Bagous tubulus

ROBERTO CALDARA, CHARLES W. O’BRIEN & MASSIMO MEREGALLI, 2017, A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults, Zootaxa 4287 (1), pp. 1-63 : 45

15. Bagous tubulus group

Diagnosis. Body elongate-cylindrical. Pronotum 0.90x or more as long as broad (char. 26.1). Rostrum cylindrical, narrow, elongate (especially in female). Elytra nearly as wide as pronotum, without calli (char. 36.0). Sclerites complex within tube of penis body with a pair of submedian subapical sclerites posteriorly joined (char. 84.2), with chela-like basal portion. Median setal brush behind orifice situated at base of subapical sclerites (char. 93.1). Penis body elongate, with sinuate and subparallel sides. Tegmen lacking parameroid lobes (char. 111.2). Arms of spiculum ventrale longer than apodemes, with long apical setae (char. 113.1). Gonocoxites elongate.

Remarks. This group is characterized by the rostrum being cylindrical, narrow, and elongate (especially in the female), and a series of non-genital characters, e.g., the elongate body, and genital character states in part shared also by the species of the B. cylindricus group. Previously, B. elegans and B. majzlani (formerly Dicranthus ) were separated in a distinct group by Caldara & O’Brien (1998) due to their very distinctive habitus, including the pattern of the dorsal vestiture (char. 39.1 and 39.2), and the shape of the elytra in which the confluence of intervals 3 and 9 is markedly raised and forms a long posteriorly-projecting process (char. 38.1). In both all-species ( Fig. 9 ) and type-species ( Fig. 10 ) analyses this group was strongly supported; the complex comprising B. tubulus / B. friwaldszkyi was sister to the complex B. elegans / B. majzlani . This group is composed of seven Palaearctic species.

Species included. PAL: §*³ Bagous elegans (Fabricius, 1801) ; *² Bagous friwaldszkyi Tournier, 1874 , ¹ B. henoni Hustache, 1927 , *4 B. majzlani ( Kodada, Holecova & Behne, 1992) ; ¹ B. mingrelicus Tournier, 1874 , ¹ B. minutus Hochhuth, 1847 , *¹ B. tubulus Caldara & O’Brien, 1994 .

Figures

FIGURE 9. Morphological analysis—all species. Bayesian consensus tree (50 % majority rule). Numbers above branches: support values, in percentage, in the following order: posterior probability (BI) / bootstrap value (ML) / symmetric resampling (MP); ‘ ns’ means that the group was present, but its support was <50 %; ‘ np’ means that the group was not present. Number below branches: Bremer support in the MP analysis. Scale bar unit: expected changes of state per character. MP analysis: length 261, consistency index = 0.61, retention index = 0.83. Outgroup taxa not reported to increase readability of the Bagoini clade.

FIGURE 10. Morphological analysis—type species only. Bayesian consensus tree (50 % majority rule). Taxa in bold font: valid taxa recognized in the present work; larger font: genera, smaller font: subgenera. Taxa in regular font: synonyms in the present work. Numbers above branches: support values, in percentage, in the following order: posterior probability (BI) / bootstrap value (ML) / symmetric resampling (MP); ‘ ns’ means that the group was present, but its support was <50 %; ‘ np’ means that the group was not present. Number below branches: Bremer support in the MP analysis. Scale bar unit: expected changes of state per character. MP analysis: length 146, consistency index = 0.70, retention index = 0.78.