Mirokempynus, Ma & Shih & Ren & Wang, 2020

Ma, Yiming, Shih, Chungkun, Ren, Dong & Wang, Yongjie, 2020, New lance lacewings (Osmylidae: Kempyninae) from the Middle Jurassic of Inner Mongolia, China, Zootaxa 4822 (1), pp. 94-100 : 95

Genus Mirokempynus gen. nov.

( Fig. 1 )

Zoobank LSID: urn:lsid:zoobank.org:act:D4912F2E-63D3-4740-8931-7FAA181A21E5

Type species. Mirokempynus profundobifurcus sp. nov. by monotypy.

Diagnosis. Hind wing: RP with a large number of branches, tightly arranged; radial crossveins dense among the proximal one-third RP branches, and sparse distally; outer gradate series well-developed; MP forked two times in the basal and distal one-third part, and intramedial area with one–three rows of regular cells; CuA long, about twice as long as CuP, with strongly pectinated branches; CuP branches pectinated at the middle.

Etymology. From the Latin prefix mir-, peculiar and kempynus , the type genus of Kempyninae , referring to the unique branches of MP in the hind wing. Gender is masculine.

Remarks. Although the holotype of Mirokempynus profundobifurcus sp. nov. does not preserve forewings, this taxon can be unequivocally assigned to Kempyninae showing the apomorphies of this subfamily, i.e. presence of multiple rows of cells in intramedial area in the hind wing ( Winterton et al. 2019; Ma et al. 2020). The new genus is characteristic due to its particular MP branchings and intramedial cells of hind wing. In the new genus, MP is forked near the base of the wing, instead of only forming distal branches, and MP2 bifurcated at the middle, so multiple rows of cells are present due to several longitudinal veins arranged in intramedial area. Among most fossil kempynines, e.g. Arbusella Khramov, 2014 and Jurakempynus Wang, Liu, Ren & Shih, 2011 , MP of hind wing is commonly branched distally, and the intramedial cells were irregularly arranged and formed by the intramedial crossveins. A similar MP branching pattern in the hind wing is also present in Ponomarenkius Khramov, Liu & Zhang, 2017. However, it is distinctly different from Mirokempynus gen. nov. due to the proximally branched MA, relatively broad hind wing, and presence of stripe-like markings. In addition, CuA of Mirokempynus has more than 20 branches that occupy approximately the half of its length, which is distinctly different from the other genera ( Wang et al. 2011; Khramov 2014b; Khramov et al. 2017).

Figures

FIGURE 1. Lance lacewing Mirokempynus profundobifurcus gen. et sp. nov., Holotype from the Middle Jurassic of Jiulongshan Formation in Daohugou, China. A. Partial body and hind wing in Part (CNU-NEU-NN2020001P). B. Counterpart (CNU- NEU-NN2020001C). Photo 1B is flipped left to right for easy comparison with A and C. C. Line drawing of the hind wing of Mirokempynus profundobifurcus gen. et sp. nov. A–B. Scale bar is 10 mm. C. Scale bar is 5 mm.