Dichotomius ( Cephagonus ) marcoscarvalhoi, Nunes & Vaz-de-Mello, 2019

Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), Journal of Natural History 53 (37), pp. 2231-2351 : 2324-2325

publication ID	https://doi.org/10.1080/00222933.2019.1692088
DOI	https://doi.org/10.5281/zenodo.3671930
persistent identifier	https://treatment.plazi.org/id/C740D609-2A3C-BB1A-FE81-E4ECE7B82E21
treatment provided by	Valdenar
scientific name	Dichotomius ( Cephagonus ) marcoscarvalhoi
status	sp. nov.

Treatment

3.18.: Dichotomius ( Cephagonus) marcoscarvalhoi View in CoL new species

( Figure 39 View Figure 39 (e-h))

Diagnosis. separated from other species in the quadraticeps group by the following combined characters: larger males pronotum with two central-rounded lobes on the anterior portion of the disc; posterior margin of pronotum having 1 – 2 rows of ocellate or/and coarse punctures; 1st ventrite with groups of 8 – 12 long hair at each side and paramera with rounded and convergent apexes.

● HOLOTYPE ( male): BRASIL: Mato Grosso. Curvelândia. Fazenda Aparecido. 15³33 ʹ 43 ʹ ’ Sı 58³00 ʹ 20 ” W. ii-2004. M Santos-Filho [at CEMT].

● PARATYPES: same data of Holotype [8 ƋƋı 2 ♀♀ at CEMT]; BRASIL. Mato Grosso. Araputanga. Fazenda Sudam. 15³15 ʹ 22 ” Sı 58³27 ʹ 12 ” Wı 352 m. Floresta semidecídua. Pitfall fezes humanas. 28-i-2013. RJ Silva [2 ƋƋ at CEMT]; Campos de Júlio. 30 km N de Uirapuru. 14³17 ’ Sı 59 ³15 ʹ W. xii-2002. A Foucart [1 Ƌı 3 ♀♀ at CEMT]; Mirassol do Oeste. Fazenda Santa Helena. 15³36 ʹ 34 ” Sı 57 ³58 ʹ 12 ” W. 26-i-2013. RJ Silva [ 1 ♀ at CEMT].

Description (Holotypeı male). BL: 20 mmı PW: 12 mm. Colour: blackı shiny. Head: dorsal surface smooth. Interocular space with fine punctures. Clypeo-genal angle obtuse (110³). Cephalic carina longer than wide with rounded apex. Pronotum: margins with 1 – 2 rows of ocellate punctures. Posterior border groovedı almost faced; posterior angles excavated. Anterior portion of disc having a pair of central lobes on anterior portion. Region adjacent to eyes with ocellate punctures. Metasternum: sides and margins of anterior lobe with ocellate setigerous punctures. Anterior portion of anterior lobe smooth and glabrous. Setigerous punctures equally spaced at sides. Elytra: striae deeply impressed having fine punctures spaced by twice their diameter. Interstriae disc smoothı with fine punctures (viewed under 20x magnification)ı flat; 1st interstriae strongly convexı also lacking microsculpture with very fine punctures (20 x). Abdomen: 1st ventrite with groups of 8 – 12 long hair at each side. Puncture of each ventrite interrupted medially as well groove along the ventrite posterior margin of sixth ventrite. Aedeagus ( Figure 39 View Figure 39 (f-h)): dorsallyı paramere apexes acuteı parallel; longitudinal excavation present taking whole paramera length; sides sinuous; basal rounded excavation present; anterior delimitation of basal excavations forming a pair of knobs. Laterallyı subtle declivity from basis towards apex; wide posterior rounded excavation (1/5). Ventrallyı each paramera with a lateral groove; subgenital plate widely emarginated at apex and less emarginated at basis.

Morphological variation. males BL: 17 – 20 mmı PW: 9 – 12 mm. Females BL: 17 – 19 mmı PW 9 – 11 mm. Distinguished from males as follows: Head: dorsal surface striated. Cephalic carina lowı not bifurcated apically. Pronotum: simply convexı lacking lobes. Abdomen: sixth ventrite three times larger than fifth and bearing a central pair of tubercles. Some individuals (males or females) mesepimeron may have dense setaeı concealing the structure.

Distribution. Brazilı state of Mato Grosso ( Figure 36 View Figure 36 ).

Etymology. named after our friend Dr. Marcos André de Carvalhoı director of Instituto de Biociências (Universidade Federal de Mato Grossoı Brasil) for many years.