Pterostichus melanarius melanarius (Illiger, 1798)

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 517

Pterostichus melanarius melanarius (Illiger, 1798)

Carabus melanarius Illiger, 1798: 163. Type locality: Prussia (inferred from title of the book). Syntype(s) probably lost (Lindroth 1966: 491).

Distribution.

This European species is adventive in North America on both coasts and is now found from Newfoundland (Lindroth 1955a: 87; Larson and Langor 1982: 592) to Vancouver Island (Lindroth 1966: 492), south to northwestern California (Del Norte County, James R. LaBonte pers. comm. 1992), northern Utah (Davis County, CMNH), northern Colorado (Miller and Peairs 2008: 34), South Dakota (Ellsbury et al. 1998: 621; French et al. 2004: 557; Larsen and Purrington 2010: 571), northeastern Iowa (Purrington and Larsen 1997: 50), east-central Ohio (Usis and MacLean 1998: 67), and central Pennsylvania (Capogreco 1989b: 4; Byers et al. 2001: 85). The first inventoried specimen collected on this continent was found in Nova Scotia in 1926 (Lindroth 1957c: 153) and in the west in Seattle in 1927 (Hatch 1933c: 120).

Records.

CAN: AB, BC (VCI), MB, NB, NF, NS (CBI), ON, PE, QC, SK USA: CA, CO, CT, IA, ID, IL, IN, MA, ME, MI, MN, MT, ND, NH, NY, OH, OR, PA, RI, SD, UT, VT, WA, WI, WY - Adventive

Note.

Two other subspecies of Pterostichus melanarius are found in Europe, Pterostichus melanarius bulgaricus Lutshnik and Pterostichus melanarius cardioderus Chaudoir (Bousquet 2003d: 499).