Tylodinus pinguis Luna-Cozar, 2014

Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. & León-Cortés, Jorge L., 2014, A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico, Zootaxa 3788 (1), pp. 1-63 : 62

36. Tylodinus pinguis Luna-Cozar , sp. nov.

( Figures 96–97 , 129 , 135 )

Diagnosis. Length male 4.8–4.9 mm. Width male 2.2–2.3 mm. Body 2.2x longer than wide, black. Head clothed with dark brown scales, punctures similar to punctures on rostrum; vertex with two patches of overlapping scales, whitish or pale yellow color, light brown on lateral margin; frons moderately concave; rostrum moderately robust, weakly carinate, apical area punctuate, eyes moderately convex. Pronotum in lateral view strongly convex, outline with anterior constriction, without posterior constriction ( Figures 10 , 97 ), in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc somewhat hollowed medially, punctures dense to contiguous at base and progressively more widely spaced, smaller and shallower to apex ( Figures 14 , 96 ); surface granulate. Elytra with tubercles as follows: I2 with round and large tubercle at declivity base; I3 with two tubercles, first subbasal, second smaller, round at middle of elytra; I4 with laminate tubercle beyond declivity; I5 with two small tubercles in same position as tubercles on I3; I7 with two tubercles in same position as tubercles on I5. Metasternum moderately concave, median section glabrous. Male abdomen with ventrite 1 moderately concave, squamous; ventrite 2 feebly concave, longer than ventrites 3–4 combined, squamous; ventrite 5 flat, without impressions, as long as ventrites 1–2. Femora armed, hind femur reaching elytral apex, tibiae with inner margin slightly sinuate, premucro present. Male genitalia ( Figure 129 ) with median lobe in lateral view weakly curved, slender; in ventral view as long as apodemes, lateral sides moderately swollen at 0.25 length from the base, convergent, apex acute; apical process in lateral view moderately large (0.26–0.34 as long as body of median lobe), straight, moderately curved downward; in ventral view not constricted beyond median orifice, lateral sides convergent.

Geographic distribution. México (Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2000–2200 m.

Derivation of specific name. Name derived from the Latin word pinguis = fat.

Material examined. Total 2 males. Holotype male ( CMNC): México: Chiapas, Angel Albino Corzo, Reserva El Triunfo , Polígono 1, elev. 2050 m (15°39'25.56''N, 92°48'32.4''W), 16–21.xi.2002, coll. R . S. Anderson. Paratype: México: Chiapas, La Concordia, 4.3 km SE Custepec, Campamento El Quetzal , elev. 2160 m (15°42'N, 92°55'59.9''W), 18.vii.2007, coll. M. G. Branstetter, 1 male ( CMNC) .

Figures

FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe.

FIGURES 120–129. Ventral and lateral habitus of median lobe: 120. Tylodinus dominicus, 121. Tylodinus noctis, 122. Tylodinus kuscheli, 123. Tylodinus pappi, 124. Tylodinus branstetteri, 125. Tylodinus immundus, 126. Tylodinus intzin, 127. Tylodinus gibbosus, 128. Tylodinus kissingeri, 129. Tylodinus pinguis.

FIGURES 134–135. Maps showing collecting sites. 134. Occurrences in Chiapas, México of: Tylodinus rufus, Tylodinus kuscheli, Tylodinus pappi, Tylodinus jonesi, Tylodinus coapillensis; 135. Occurrences in Chiapas, México of: Tylodinus spiniventris, Tylodinus lum, Tylodinus noctis, Tylodinus pinguis, Tylodinus rugosus.

FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic).

FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth.