Campanula tatrae tatrae Borbas , Magyar Bot. Lapok 1: 319 (1902)

Novikov, Andriy, 2023, An annotated nomenclatural checklist of endemic vascular plants distributed in the Ukrainian Carpathians, Biodiversity Data Journal 11, pp. 103921-103921 : 103921

Campanula tatrae tatrae Borbas, Magyar Bot. Lapok 1: 319 (1902)

Campanula kladniana = Campanula kladniana subsp. polymorpha Witasek, Magyar Bot. Lapok 5: 239 (1906); GBIF: https://www.gbif.org/species/5410038; IPNI: https://www.ipni.org/n/77255487-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000827554; POWO: https://powo.science.kew.org/taxon/77255487-1; BHL: https://www.biodiversitylibrary.org/page/50344447#page/645; JACQ: https://wu.jacq.org/WU0071526

Campanula kladniana = Campanula kladniana var. polymorpha (Witasek) Pawł., Acta Soc. Bot. Pol. 1: 5 (1923); GBIF: https://www.gbif.org/species/8038018; WFO: http://www.worldfloraonline.org/taxon/wfo-0000827555; POWO: https://powo.science.kew.org/taxon/364966-4

Campanula kladniana = Campanula kladniana subsp. stenophylla (Schur) Witasek, Magyar Bot. Lapok 5: 238 (1906); GBIF: https://www.gbif.org/species/5410040; IPNI: https://www.ipni.org/n/77256020-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000827556; POWO: https://powo.science.kew.org/taxon/77256020-1; BHL: https://www.biodiversitylibrary.org/page/50344446#page/644

Campanula polymorpha = Campanula polymorpha (Witasek) Prain, Index Kew. Suppl. 4: 35 (1913), non Banks & Sol. ex A.DC. *; GBIF: https://www.gbif.org/species/5410057; IPNI: https://www.ipni.org/n/77237988-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828219; POWO: https://powo.science.kew.org/taxon/77237988-1; BHL: https://www.biodiversitylibrary.org/item/133932#page/47

Campanula polymorpha = Campanula polymorpha var. intercedens Hruby, Magyar Bot. Lapok 29: 199 (1930), non C. witasekiana var. intercedens Hruby; GBIF: https://www.gbif.org/species/5410061; IPNI: https://www.ipni.org/n/77266950-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828224; POWO: https://powo.science.kew.org/taxon/77266950-1

Campanula polymorpha = Campanula polymorpha var. intercedens f. angustifolia Hruby, Magyar Bot. Lapok 29: 200 (1930); GBIF: https://www.gbif.org/species/5410043; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828220; POWO: https://powo.science.kew.org/taxon/365601-4

Campanula polymorpha = Campanula polymorpha var. intercedens f. exigua Hruby, Magyar Bot. Lapok 29: 200 (1930); GBIF: https://www.gbif.org/species/5410058; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828222; POWO: https://powo.science.kew.org/taxon/365603-4

Campanula polymorpha = Campanula polymorpha var. intercedens f. latifolia Hruby, Magyar Bot. Lapok 29: 199 (1930); GBIF: https://www.gbif.org/species/5410046; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828226; POWO: https://powo.science.kew.org/taxon/365607-4

Campanula polymorpha = Campanula polymorpha var. intercedens f. reflectans Hruby, Magyar Bot. Lapok 29: 200 (1930); GBIF: https://www.gbif.org/species/5410039; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828233; POWO: https://powo.science.kew.org/taxon/365614-4

Campanula polymorpha = Campanula polymorpha var. intercedens f. umbrosa Hruby, Magyar Bot. Lapok 29: 200 (1930); GBIF: https://www.gbif.org/species/5410036; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828236; POWO: https://powo.science.kew.org/taxon/365619-4

Campanula polymorpha = Campanula polymorpha var. lepida Nyár. ex Hruby, Magyar Bot. Lapok 29: 201 (1930)

Campanula polymorpha = Campanula polymorpha var. pluriflora Nyár. ex Hruby, Magyar Bot. Lapok 29: 198 (1930)

Campanula polymorpha = Campanula polymorpha var. praticola Hruby, Magyar Bot. Lapok 29: 198 (1930), non C. witasekiana var. praticola Hruby; GBIF: https://www.gbif.org/species/5410053; IPNI: https://www.ipni.org/n/77268347-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828231; POWO: https://powo.science.kew.org/taxon/77268347-1

Campanula polymorpha = Campanula polymorpha var. praticola f. hirta ( Nyár.) Hruby, Magyar Bot. Lapok 29: 199 (1930)

Campanula polymorpha = Campanula polymorpha var. praticola f. pluriflora ( Nyár.) Hruby, Magyar Bot. Lapok 29: 199 (1930)

Campanula polymorpha = Campanula polymorpha var. stenophylla (Schur) Hruby, Magyar Bot. Lapok 29: 205 (1930); GBIF: https://www.gbif.org/species/5410054; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828235; POWO: https://powo.science.kew.org/taxon/365616-4

Campanula polymorpha = Campanula polymorpha var. stenophylla f. brachyphylla Hruby, Magyar Bot. Lapok 29: 206 (1930); GBIF: https://www.gbif.org/species/5410056; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828221; POWO: https://powo.science.kew.org/taxon/365602-4

Campanula polymorpha = Campanula polymorpha var. stenophylla f. genuina Hruby, Magyar Bot. Lapok 29: 206 (1930)

Campanula polymorpha = Campanula polymorpha var. stenophylla f. gracilis Hruby, Magyar Bot. Lapok 29: 206 (1930); GBIF: https://www.gbif.org/species/5410050; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828223; POWO: https://powo.science.kew.org/taxon/365604-4

Campanula polymorpha = Campanula polymorpha var. typica Hruby, Magyar Bot. Lapok 29: 198, 201 (1930); GBIF: https://www.gbif.org/species/5410048; POWO: https://powo.science.kew.org/taxon/365617-4

Campanula polymorpha = Campanula polymorpha var. typica f. fasciculata Nyár. ex Hruby, Magyar Bot. Lapok 29: 202 (1930)

Campanula polymorpha = Campanula polymorpha var. typica f. fasciculata subf. deltoidea Hruby, Magyar Bot. Lapok 29: 203 (1930)

Campanula polymorpha = Campanula polymorpha var. typica f. kladnianioides Nyárady ex Hruby, Magyar Bot. Lapok 29: 201 (1930); GBIF: https://www.gbif.org/species/8517923; GBIF: https://www.gbif.org/species/5410051; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828225; POWO: https://powo.science.kew.org/taxon/365606-4

Campanula polymorpha = Campanula polymorpha var. typica f. latifolia Hruby, Magyar Bot. Lapok 29: 201 (1930); GBIF: https://www.gbif.org/species/5410046; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828226; POWO: https://powo.science.kew.org/taxon/365607-4

Campanula polymorpha = Campanula polymorpha var. typica f. latifolia subf. umbrosa Hruby, Magyar Bot. Lapok 29: 202 (1930); GBIF: https://www.gbif.org/species/5410036; POWO: https://powo.science.kew.org/taxon/365618-4

Campanula polymorpha = Campanula polymorpha var. typica f. lepida ( Nyár.) Hruby, Magyar Bot. Lapok 29: 202 (1930); GBIF: https://www.gbif.org/species/5410037; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828227; POWO: https://powo.science.kew.org/taxon/365608-4

Campanula polymorpha = Campanula polymorpha var. typica f. lepida subf. reflectans Hruby, Magyar Bot. Lapok 29: 202 (1930); GBIF: https://www.gbif.org/species/5410039; POWO: https://powo.science.kew.org/taxon/365613-4

Campanula polymorpha = Campanula polymorpha var. typica f. saxiphila Hruby, Magyar Bot. Lapok 29: 204 (1930)

Campanula polymorpha = Campanula polymorpha var. typica f. saxiphila subf. reflectans Hruby, Magyar Bot. Lapok 29: 205 (1930)

Campanula rotundifolia = Campanula rotundifolia subsp. polymorpha (Witasek) Tacik in Jasiewicz, Monogr. Bot. 20: 254 (1965); GBIF: https://www.gbif.org/species/7685150; IPNI: https://www.ipni.org/n/77255879-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828685; POWO: https://powo.science.kew.org/taxon/77255879-1

Campanula scheuchzeri = Campanula scheuchzeri [unranked] β Campanula dacica Porcius, Enum. Pl. Phan. Naszód.: 37 (1878)

Campanula scheuchzeri = Campanula scheuchzeri var. dacica Porcius, Fl. Naseud.: 98 (1885); GBIF: https://www.gbif.org/species/5410055; IPNI: https://www.ipni.org/n/77268262-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828824; POWO: https://powo.science.kew.org/taxon/77268262-1

Campanula scheuchzeri = Campanula scheuchzeri var. stenophylla Schur, Enum. Pl. Transsilv.: 443 (1866); GBIF: https://www.gbif.org/species/5410059; IPNI: https://www.ipni.org/n/77268259-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828864; POWO: https://powo.science.kew.org/taxon/77268259-1; BHL: https://www.biodiversitylibrary.org/page/10544494#page/465

Campanula stenophylla = Campanula stenophylla (Schur) Prain, Index Kew. Suppl. 4: 35 (1913) [nom. inval.]; GBIF: https://www.gbif.org/species/5410042; IPNI: https://www.ipni.org/n/77238117-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000829052; POWO: https://powo.science.kew.org/taxon/77238117-1; BHL: https://www.biodiversitylibrary.org/item/133932#page/47

Campanula stenophylla = Campanula stenophylla (Schur) Witasek, Magyar Bot. Lapok 5: 238 (1906), non Boiss. & Heldr.; GBIF: https://www.gbif.org/species/7633288; IPNI: https://www.ipni.org/n/141149-1; BHL: https://www.biodiversitylibrary.org/item/202161#page/644

Campanula carnica - Campanula carnica auct. fl. transsilv., non Schiede

Campanula consanguinea - Campanula consanguinea Simonk., Enum. Fl. Transsilv.: 385 (1886) [p. p.], non Schott; BHL: https://www.biodiversitylibrary.org/item/40105#page/449

Campanula kladniana - Campanula kladniana (Schur) Witasek, Abh. Zool.-Bot. Ges. Wien 1: 39 (1902) [p. p. min., non sensu Schur orig.]; GBIF: https://www.gbif.org/species/5411315; IPNI: https://www.ipni.org/n/140522-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000827551; POWO: https://powo.science.kew.org/taxon/140522-1

Campanula linifolia - Campanula linifolia auct. [e.g., Wahlenb.], non Jacq.

