Bolla subgisela Strand, 1921

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Genomic analysis reveals new species and subspecies of butterflies, The Taxonomic Report of the International Lepidoptera Survey 11 (6), pp. 1-63 : 34-35

Bolla subgisela Strand, 1921 is a junior subjective synonym of Staphylus melangon epicaste Mabille, 1903 and not of Bolla eusebius (Plötz, 1884)

Genomic analysis of the holotype of Bolla subgisela Strand, 1921 (type locality in Colombia), currently regarded as a junior subjective synonym of Bolla eusebius (Plötz, 1884) (type locality in Central America) reveals that it is not monophyletic with Bolla Mabille, 1903 (type species Bolla pullata Mabille, 1903 treated as a junior subjective synonym of Staphylus imbras Godman and Salvin, 1896 ), but instead is placed within specimens of Staphylus melangon (Mabille, 1883) (type locality in South America) and away from Staphylus tucumanus (Plötz, 1884) (type locality in Argentina), a sister species of S. melangon ( Fig. 33). Therefore, B. subgisela is conspecific with S. melangon . The phylogenetic analysis we used does not differentiate between subspecies of S. melangon ( Fig. 33), and we assign B. subgisela to subspecies by a combination of wing pattern characters and locality. Only Staphylus melangon epicaste Mabille, 1903 (type locality in Brazil) possesses brown ventral hindwing without dominant white overscaling toward tornus and inner margin, similar to B. subgisela , and it is the only subspecies documented from Colombia ( Evans 1953). Therefore, we propose that Bolla subgisela Strand, 1921 is a junior subjective synonym of Staphylus melangon epicaste Mabille, 1903 .

from Gorgythion begga (Prittwitz, 1868)

Gorgythion marginata Schaus, 1902 ( type locality in Peru) ( Fig. 34 red) currently regarded as a junior subjective synonym of Gorgythion begga pyralina (Möschler, 1877) ( type locality in Suriname) ( Fig. 34 blue, part) is genetically differentiated from it and generally from Gorgythion begga (Prittwitz, 1868) ( type locality in Brazil: Rio de Janeiro) at the species level ( Fig. 34), e.g., Fst /COI barcode difference of 0.26/2.9% (19 bp). Therefore, we propose that Gorgythion marginata Schaus, 1902 , stat. rest. is a species-level taxon distinct from Gorgythion begga (Prittwitz, 1868) .