Ceratopogon aeschnosuga de Meijere
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publication ID |
https://doi.org/ 10.11646/zootaxa.5261.1.1 |
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publication LSID |
lsid:zoobank.org:pub:06188017-222F-47B8-AF52-1127FBB3601B |
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persistent identifier |
https://treatment.plazi.org/id/C05987B9-FF8D-FFBD-FF67-CF0510F026BB |
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treatment provided by |
Plazi |
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scientific name |
Ceratopogon aeschnosuga de Meijere |
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Subgenus PTEROBOSCA Macfie
PTEROBOSCA Macfie, 1932: 266. Unavailable name; proposed after 1930 without type species designation. PTEROBOSCA Macfie, 1940c: 16. Type species: Ceratopogon aeschnosuga de Meijere View in CoL , by original designation.
incubans (Macfie), 1937b: 111 View in CoL (Pterobosca; female). Type locality: Belize, Cayo District, Mtn. Pine Ridge. ST F (BMNH). Distr.: Mexico (San Luis Potosi, Veracruz), Puerto Rico to Argentina (Buenos Aires: Otamendi, Estación Experimental INTA-Delta, 34°10′30.44″S, 58°51′39.75″W; Tigre, 34°25′29.03″S, 58°32′31.80″W; Campana, Paraná de las Palmas, Isla del INTA, 34°10′27.09″S, 58°52′01.17″W). Refs.: Cavalieri & Chiossone, 1965: 65; Marino & von Ellenrieder, 1999: 39.
Subgenus SALIOHELEA Wirth & Ratanaworabhan
SALIOHELEA Wirth & Ratanaworabhan, 1978: 494 (as subgenus of Forcipomyia View in CoL ). Type species: Forcipomyia leei Wirth & Ratanaworabhan View in CoL , by original designation.
leei Wirth & Ratanaworabhan, 1978: 498 View in CoL (male, female). Type locality: Colombia, Valle, Río Raposo. HT F (USNM). Distr.: Eastern USA and Caribbean to southern Brazil (Amazonas, Pará, Bahia, Rio de Janeiro), Argentina (Buenos Aires: Ensenada, Reserva Natural Integral Punta Lara, marginal forest, 34º47′21.2′′S, 57º59′56.8′′W; Reserva Parque Provincial Ernesto Tornquist, unnamed stream, 38º01′09.4′′S, 62º00′56.6′′W). Refs.: Marino et al., 2011a: 202; Cazorla et al., 2018: 6.
New record: Misiones: Aristóbulo del Valle, Arroyo Cuña Pirú, 27°05′16.30″S, 54°57′08.64″W GoogleMaps .
Subgenus SYNTHYRIDOMYIA Saunders
SYNTHYRIDOMYIA Saunders, 1957: 688 (as subgenus of Forcipomyia ). Type species: Lasiohelea acidicola Tokunaga , by original designation.
murina (Winnertz), 1852: 26 ( Ceratopogon murinus ; male, female). Type locality: Germany. ST F (BMNH). Distr.: Widely spread in the Holarctic and Afrotropical regions, reported also from the Oriental region (Taiwan) and in the Neotropical region only in Argentina ( Salta: Cafayate , motel ACA, 26°03′54.14′′S, 65°58′21.99″W; Cachi, 25°07′05.72″S, 66°09′47.78″W. La Rioja: Río Miranda at NRt. 40, 29º20′50.4′′S, 67º42′12.8′′W, 1590 m; Chilecito, 29°10′27.22″S, 67°28′45.61″W. Santiago del Estero: Añatuya, 28°27′38.08″S, 62°50′03.71″W. Córdoba: paraje Las Jarillas, Río San Antonio, 31º32′02.8′′S, 64º33′01.7′′W, 820 m; Huerta Grande, Río Suquía at Arroyo Huerta Grande, 31º05′00.6′′S, 64º30′05.2′′W, 934 m. Corrientes: Esteros del Iberá, Colonia Pellegrini, 28°32′17.58″S, 57°10′56.86″W. Buenos Aires: Alberti, 35°01′28.63″S, 60°15′36.92″W. Río Negro: Meseta de Somuncura, Chipauquil, spring Arroyo Valcheta, 40°58′25′′S, 66°39′14.7′′W, 630 m; Meseta de Somuncura, Estancia El Rincón, spring Arroyo Valcheta, 40°59′24.1′′S, 66°40′35.7′′W, 620 m; Cinco Saltos, 38°49′S, 68°04′′W). Refs.: Spinelli & Marino, 2018: 3.
calchaqui Spinelli & Marino, 1997: 188 View in CoL (as belonging to subgenus Thyridomyia ). Type locality: Argentina, Salta, Cafayate, motel ACA , 26°03′54.14′′S, 65°58′21.99″W. HT F (MLPA).
