Achilia larvata (Reitter, 1885)

Giorgio Sabella, Sergey A. Kurbatov & Giulio Cuccodoro, 2017, A revision of the Chilean Brachyglutini - Part 2. Revision of Achilia Reitter, 1890: A. crassicornis, A. tumidifrons, A. bifossifrons, and A. lobifera species groups (Coleoptera: Staphylinidae: Pselaphinae), Revue suisse de Zoologie 124 (1), pp. 119-140 : 136-137

Achilia larvata (Reitter, 1885)

Additional material (1284 ex.): FMNH (Orlando Park Pselaphidae collection, ex F. C. Fletcher collection); 2 ♀; Chile. – CENTRAL CHILI: Región Los Lagos: Chiloé prov.: MHNS; 2 ♀ (mislabelled paratypes of Achilia tumidifrons n° 1640 and 1642); Chepu; 03.X.1958; G. Kuschel.– MHNS; 1 ♂ (mislabelled paratypes of Achilia tumidifrons n° 1644); Chepu; 04.X.1958; G. Kuschel. – MHNS; 3 ♀ (mislabelled paratypes of Achilia tumidifrons n° 1633, 1635 and 1639); Chepu; 15.X.1958; G. Kuschel. – MHNG; 7 ♂ and 21 ♀; Chiloé Island, Huillinco Lake; 31.I.1983; T. Cekalovic. – DBUC; 1 ♂ and 6 ♀, same data; T. Cekalovic. – MSNG; 1 ♂; same locality; TC-278; 21.II.1991, T. Cekalovic. – MSNG; 1 ♀ and 18 ♀; Chiloé Island, 1 km W of Huillinco Lake; TC-564; 24.I.1998; T. Cekalovic. – MHNG; 1 ♀; Chiloé Island, Río Pudeto; 28.II.1972; T. Cekalovic. – MHNG; 2 ♀; same locality; 27.I.1983; T. Cekalovic. – DBUC; 1 ♂ and 1 ♀; same data; T.Cekalovic. – MSNG, 1 ♂ and 1 ♀; same locality; SyTC-226; 21.II.1989; S. & T. Cekalovic. – UNHC; 1 ♂; Chiloé Island, 2 km N Puente Río Pudeto; 20.I.2000; T. Cekalovic. – MHNG; 1 ♀; San Pedro; 22.II.1976; T. Cekalovic. – FMNH (FMHD #97-21); 8 ♂ and 9 ♀; Puente La Caldera, 9.8 km E of Cucao; 42° 39.96’S 74° 00.70’W; 10 m; 14.I.1997; A. Newton & M. Thayer 991; valdivian raiforest, berlese, leaf & log litter. – MSNG; 20 ♂ and 17 ♀; Chiloé Island, Puente La Caldera; TC-466; 15.II.1996, T. Cekalovic. – MSNG; 2 ♂; same locality; TC-524; 18. II.1997; T. Cekalovic. – MSNG; 1 ♂; Chiloé Island, Chepu; TC-275; 19.II.1991; T. Cekalovic. – MSNG; 2 ♂ and 2 ♀; same locality; TC- 580; 09.II.1999; T. Cekalovic. – MSNG; 14 ♀; same locality; TC-610; 20.I.2000; T. Cekalovic. – MSNG; 12 ♂ and 2 ♀; same locality; TC- 624; 26.I.2000, T. Cekalovic. – MSNG; 6 ♂ and 3 ♀; same locality; TC-625; 26.I.2000, T. Cekalovic. – MSNG; 1 ♂ and 1 ♀; Chiloé Island, Estero Tablin; TC-609; 19.I.2000, T. Cekalovic. – MNSG; 1 ♀; Chiloé Island, 1 km N of Puente Notuco; TC-528; 20.II.1997; T. Cekalovic. – MSNG; 4 ♀; Chiloé Island, Puente Milildeo; TC-471; 15.II.1995; T. Cekalovic. – MSNG; 1 ♀; Chiloé Island, San Juan de Chadmo; TC- 555; 18.I.1998, T. Cekalovic. – MSNG; 2 ♂; Chiloé Island, 5 km SW Chonchi; 14.I.1999, T. Cekalovic.