Pimpinella saxifraga L. subsp. rupestris Weide, 1962 (Weide, Fedd. Repert. 64: 259. 1962; Sourek , Preslia 39: 70)

Kwiatkowski, Pawel, Sida, Otakar & Urbaniak, Jacek, 2022, Pimpinella saxifraga subsp. rupestris (Apiaceae) - taxonomy and nomenclature of stenoendemic taxon from Karkonosze Mountains (Sudetes, Poland), PhytoKeys 213, pp. 111-118 : 111

Pimpinella saxifraga L. subsp. rupestris Weide, 1962 (Weide, Fedd. Repert. 64: 259. 1962; Sourek, Preslia 39: 70)

Holotype.

-Poland. Europe, Sudetes: Kleine Schneegrube des Riesengebirges (Hirte, Naturwissenschaftlichen Museum Coburg) [missing, Heimo Rainer, pers. comm.]. - Lectotype (designated here): Poland. Flora des Westsudeten. Basalt in der Kleinen Schneegrube [Karkonosze Mountains-Mały Śnieżny Kocioł Cirque, basaltic rocks, ca 1300 m a.s.l.], 14 August 1891, leg. G. Hirte, (PR 162605! - Fig. 1 ; isolectotypes: G00379179, G00379180, G00848072, JE00028396, JE00028397, JE00028398, PR162596).

All specimens (isolectotypes) come from the same collection by G. Hirte (August 14, 1891) and were sent as duplicates to various herbaria (Geneva, Jena, Pruhonice).

Description.

Plants (5-)10-35 cm tall. Rosette leaves with ± long petioles, 2-pinnate with 3-6(-8) pairs of sessile leaflets; leaflets rounded to ovate, evenly dentate or serrate. Stems cylindrical, sometimes slightly striate, rarely branched in the upper part, leafless or with 1-3 cauline leaves. Lower cauline leaves 1-pinnate, dentate with obtuse teeth, light green; middle cauline leaves 2-pinnate, sessile with short sheaths; leaflets of upper cauline leaves reduced, linear or lanceolate. Umbels small, with 7-14 rays of uneven length; rays smooth or ± ciliate; involucres and involucels usually absent. Petals whitish, yellowish, sporadically pink, up to 0.7-1.0 mm long, cordate, incurved at tips. Fruits 1.0-2.0(-2.5) × 0.5-1.5(-2.0) mm, ovoid, slightly compressed, smooth with ribs distinct only at maturity (Fig. 2 ). The most important differences in the morphological structure between the nominative taxa of Pimpinella saxifraga subsp. saxifraga and the discussed P. saxifraga subsp. rupestris are given in Table 1 .

Poland. Sudetes: Riesengebirge [Karkonosze Mountains]: Kleine Schneegrube [ Mały Śnieżny Kocioł Glacial Cirque], 14 August 1891, Hirte (Flora silesica exsiccata No 375, 1891), ut Pimpinella saxifraga L. var. alpestris Spreng. (G00379179, G00848072, JE00028397); Am Basalt in der Kleinen Schneegrube [basalt outcrop in the Mały Śnieżny Kocioł Glacial Cirque], 14 August 1891, Hirte, ut Pimpinella saxifraga L. var. alpestris Spreng. (G00379180, JE00028396, JE00028398, PR162605); M. Sněžná jáma [ Mały Śnieżny Kocioł Glacial Cirque], August 1921, Pilát, ut Pimpinella saxifraga var. petraea (PR162596).

Figures

Figure 1. Lectotype of Pimpinella saxifraga L. subsp. rupestris Weide, 1962: National Museum in Prague, Czech Republic, PR 162205.

Figure 2. Living plants of Pimpinella saxifraga L. subsp. rupestris Weide 1962 A general view of type locality - locus classicus, Maly Sniezny Kociol glacial cirque, Karkonosze Mountains, Sudetes, Poland, Europe B stem C pinnate leaf D leaf rosette E inflorescence F infrutescens (A photo by P. Kwiatkowski; B-F photo by L. Przewoznik).