Bagous bipunctatus

ROBERTO CALDARA, CHARLES W. O’BRIEN & MASSIMO MEREGALLI, 2017, A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults, Zootaxa 4287 (1), pp. 1-63 : 36-37

5. Bagous bipunctatus group

Diagnosis. Tube of penis body with a pair of lateral concentrations of more or less numerous, fine to large, mostly acute denticles (char. 87.1). Ventrobasal margin of penis body with short, ventrobasal plate extending between apodemes, therefore visible from above (excluding B. kagiashi and B. picturatus from Japan) (char. 70.1).

Remarks. The species presently included in this large widespread group are defined by one distinctive character of the endophallus, which consists of a pair of regions with distinctly concentrated and usually minute (rarely large) denticles toward the apex. Otherwise the group is heterogeneous in external structure such as tarsal shape and tibial curvature and dentition. Among the species of the complex, the females of B. bipunctatus and B. puncticollis possess a bursa with sclerites more or less bilobed and convex (as in unrelated species like those of the B. argillaceus group) (char. 116.2). Monophyly of this group is suggested also by the phylogenetic analysis, its species being included in a clade, although not statistically supported (44% pp). It is distributed worldwide except for Australia.

Species included. AFR: 4 Bagous coenosus Gyllenhal, 1836 , *³ B. fastosus Hartmann, 1904 , 4 B. madecassus Fairmaire, 1897 , 4 B. quadrinodulosus Hustache, 1932 . NAR: ² B. californicus LeConte, 1876 , ² B. chandleri Tanner, 1943 , ² B. dietzi Tanner, 1954 , ² B. floridanus Tanner, 1943 , 5 B. lunatoides O'Brien, 1979 , ² B. lunatus Blatchley, 1916 , *10 B. mamillatus Say, 1831 , ² B. obliquus LeConte, 1876 , ² B. planatus LeConte, 1876 , ² B. puritanus Blatchley, 1916 , ² B. restrictus LeConte, 1876 , 7 B. tuberosus ( Tanner, 1943) . ORR: *7 B. difficilis O'Brien, 1995 , *8 B. vicinus Hustache, 1926 . PAL: ² B. amurensis Egorov & Gratshev, 1990 , §*4 B. bagdatensis Pic, 1904 , *¹ B. bipunctatus ( Kôno, 1934) , §² B. glabrirostris (Herbst, 1795) , *9 B. kagiashi Chujo & Morimoto, 1959 , §² B. lutulentus (Gyllenhal, 1813) , *6 B. occultus O'Brien, 1995 , ² B. olcesei Tournier, 1874 , *5 B. picturatus Egorov & Gratshev, 1990 , ¹ B. puncticollis Boheman, 1845 , §*² B. robustus Brisout de Barneville, 1863 , ² B. spiculatus O'Brien & Morimoto, 1994 , §² B. subcarinatus Gyllenhal, 1836 , ² B. subcordatus O'Brien & Morimoto, 1994 , *11 B. yamazakii O'Brien & Morimoto, 1995 .