soibelzoni Marino & Spinelli, 2001c: 14 View in CoL . Type locality: Argentina, Río Negro GoogleMaps , Meseta de Somuncura GoogleMaps , Chipauquil, spring Arroyo Valcheta, 40°58′25′′S, 66°39′14.7′′W, 630 m. HT F (MLPA)
sanctaeclarae Wirth, 1952b: 90 View in CoL (male, female). Type locality: Chile, Juan Fernández Islands, Santa Clara, El Corral. HT M (MUNS) View in CoL . Distr.: Chile, Argentina (Río Negro: Meseta de Somuncura, Estancia La Rinconada, springs Arroyo Comi-Có, 41°08′35.1′′S, 67°27′34.6′′W, 1000 m. Chubut: Meseta de la Sierra Cuadrada, Estancia Don Eduardo, 44º39′25.9′′S, 68º00′09.9′′W. Santa Cruz: Río Pinturas, 47º09′21.8′′S, 70º39′31′′W, 570 m). Refs.: Marino & Spinelli, 2001c: 14; Muzón et al., 2010: 115.
New records: Jujuy: Arroyo Condorcito at NRt. 9, 119 km S La Quiaca, 22°56′53.82″S, 65°24′54.44″W GoogleMaps . Salta: Cafayate, 26°03′54.14′′S, 65°58′21.99″W. La Rioja: Refugio del Peñon , 28°28′31.2″S, 68°50′13.5″W, 3600 m GoogleMaps ; Chilecito , 29°10′27.22″S, 67°28′45.61″W GoogleMaps ; Río Miranda at NRt. 40, 29º20′50.4′′S, 67º42′12.8′′W, 1590 m GoogleMaps . Mendoza: Mendoza city, Zoo , 32°53′03.75″S, 68°53′20.74″W GoogleMaps .
Subgenus THYRIDOMYIA Saunders
THYRIDOMYIA Saunders, 1925: 268 . Type species: Thyridomyia palustris Saunders View in CoL (= Ceratopogon monilicornis Coquillett View in CoL ), by original designation.
REFERENCE: Wirth, 1970 (key to males and females).
nana (Macfie), 1939b: 171 View in CoL ( Lasiohelea View in CoL ; female). Type locality: Brazil, Santa Catarina, Nova Teutônia. HT F (BMNH) View in CoL . Distr.: Mexico (Veracruz), Jamaica through Caribbean to southern Brazil (Santa Catarina), Argentina (Buenos Aires: Ensenada, Reserva Natural Integral Punta Lara, marginal forest, 34º47′21.2′′S, 57º59′56.8′′W). Refs.: Spinelli et al., 2010: 130; Cazorla et al., 2018: 6.
riojana Spinelli & Marino, 1997: 188 View in CoL (male). Type locality: Argentina, La Rioja, Chilecito View in CoL , 29°10′27.22″S, 67°28′45.61″W. HT M (MLPA). Distr.: Argentina (La Rioja: type locality. Río Negro: Meseta de Somuncura, Arroyo Comi-Có, 41°06′23.7′′S, 67°27′43.1′′W, 870 m; Meseta de Somuncura, Estancia El Rincón, spring Arroyo Valcheta, 40°59′24.1′′S, 66°40′35.7′′W, 620 m; Meseta de Somuncura, Chipauquil, spring Arroyo Valcheta, 40°58′25′′S, 66°39′14.7′′W, 630 m. Chubut: Meseta de la Sierra Cuadrada, Estancia Don Eduardo, 44º39′25.9′′S, 68º00′09.9′′W). Refs.: Marino & Spinelli, 2001c: 16; Muzón et al., 2005: 62.
New records: La Rioja: Río Miranda at NRt. 40, 29º20′50.4′′S, 67º42′12.8′′W, 1590 m GoogleMaps . San Luis: PRt. nº 5, Bajo de Veliz , 32º08′48.2′′S, 65º24′43.3′′W, 583 m. GoogleMaps Córdoba: paraje Las Jarillas, Río San Antonio , 31º32′02.8′′S, 64º33′01.7′′W, 820 m. GoogleMaps
Subgenus TRICHOHELEA Goetghebuer
APELMA Kieffer, 1919: 64 (preoccupied by Billberg, 1820). Type species: Apelma auronitens Kieffer View in CoL (= Trichohelea tonnoiri Goetghebuer View in CoL ), designation by Macfie, 1940c: 16.
TRICHOHELEA Goetghebuer, 1920: 39 . Type species: Trichohelea tonnoiri Goetghebuer View in CoL , by original designation.
