Exoprosopa major Ricardo, 1901 : ‘Zululand’* & Umfolozi reserves (Fiedler & Kluge (1954)) Exoprosopa luteicosta Bezzi, 1921 : Manguzi River Exhyalanthrax flammiger (Walker, 1849) : (nr Maputa xi–xii.1945), Shangawana (x.1938) ‘Zululand’

Exoprosopa morosa Loew, 1860 : ‘Zululand’* Exhyalanthrax lugens (Loew, 1860) : Hluhluwe Heteralonia (Acrodisca) katonae (Bezzi, 1924) : St (viii.1947, viii.1948), Umfolozi (viii.1947) Lucia Bay (x.1920), Shangawana (x.1938) Reserves (also Fiedler & Kluge (1954))

Ligyra nigripennis (Loew, 1852) : ‘Zululand’ Exhyalanthrax transiens (Bezzi, 1921) : Ligyra vittata (Ricardo, 1901) : ‘Zululand’ ‘Zululand’*

APPENDIX 3

Bombyliidae collected in Shimba Hills Nature Reserve, Kenya, 1962–82 and species from similar reserves on the Kenya coast that may be expected to be found there. Months are given as Roman numerals. The dry seasons are December–March and June–September. The term ‘savanna’ is used to indicate drier savannas away from the coast, and ‘coast’ for the wetter savannas and woodland on the coast. One asterisk indicates species present at Mkhuze Phinda; two asterisks indicate other species common to Maputaland and the Kenya coast. Square brackets indicate species recorded from around Shimba Hills but not in the park, although they most probably do occur there.

Toxophorinae : Gerontini Bombylella rufiventris (Macquart, 1846) : (v) East

Geron psammobates Hesse, 1938 : (i, v, x, xii) African savannas** Eastern Africa Bombylisoma gemmeum (Bezzi, 1924) : (xii) East Geron nasutus Bezzi, 1924 : (v) East & West African coasts African savannas Bombylisoma nucale (Bezzi, 1924) : (viii, xii) East

& West African savannas

[ Bombylisoma simba Bowden, 1973 : East Kenya

Toxophorinae : Toxophorini savannas] Toxophora coeruliventris Karsch, 1887 : (vi) South Eurycarenus dichopticus Bezzi, 1920 : (i, v, xii) & East African savannas African savannas Toxophora diploptera Speiser, 1910 : (v–vi) South Eurycarenus loewi Hesse, 1938 : (v) East African & East African savannas** savannas

Eurycarenus melanurus Bezzi, 1924 : (i, v–vi) East

African coasts

Toxophorinae : Systropodini

Systoechus flavicapillis Bowden, 1959 : (i, x–xi, iv)

Systropus leucoproctus Bowden, 1967 : Kenya Kenya, Tanzania coast Systoechus niveicomatus Bowden, 1959 : (xii) Bombyliinae : Bombyliini Kenya, Tanzania [ Bombomyia discoidea (Fabricius, 1794) :African savannas]* Anthracinae : Exoprosopini Bombylella auricoma (Bezzi, 1924) : (viii) North- Exoprosopa aurivestrix François, 1962 : (i, vi, xii) east & East African savannas East African savannas Bombylella delicata (Wiedemann, 1830) : (v, viii, Exoprosopa decastroi Hesse, 1950 : (v–vi) East xii) Eastern & southern Africa* African coasts Bombylella elegans (Wiedemann, 1828) : (xii) Exoprosopa luteicosta Bezzi, 1921 : (v–vi, x) South & East African savannas** Savannas of north-eastern South Africa ** Bombylella kilimanjarica (Speiser, 1910) : (v–vi, Exoprosopa punctulata Macquart, 1840 : (i, vi, viii, viii, x) East African savannas, Angola ** xii) Savannas of north-eastern South Africa * Bombylella plorans (Bezzi) : (i) West & East Africa Exoprosopa rubescens Bezzi, 1924 : (v, viii) forest Eastern Africa

Exoprosopa schmidti Karsch, 1888 : (v–vi) Kenya [ Litorhina pseudocollaris (Bezzi, 1924) : Southern & Tanzania coasts tropical Africa woodland]

Exoprosopa selenops Greathead, 2001 : (v) Kenya coast Anthracinae : Villini

Exoprosopa sp. n. near claripennis Hesse, 1956: Exhyalanthrax alliopterus (Hesse: 1956): (i, v–vi, (v, xii) Kenya coast viii, xii) Kenya & Tanzania coasts

Exoprosopa thoracica Bezzi, 1924 : (i, v–vi) Exhyalanthrax transiens (Bezzi, 1921) : (v, viii) Eastern Africa East & South Africa *

Ligyra nigripennis (Loew, 1852) : (x) Eastern Villa galla Greathead, 1967 : (vii) West & East African savannas** African savannas

[ Ligyra paris (Bezzi, 1923: West & East African savannas]** Anthracinae : Xeramoebini

Ligyra sisyphus (Fabricius, 1805) : (vi) West & East [ Petrorossia fulvipes (Loew, 1860) : East & South African savannas Africa woodland/savanna]**

[ Litorhina basalis Ricardo, 1901 ): East African Petrorossia tropicalis Bezzi, 1921 : (i) African savannas] savannas