Aetana ondawamei Huber, 2019

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96 : 78-79

Aetana ondawamei Huber sp. n.

Figs 270–271 , 286–287 , 290 , 297–299

Aetana Ind 101: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 7.

Type material. INDONESIA: ♂ holotype, ZFMK (Ar 20657), West Papua, Manokwari, Gunung Meja (0.860°S, 134.084°E), 190 m a.s.l., among rocks, 8.xi.2009 (S. Sutono) .

Other material examined. INDONESIA: 4♂ 7♀, ZFMK (Ar 20658–59), and 2♀ 1 juv. in pure ethanol, ZFMK (Ind198), same data as holotype .

Etymology. The species is named for Papuan activist John Ondawame (1953–2014) who promoted a peaceful solution to the conflict in West Papua.

Diagnosis. Males differ from most known congeners (except A. mokwam ) by pair of finger-shaped apophyses on male palpal trochanter (cf. Fig. 281 ) and by shape of procursus ( Fig. 286 ; three distal elements: whitish oval element with two heavily sclerotized rounded apophyses, and two long hinged elements); from A. mokwam by much smaller palps (compare Figs 284 and 286 ) and by wider dorsal hinged element of procursus (12 x longer than wide versus 19 x in A. mokwam ). Females differ from most congeners (except A. mokwam and A. ternate ) by elongated female internal genitalia and pore plates ( Fig. 287 ); females differ from A. mokwam and A. ternate by distinctive curved internal folds visible through cuticle ( Fig. 297 ); from A. mokwam also by anterior part of internal genitalia barely narrower than posterior part ( Figs 298–299 ). Males and females also differ from A. mokwam by pair of marks ventrally on abdomen (unpaired median mark in A. mokwam ; compare Figs 289 and 290 ).

Description. Male (holotype). MEASUREMENTS. Total length 3.5, carapace width 1.1. Distance PME-PME 340 µm; diameter PME 110 µm; distance PME-ALE 40 µm; AME absent. Leg 1: 38.5 (9.0 + 0.5 + 9.0 + 16.7 + 3.3), tibia 2: 5.5, tibia 3: 3.6, tibia 4: 5.5; tibia 1 L/d: 90.

COLOR (in ethanol). Carapace ochre yellow with narrow dark brown lateral marginal bands and narrow median band including posterior part of ocular area; clypeus with dark brown pattern in distal half; sternum monochromous light brown; legs light brown, without darker rings, tips of femora and tibiae lighter. Abdomen pale gray, densely covered with dark internal marks dorsally and laterally; ventrally with pair of dark marks behind gonopore and median mark in front of spinnerets.

BODY. Habitus as in Fig. 270 . Eye triads on very short stalks directed towards lateral. Thoracic furrow absent (only black median line). Clypeus with pair of short obtuse apophyses close to distal margin (cf. Fig. 283 ). Sternum wider than long (0.72/0.56), unmodified.

CHELICERAE. As in A. mokwam (cf. Fig. 283 ), with pair of long lateral apophyses directed towards lateral and anterior, and pair of small proximal lateral processes.

PALPS. Almost identical in shape to A. mokwam (cf. Figs 281–282 ) but much smaller (compare Figs 284 and 286 ); dorsal hinged element of procursus relatively thicker (12 x as long as thick in mid-section versus 19 x in A. mokwam ).

LEGS. Without spines; few vertical hairs; with curved hairs on tibiae and metatarsi 2; retrolateral trichobothrium of tibia 1 at 3%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, distally distinct.

Male (variation). Tibia 1 in 5 males (including holotype): 8.3–9.6 (mean 9.1).

Female. In general similar to male ( Fig. 271 ) but eye triads on low humps and closer together (distance PME- PME 200 µm), clypeus unmodified, abdomen anteriorly above pedicel with slightly sclerotized area acting against barely elevated small brown area medially on carapace; sometimes with indistinct darker rings on leg femora (subdistal) and tibiae (proximal and subdistal). Tibia 1 in 6 females: 7.9–8.6 (mean 8.3). Epigynum long, resulting in relatively posterior position of genital opening ( Fig. 290 ); anterior plate barely modified in anterior part, internal valve visible through cuticle ( Fig. 297 ), with strong transversal ridges in posterior part and dark internal folds visible through cuticle; posterior plate simple, pockets possibly present in the membrane on both sides of genital opening (arrows in Fig. 287 ). Internal genitalia with very long pore plates (actually fields of pores), apparently without internal pockets ( Figs 287 , 299 ).

Distribution. Known from type locality only ( Fig. 349 ).

Natural history. This species was found among karstic rocks near the ground, in domed sheet webs of ~ 30 cm diameter. The webs were rather exposed and the spiders moved slowly when disturbed.

Figures

FIGURE 349. Collecting sites of newly described species of Aetana in eastern Indonesia and of Giloloa sofifi sp. n. on Halmahera. Red mark: two undescribed species deposited in Bernice P. Bishop Museum, Honolulu, Hawaii.

FIGURES 264–271. Giloloa sofifi sp. n. and Aetana spp. live specimens. 264. Giloloa sofifi sp. n., male from near Sofifi. 265–266. Aetana ternate sp. n., male and female from Ternate. 267–269. Aetana mokwam sp. n., male and female from Mokwam. 270–271. Aetana ondawamei sp. n., male and female with egg sac from Gunung Meja.

FIGURES 283–287. Aetana mokwam sp. n. (283–285, ZFMK, Ar 20653–54) and Aetana ondawamei sp. n. (286–287, ZFMK, Ar 20658–59). 283. Male prosoma and chelicerae, oblique frontal view. 284, 286. Left procursi, prolateral views (at same scale). 285, 287. Cleared female genitalia, dorsal views; arrows point at possible membranous pockets. Scale lines: 0.5.

FIGURES 288–293. Aetana spp., female abdomens in ventral views, epigynum in ventral view, and cleared female genitalia in ventral and dorsal views. 288, 291–293. A. ternate sp. n. (ZFMK, Ar 20651). 289. A. mokwam sp. n. (ZFMK, Ar 20654). 290. A. ondawamei sp. n. (ZFMK, Ar 20659). Scale lines: 0.5.

FIGURES 294–299. Aetana spp., epigyna in ventral views, and cleared female genitalia in ventral and dorsal views. 294–296. A. mokwam sp. n. (ZFMK, Ar 20654). 297–299. A. ondawamei sp. n. (ZFMK, Ar 20659); arrows point at possible membranous pockets. Scale lines: 0.5.

FIGURES 281–282. Aetana mokwam sp. n. (ZFMK, Ar 20653), left male palp, prolateral and retrolateral views. em, embolus; gb, genital bulb; pr, procursus; tr, trochanter. Scale line: 1.0.