REFERENCE: Marino & Spinelli, 2004b (key to males and females from Patagonia).
goniognatha Wirth and Messersmith, 1971: 20 View in CoL (male, female). Type locality: Mexico GoogleMaps , Sonora GoogleMaps , 5 mi W Alamos. HT F (USNM). Distr.: USA ( Arizona GoogleMaps , Kansas GoogleMaps , Texas GoogleMaps ), Mexico GoogleMaps ( Baja California GoogleMaps , Sonora GoogleMaps ), Argentina GoogleMaps ( Córdoba GoogleMaps : Capilla del Monte, 30°51′31.55″S, 64°32′26.90″W). First Argentina record.
leptognatha Wirth and Messersmith, 1971: 21 View in CoL (male, female). Type locality: USA, Kansas, Pottawatomie Co. HT F (USNM). Distr.: USA ( Arizona GoogleMaps , Iowa GoogleMaps , Kansas GoogleMaps ), Mexico GoogleMaps ( Sonora GoogleMaps ), Argentina GoogleMaps ( Misiones GoogleMaps : Corpus GoogleMaps , 27°06′22.43′′S, 55°31′24.71″W. Corrientes: Garapé GoogleMaps , 27°36′05.30″S, 56°14′44.44″W). First Argentina records.
limnetis Ingram & Macfie, 1931: 169 View in CoL (male). Type locality: Argentina, Neuquén, PN Nahuel Huapi, Lago Correntoso, 40°43′58.90″S, 71°40′15.97″W. HT M (BMNH). Distr.: Argentina (Neuquén: type locality; Arroyo Pedregoso at PRt. 65, 5 km E El Portezuelo, 40°37′27.18″S, 71°35′29.45″W. Río Negro: Arroyo Challhuaco, 41°13′14.15″S, 71°17′56.71″W; Lago Guillelmo, 41°22′42.08″S, 71°30′26.30″W). Refs.: Ingram & Macfie, 1931: 169; Marino & Spinelli, 2004b: 2254 View Cited Treatment .
shannoni (Ingram & Macfie), 1931: 171 View in CoL ( Lasiohelea View in CoL ). Type locality: Argentina, Neuquén, PN Nahuel Huapi, Lago Correntoso, 40°43′58.90″S, 71°40′15.97″W. HT F (BMNH).
sayhuequei Marino & Spinelli, 2004b: 2257 View in CoL View Cited Treatment (female). Type locality: Argentina, Chubut, PN Los Alerces, El Alerzal, 42°36′40.50″S, 71°53′29.78″W. HT F (MLPA). Distr.: Chile, Argentina (Chubut: type locality). Refs.: Marino & Spinelli, 2004b: 2257 View Cited Treatment .
tehuelche Marino & Spinelli, 2004b: 2260 View in CoL View Cited Treatment (male, female). Type locality: Argentina, Santa Cruz, Lago del Desierto, 49°05′0.71′′S, 72°53′32.61″W. HT M (MLPA). Distr.: Argentina (Santa Cruz: type locality). Refs.: Marino & Spinelli, 2004b: 2260 View Cited Treatment .
New record: Neuquén: Laguna Epulaufquen , 36°50′09.49′′S, 71°01′01.08′′W GoogleMaps .
Subgenus WARMKEA Saunders
WARMKEA Saunders, 1957: 671 (as subgenus of Forcipomyia View in CoL ). Type species: Forcipomyia bicolor Saunders View in CoL (= Forcipomyia lesliei Wirth View in CoL ), by original designation.
REFERENCE: Wirth & Soria, 1980 (key to males and females).
aeria Saunders, 1957: 675 View in CoL (larva, pupa, male, female). Type locality: Puerto Rico, Mayaguez. HT F (CNCI). Distr.: Puerto Rico, Colombia, Argentina (Tucumán: Finca Santa Bárbara (?)). Refs.: Wirth & Soria, 1980: 143.
New record: Misiones: Posadas , 27°24′26.66″S, 55°57′29.29″W GoogleMaps .
tuberculata Saunders, 1957: 677 View in CoL (larva, pupa, male, female). Type locality: Trinidad and Tobago, North Range. HT F (CNCI). Distr.: Costa Rica, Colombia, Trinidad and Tobago, Brazil (Amazonas, Pará), Argentina (Buenos Aires: Berisso, Los Talas, 34°52′16.47′′S, 57°51′08.76′′W). Refs.: Spinelli & Balseiro, 1982: 226.
New records: Misiones: Arroyo Anyico at km 813 NRt. 14, 27°39′52.76″S, 55°36′31.35″W GoogleMaps . Buenos Aires: Estación Otamendi, INTA Delta , 34°10′30.44″S, 58°51′39.75″W GoogleMaps .
SUBFAMILY CERATOPOGONINAE NEWMAN, 1834: 388 View in CoL
TRIBE CULICOIDINI KIEFFER, 1911c: 1, 1911b: 319 View in CoL
Genus CULICOIDES Latreille View in CoL
CULICOIDES Latreille, 1809: 251 View in CoL . Type species: Culicoides punctatus Latreille (= Ceratopogon punctatus Meigen View in CoL ), by monotypy.