Campanula polymorpha - Campanula polymorpha f. sciaphila Hruby, Magyar Bot. Lapok 29: 204 (1930) [nom. et des. inval.]; GBIF: https://www.gbif.org/species/5410041; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828234; POWO: https://powo.science.kew.org/taxon/365615-4

Campanula rotundifolia - Campanula rotundifolia L., Sp. Pl. 1: 163 (1753) [p. p. Festuca minor , tantum quod plantas ucrain. carpat.], non alior *; GBIF: https://www.gbif.org/species/5410907; IPNI: https://www.ipni.org/n/30036649-2; WFO: http://www.worldfloraonline.org/taxon/wfo-0000828550; POWO: https://powo.science.kew.org/taxon/30036649-2; BHL: https://www.biodiversitylibrary.org/page/358182#page/175

Campanula pusilla - Campanula pusilla auct. fl. ucrain. carpat., non Haenke

Campanula scheuchzeri - Campanula scheuchzeri auct. [e.g., Reuss, Května Slov.: 278 (1853); Sagorski & Schneider, Fl. Centralkarp.: 369 (1891)], non Vill. *

Conservation status

In Ukraine - LC ( Onyshchenko et al. 2022).

Distribution

Pancarpathian endemic.

Notes

In general, there are three subspecies of C. tatrae (https://powo.science.kew.org/taxon/141196-1, accessed on 05.06.2023) - subsp. Campanula tatrae tatrae , subsp. Campanula tatrae mentiens (Witasek) Kovanda and subsp. Campanula mentiens sudetica (Hruby) Kovanda. The last two subspecies occur exclusively in the Western Carpathians; therefore, only C. tatrae subsp. tatrae is represented in the flora of the Ukrainian Carpathians.

However, C. tatrae subsp. tatrae has a long and complex naming history. In the Ukrainian Carpathians, it combines two main taxa that are often distinguished as independent species (i.e. C. polymorpha (Witasek) Prain and C. rotundifolia L., p.p.) by certain authors (e.g. Dremliuga and Zyman (2012), Chopyk and Fedoronchuk (2015)). However, Kovanda (1975) conducted nomenclatural explorations and stressed the taxonomic concept that Witaseks applied to Schur's C. kladniana in her early publication ( Witasek 1902). To avoid misinterpretation of plants from the Tatrae Mts. that were often incorrectly identified as C. kladniana (Schur) Witasek and synonymised with C. polymorha , Kovanda (1975) proposed using the name C. tatrae that has been published earlier by Borbás (1902) and had an unambiguous sense. Many infraspecific taxa from Campanula scheuchzeri auct., non Vill. seem to be a part of C. tatrae subsp. tatrae too ( Geslot 1971). On the other hand, in the recent checklists (e.g. POWO, WFO and Euro+Med - https://powo.science.kew.org/taxon/77168970-1, https://list.worldfloraonline.org/wfo-0000838737, https://europlusmed.org/cdm_dataportal/taxon/c454d5af-c66c-4d0a-b009-a0e8754e9c9c, all databases being accessed on 05.06.2023), C. scheuchzeriformis Hayek and derived C. balcanica var. scheuchzeriformis (Hayek) Hruby are wrongly listed amongst synonyms of C. tatrae subsp. Campanula tatrae tatrae . Campanula Festuca scheuchzeriformis occurs in North Albania ( Hayek 1921, Hruby 1930). Such synonymisation probably results from confusion because Hayek (1921), in the protologue of C. scheuchzeriformis , mentioned that it is similar to C. scheuchzeri. Another taxon, C. rotundifolia var. alpicola Hayek (≡ C. rotundifolia f. alpicola (Hayek) Hruby), is also mistakenly indicated as a synonym of C. tatrae subsp. tatrae in the Worldplants checklist (https://www.worldplants.de/world-plants-complete-list/complete-plant-list?name=Campanula-tatrae, accessed on 07.06.2023) and GBIF (https://www.gbif.org/species/7222073, accessed on 05.06.2023). Finally, Worldplants wrongly indicate the presence of C. carnica subsp. carnica Mert. & W.D.J.Koch for Ukraine. This species and subspecies do not occur in Ukraine. Such confusion of C. rotundifolia f. alpicola probably results from the mistaken synonymisation of C. kladniana with C. carnica subsp. carnica due to ambiguous interpretation of the taxonomic limits of C. kladniana ( Podlech 1965, Kovanda 1975).