– MSNG; 4 ♂ and 4 ♀; same locality; TC-560; 21.I.1998; T. Cekalovic. – MSNG; 11 ♂ and 19 ♀; Chiloé Island, Estero Llicaldad; TC-608; 19.I.2000; T. Cekalovic. – MSNG; 1 ♀; Quinchao Island, Laguna Pulul; TC-615; 22.I.2000; T. Cekalovic. – Llanquihue prov.: MHNG; 1 ♂ and 5 ♀; La Arena, 45 km SE Puerto Montt; 100 m; 25.XII.1984; S. & J. Peck. – MHNG; 3 ♀; Petrohué; 30.I.1979;A. De Chambrier. – FMNH (FMHD #85-995, #85-112); 2 ♂; Vicente Perez National Park, Salto Petrohué; 150 m; 04.II.1985; S. & J. Peck; mixed forest litter, berlese. – FMNH (FMHD #85-947, #85-63); 4 ♀; Lenca, 45 km SE Puerto Montt; 100 m; 25.XII.1984; S. & J. Peck; forest remnant, leaf stick litter, berlese. – MHNS; 1 ♂ and 1 ♀ (mislabelled paratypes of Achilia globiceps n° 1647 and 1657); Frutillar; 20.IX.1954; G. Kuschel. – MHNG; 1 ♂ and 1 ♀; Frutillar Bajo, Universidad Chile Forest Reserve; 100 m; 22.XII.1984/02. II.1985; S. & J. Peck; FIT ravine mixed forest. – DBUC; 1 ♂ and 1 ♀, same data; S. & J. Peck. – FMNH (FMHD #2002-085); 3 ♀; Ensenada, Las Cascadas road, 0.9 km N of La Burbuja; 41° 11.27’S 72° 32.6’W; 80 m; 16.XII.2002; M. Chani; Nothofagus dombeyi , mixed hardwoods, berlese, litter. – FMNH (FMHD #97-38); 3 ♂ and 7 ♀; Vicente Perez Rosales National Park, SW slope Vn Osorno, km 4 to La Burbuja; 41° 09.95’S 72° 30.80’W; 310 m; 27.I.1997;A. Newton & M. Thayer 1007; secondary valdivian rainforest w/ Nothofagus dombeyi- Eucryphia cordifolia berlese, leaf & log. – Osorno prov.: MHNG; 3 ♂; environs of Osorno; H. Franz. – FMNH (FMHD #96-248); 4 ♂ and 8 ♀; 15.1 km W Puaucho; 40° 34.97’S 73° 37.68’W; 50 m; 30.XII.1996; A. Newton & M. Thayer 984; valdivian rainforest remnant in sm. ravine, w/large ferns, berlese, leaf & log litter. – FMNH; 4 ♂ and 12 ♀; Puyehue National Park, Antillanca road; 470 m; 20-25.XII.1982; A. Newton & M. Thayer; valdivian rainforest, vouchers associated with larvae, berlese, leaf & log litter, forest floor. – FMNH (FMHD #2002-082); 12 ♂ and 23 ♀; Vicente Perez Rosales National Park, SW slope Volcàn Osorno, road to Ref. La Picada; 41° 03.25’S 72° 30.18’W; 660 m; 16.XII.2002; A. Solodovnikov, A. Newton & M. Thayer 1067; Nothofagus dombeyi w/conifers dense Chusquea bamboo understory, flat area, berlese, leaf & log litter. – MHNG; 17 ♂ and 53 ♀; Puyehue National Park,Anticura Repucura trail; 500 m; 06.II.1985; S. & J. Peck; forest litter. – DBUC; 1 ♀; same data; S. & J. Peck; forest litter. – FMNH (FMHD #85-996, #85-113); 8 ♀; same data; S. & J. Peck. – DBUC; 1 ♂ and 1 ♀; same data; S. & J. Peck. – FMNH; 1 ♂ and 2 ♀; Puyehue National Park, 4.1 km E Anticura, trap site 662; 430 m; 19-26. XII.1982; A. Newton & M. Thayer; valdivian rainforest. – FMNH 1 ♂ and 1 ♀; same data; A. Newton & M. Thayer; valdivian rainforest, window traps. – FMNH; 8 ♂ and 48 ♀; same data; A. Newton & M. Thayer; valdivian rainforest, vouchers associated with larvae, berlese, leaf & log litter, forest floor. – UNHC; 1 ♂; 4.1 km W Anticura, site 663; 270 m; 19-25.XII.1982; A. Newton & M. Thayer; valdivian rainforest, flight intercept (windows) trap. – FMNH; 1 ♂; same data; A. Newton & M. Thayer. – FMNH; 4 ♂ and 4 ♀; same data; A. Newton & M. Thayer; valdivian rainforest, vouchers associated with larvae – FMNH (FMHD #97-40); 17 ♂ and 58 ♀; Puyehue National Park, 4 km E Anticura; 40° 39.73’S 72° 08.10’W; 460 m; 30.I.1997; A. Newton & M. Thayer 985-2; valdivian rainforest w/large, Saxegothea , berlese, leaf and log litter. – FMNH (FMHD #97-4); 6 ♂; same data; 01- 30.I.1997; A. Newton & M. Thayer 985-2; valdivian rainforest w/large, Saxegothea , flight intercept trap. – FMNH (FMHD #97-39); 25 ♂ and 61 ♀; same data; A. Newton & M. Thayer 985-3; valdivian rainforest w/large, Saxegothea berlese, leaf & log litter. – FMNH (FMHD #97- 41); 5 ♂ and 9 ♀; same data; 30.I.1997; A. Newton & M. Thayer 985-1; valdivian rainforest w/large, Saxegothea , berlese, leaf & log litter. – FMNH (FMHD #96-250); 7 ♂ and 1 ♀; same data; A. Newton & M. Thayer 985-1; valdivian rainforest w/large, Saxegothea berlese, flight intercept trap. – FMNH (FMHD #2002-90); 8 ♂ and 14 ♀; Puyehue National Park, Ruta 215; km 4.5 of Aduana station; 40° 40.23’S 72° 05.21’W; 580 m; 19.XII.2002; A. Newton, M. Thayer, D. J. Clarke & M. Chani 1071; valdivian rainforest, berlese, leaf & log litter. – MHNG; 14 ♂ and 76 ♀; Puyehue National Park,Aguas Calientes, Pionero track; 500 m; 28.XII.1984; S. & J. Peck; sifted forest stick litter. – DBUC; 3 ♂; same data; S. & J. Peck. – MHNG; 6 ♂ and 29 ♀; Puyehue National Park, Aguas Calientes, station 25a; 400-500 m; 31.XII.1990/01.I.1991; M. Agosti & D. Burckhardt. – FMNH (FMHD #85-928, #85-43); 4 ♀; Puyehue National Park, Aguas Calientes; 500 m; 20.XII.1984; S. & J. Peck; forest litter on trail, sifting. – PHPC; 5 ♂; Puyehue National Park, 26.2 km E Entre Lagos, near Termas Aguas Calientes; 40° 44.130’S 72° 18.427’W; 460 m; 09-12.III.2008; H. Wood & C. Griswold; sifting litter. – MHNG; 2 ♂ and 3 ♀; Pucatrihue, 65 km W Osorno, station 21; 40° 28’S 73° 43’W; 150 m; 04.XII.1984; D. Burckhardt; valdivian rainforest, sifting of moss on dead tree trunks, branches and rocks and of vegetable detritus. – FMNH (FMHD #85-933, #85-48); 4 ♂ and 11 ♀; 3 km S Maicolpué, Bahìa Mansa; 200 m; 21.XII.1984; S. & J. Peck; mixed forest litter, berlese. – MHNG; 20 ♂ and 73 ♀; 3 km S Maicolpué, Bahìa Mansa; 21.XII.1984; S. & J. Peck; mixed forest litter. – MHNG; 9 ♂ and 46 ♀; same locality; 03.II.1985; S. & J. Peck. – FMNH; 1 ♂ and 8 ♀; same data, S. & J. Peck. – DBUC; 2 ♂ and 2 ♀; same data; S. & J. Peck. – FMNH (FMHD #96-247); 1 ♂ and 9 ♀; Hills S of Maicolpué; 40° 36.57’S 73° 44.91’W; 160 m; 30.XII.1996; A. Newton & M. Thayer 983; disturbed valdivian rainforest, berlese, leaf & log litter. – UNHC; 2 ♂ and 3 ♀; Hills S of Maicolpué; 160 m; 21.XII.1982; A. Newton & M. Thayer; 2° valdivian forest, berlese, leaf & log litter, forest floor. – FMNH; 3 ♂ and 40 ♀; same data; A. Newton & M. Thayer. – MSNG; 1 ♂ and 2 ♀; Los Ñilques; TC-257; 10.I.1990; T. Cekalovic. – MSNG; 5 ♂ and 11 ♀; same locality; TC-260; 13.I.1990; T. Cekalovic. – MSNG; 1 ♂ and 2 ♀; same locality; TC-553; 17.I.1998, T. Cekalovic. – Región Los Ríos: Ranco prov.: MHNG; 5 ♂ and 21 ♀; 34 km WNW La Unión, station 36; 700 m; 17.XII.1984; S. & J. Peck; litter mixed evergreen forest. – DBUC; 1 ♂ and 2 ♀; same data; S. & J. Peck. – FMNH (FMHD #85-997, #85-114); 1 ♀; 35 km WNW La Unión; 700 m; 07.II.1985; S. & J. Peck; litter mixed evergreen forest, berlese. – Valdivia prov.: UNHC; 2 ♂; Oncol Park; 485 m; 20.I.2001; T. Cekalovic. – PHPC; 8 ♂ and 