Mirnapis Urban.

Packer, Laurence & Dumesh, Sheila, 2012, Mirnapis ohloweni Packer and Dumesh, new species with notes on M. inca Urban (Hymenoptera: Apidae: Eucerini), Zootaxa 3478, pp. 113-122 : 114-116

Mirnapis Urban.

(Figs. 1–3, 4C)

Type species Mirnapis inca Urban 1997 , by original designation.

FIGURE 1. Important characters for generic level identification of Mirnapis . A–E, labra; A. male M. ohloweni (this is the longest and least strongly apically concave labrum of any male of the genus available to us); B and C, male M. inca , (the latter has the most obtuse lateral lobes of any individual of the species available to us); D. female M. ohloweni ; E. female M. inca ; F. part of forewing of M. ohloweni showing how length (blue) and width (red) of 2nd submarginal cell were measured; G. midtibia of male M. inca to show the lanceolate hairs; H. pygidial plate of female M. inca ; I. pygidial plate of male M. inca ; J. S2 of female M. inca to show the angle of the sternal gradulus.

Diagnosis: For the male each of i) labrum strongly excised apically with the sides of the emargination at approximately a right angle to each other and ii) S7 with inner lobe weakly sclerotized, transversely scrolled and sparsely setose, are unique among Neotropical Eucerini . For the female, the labrum with deep apicomedian concavity extending for approximately 0.4X the maximum length of the labrum is diagnostic. This character has to be observed for the ventral (oral) surface of the labrum as the concavity is hidden by a dense tuft of hairs on the dorsal (aboral) surface.

Description: Labrum long, length more than 0.6X width in male (Figs. 1A–C), sometimes slightly less in female (Figs. 1D and E); apex with concavity deep and narrow in female, extending for ~0.4X greatest length; shallower and triangular, <0.2X greatest length, in male; lobes on either side approximately right angular, sides of emargination approximately right angular in male, acutely angulate in female. Galea 1.05–1.1X as long as compound eye. Mandible without inner basal tooth, subapical tooth obtuse or absent, apex triangularly excised in fresh specimens, condylar ridge of normal saliency. Maxillary palpomeres 5 (rarely 4); mouthparts lacking hooked hairs. Clypeal margin unmodified; clypeocular distance 0.75X minimum diameter of F1; paraocular carina absent below thus not united with lateral clypeal carina; clypeus produced for slightly more than half width of compound eye (20:42). Malar space short, ~0.3X basal depth of mandible. Vertex elevated such that ocelli face somewhat anteriorly; distinct acute tubercle lateral to lateral ocellus. Inner margin of compound eyes subparallel. Female scape length ~ 3X maximum width; male F1 minimum length at most 1/6 as long as F2, maximum length 1/4–1/3 as long as F2; male antenna attaining T4, flagellum slightly flattened, depth 0.8X width. Scutellum approximately one third length of mesoscutum; anterior margin of tegula convex. Submarginal cells 3; submarginal cell 2 shorter than wide (Fig. 1F) or with length and width subequal; stigma much shorter on posterior margin than prestigma (Fig. 1F); jugal lobe more than half as long as vannal lobe; hindwing cu less than half as long as 2nd abscissa of M+Cu; wing hairs sparse. Forecoxal spine absent; male midtibia with lanceolate hairs apicoventrally (Fig. 1G), legs otherwise unmodified; keirotrichiate area on inner surface of hind tibia of female broad, extending almost to base; tibial and basitarsal scopal hairs long, plumose ( Fig. 2A, 2B and 4 C), apical bare area short, ~0.15X length of tibia; basitibial plate largely obscured by hairs, only posterior margin visible; dense dorsobasal hair patch on hind tibia shorter than basitibial plate; arolia present. Apical impressed areas of metasomal terga extensive,>0.5 tergal length on T2, these areas translucent only towards apex. Spatuloplumose hairs absent; metasomal terga lacking distinct basal hair bands (the female T 2 may have a weak pale band basolaterally); patch of appressed, plumose, white hairs sublaterally on T4 and laterally on T5 of female, sublaterally on both T4 and T5 of male ( Figs. 2A , C, E and G). Pygidial plate acute <45° in female (Fig. 1H), narrowly subtruncate in male with apical margin <MOD (Fig. 1I); gradulus of female on T2–T6 carinate laterad, untoothed; male gradulus on T2–T7 carinate laterally, T4–T5 with apex of gradular carina right angular, sometimes somewhat obtuse or acute, T6 with acute lateral tooth, T7 without tooth; female S2 gradulus concave medially, angle subtended by sides of concavity ~140° (Fig. 1J). Female with sternal hairs suberect, male with subapical fringe of long hairs on S2–S5. Male S6 with converging carinae, with shallow median depression margined with posteriorly oriented setae; S7 disc flat and apically weakly bilobed, inner lobes somewhat membranous, scrolled, sparsely setose; outer lobes strongly sclerotized, ventrally concave ( Fig. 3 A and C); S8 with apex of apical lobe broadly concave, apicodorsal swelling broad and deep ( Figs. 3 A and B); gonostylus 2/3 as long as maximum length of gonocoxite, spatha ~ 2X as wide as long ( Figs. 3 C and D).

Figures

FIGURE 3. Terminalia of male Mirnapis, ventral views. A and C, M. ohloweni (S 7 – S 8 and genital capsule respectively); B and D, M. inca (S 7 – S 8 and genital capsule respectively).

FIGURE 4. Characters to aid in identification of Mirnapis based upon keys in the text. A. Labrum of male Svastrides melanura to show strongly obtuse emargination and apicolateral margin. B. tibial scopa of M. inca to show plumose hairs. C. tibial scopa of S. melanura to show mostly simple hairs.

FIGURE 2. A, B, E, F Mirnapis ohloweni Packer & Dumesh, new species. C, D, G, H. M. inca. A and C, female habitus lateral view; B and D, female T 1 – T 3, dorsal view; E and G, male habitus, lateral view; F and H, male T 1 – T 3, dorsal view.