16 ♀; Oncol Park, 12 km NW Valdivia, Sendero Bonifacio, WDS-T-201; 39° 42’S 73° 19’W; 22.II.2008; W. D. Shepard; sifting litter. – DBUC; 1 ♂ and 1 ♀; same data; W. D. Shepard. – FMNH (FMHD #97-19); 1 ♀; Rincón de La Piedra, turnoff, 14.8 km SE Valdivia; 39° 55’ 32’’S 73° 06’ 27’’W; 50 m; 11.I-01.II.1997; A. Newton & M. Thayer 990; disturbed Valdivian rainforest, with Nothophagus dombeyi and Podocarpus saligna , carrion trap (squid). – FMNH (FMHD #97-20); 3 ♂ and 10 ♀; same locality; 11.I.1997; A. Newton & M. Thayer 990; berlese, leaf & log litter. – FMNH (FMHD #97-42); 1 ♂; same locality; 02.II.1997; A. Newton & M. Thayer 990; disturbed Valdivian rainforest, with Nothophagus dombeyi and Podocarpus saligna , berlese, litter in ground bromeliad (? Puya sp.). leaf axils (live & dead). – Región Araucanía: Cautín prov.: MHNG; 2 ♂; 15 km NE Villarica, Flor del Lago; 500 m; 10.II.1985; S. & J. Peck; forest litter, berlese. – FMNH (FMHD #85-999, #85-116); 1 ♂; same data; S. & J. Peck. – MHNG; 2 ♂ and 3 ♀; 15 km NE Villarica, Flor del Lago, station 16; 300 m; 14.XII.1984; S. & J. Peck; sifted Boletus in spruce plantation. – UNHC; 1 ♂ and 1 ♀; Bellavista, North shore Lago Villarica, site 655; 310 m; 15-30.XII.1982; A. Newton & M. Thayer; valdivian rainforest, flood debris forest stream. – FMNH; 2 ♂ and 1 ♀; same data; A. Newton & M. Thayer. – FMNH; 4 ♀; same data; A. Newton & M. Thayer; valdivian rainforest, flood vouchers associated with larvae. – FMNH; 6 ♀; Ñielol National Park, near Temuco, site 652; about 250 m; 14/30.XII.1982; A. Newton & M. Thayer; native forest remnants with Nothophagus, vouchers associated with larvae, leaf & log litter, forest floor. – Malleco prov.: MHNG; 4 ♂ and 37 ♀; Purén, Contulmo Natural Monument; 350 m; 13.II.1985; S. & J. Peck; mixed forest litter. – DBUC; 1 ♂ and 2 ♀; same data; S. & J. Peck. – UNHC; 1 ♂; Contulmo National Park, 10 km W Purén; 240 m; 12.

XII.1982;A. Newton & M. Thayer; mixed hdwd. forest with Chusquea , berlese, vouchers associated with larvae, leaf & log litter, forest floor. – FMNH; 5 ♂ and 14 ♀; same data; A. Newton & M. Thayer. – FMNH (FMHD #2002-64); 1 ♂ and 5 ♀; Contulmo Natural Monument, Sendero Lemu Mau; 38° 00.74’S 73° 11.13’’W; 410 m; 08.XII.2002; A. Newton & A. Solodovnikov 1059; Nothophagus obliqua-Eucryphia cordifolia w/fern & bamboo understory, flood debris, small stream. – FMNH; 3 ♀; same data; D. J. Clarke & A. Solodovnikov 1059; Nothophagus obliqua-Eucryphia cordifolia w/fern & bamboo understory, sifted litter, hand-collected. – Región Bío Bío: Concepción prov.: MHNG; 12 ♂ and 3 ♀; Pinares; 18.III.1973; T. Cekalovic. – MSNG; 1 ♂ and 2 ♀; El Manzano; TC-329; 08.XI.1992; T. Cekalovic. – Bío Bío prov.: MHNS; 1 ♂ (mislabelled paratypes of A. globiceps n° 1659); Pemehue; I.1896; P. Germain. – MNHN; 1 ♀ (sub A. globocephala ?); Cordillera of Pemehue; 38° 00’S; P. Germain. – Ñuble prov.: MHNS; 1 ♀ (mislabelled paratypes of A. globiceps n° 1658); Chillán; P. Germain. – MNHN; 1 ♂ (sub A. globocephala ?); Cordillera of Chillán, 36° 54’S; P. Germain.