PADROSIA Rafinesque, 1815: 130 (unnecessary replacement name for Culicoides Latreille View in CoL ). Type species: Culicoides punctatus Latreille (= Ceratopogon punctatus Meigen View in CoL ), automatic.
REFERENCES: Forattini, 1957 (key to females of Neotropical species); Wirth & Blanton, 1959 (key to females and some males from Panama); Wirth & Blanton 1973 (key to females and some males from Amazon Basin); Wirth & Blanton, 1974 (key to females and some males from the Caribbean); Aitken et al., 1975 (key to females from Trinidad and Tobago); Blanton & Wirth, 1979 (key to larvae, males and females from Florida ( USA)); Spinelli & Wirth, 1986 (key to females south of the Amazon Basin); Wirth et al., 1988 (some meristic character states and photographs of wings of all species); Huerta, 1996 (key to females from Mexico); Ronderos & Spinelli, 1998 (key to females and some males from the Yacyretá Dam, between Argentina and Paraguay); Spinelli et al., 2005a (key to females and some males from Argentina).
There is no reliable key to the subgenera and species groups of Neotropical Culicoides View in CoL . For species outside of local areas for which revisions are available (as listed above), the easiest guide to species identification is that of Wirth et al., 1988. From there readers may refer to keys to subgenera or species groups referred to below.
Subgenus ANILOMYIA Vargas
ANILOMYIA Vargas, 1960: 37 (as subgenus of Culicoides View in CoL ). Type species: Culicoides covagarciai Ortíz View in CoL , by original designation.
REFERENCES: Wirth & Blanton, 1956 (key to males and females of covagarciai View in CoL group); Wirth & Blanton, 1970 (key to key to males and females of nigrigenus View in CoL group).
nigrigenus Wirth & Blanton, 1956: 222 View in CoL (male, female). Type locality: Panama, Bocas del Toro, Almirante. HT F (USNM). Distr.: Mexico (Veracruz) to Colombia, Trinidad, Argentina ( Salta: PN El Rey , 24°41′60.00″S, 64°37′60.00″W). Refs.: Ronderos & Spinelli, 1990: 85.
Subgenus AVARITIA Fox
AVARITIA Fox, 1955: 218 (as subgenus of Culicoides View in CoL ). Type species: Ceratopogon obsoletus Meigen View in CoL , by original designation.
REFERENCES: Rodríguez & Wirth, 1986 (revision of andicola View in CoL group); Wirth & Mullens, 1992 (key to males and females in pusillus View in CoL group).
pusillus Lutz, 1913: 52 View in CoL (male, female). Type locality: Brazil, Rio de Janeiro, Manguinhos. Type material (?). Distr.: USA (Florida), Mexico (Chiapas) to Argentina (Misiones: Corpus, 27°06′22.43′′S, 55°31′24.71″W; Candelaria, 27°27′33.49″S, 55°45′28.76″W; Posadas, Arroyo Mártires, 27°24′26.66″S, 55°57′29.29″W). Refs.: Ronderos & Spinelli, 1998: 82; Ronderos et al., 2003b: 22.
New records: Misiones: Posadas, club San Francisco, 27°23′31.20″S, 55°55′59.37″W GoogleMaps ; Posadas , Arroyo Zaiman , 27°25′46.64″S, 55°54′8.56″W GoogleMaps ; Posadas , airport, 27°22′18.82″S, 55°58′44.36″W GoogleMaps ; Arroyo Anyico at km 813 NRt. 14, aprox. 30 km N San José, 27°39′52.76″S, 55°36′31.35″W GoogleMaps . Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W GoogleMaps .
Subgenus COTOCRIPUS Brèthes
COTOCRIPUS Brèthes, 1912: 451 . Type species: Cotocripus caridei Brèthes View in CoL , by monotypy.
bambusicola Lutz, 1913: 62 View in CoL (female). Type locality: Brazil, Rio de Janeiro, Petrópolis. ST F (FIOC). Distr.: Colombia, Venezuela, Brazil (Bahia, Espírito Santo, Rio de Janeiro, S„o Paulo), Argentina (Misiones: PN Iguazú, 25°39′S, 54°18′W; Corpus, Puerto Maní, 27°06′22.43′′S, 55°31′24.71″W; Parque Provincial Saltos del Moconá, 27°09′10.38′′S, 53°53′29.00′′W. Corrientes: Garapé, 27°36′05.30″S, 56°14′44.44″W; Estación Biológica Corrientes (EBCo), 27º32′51.8′′S, 58º40′44.8′′W, 52 m. Buenos Aires: City Bell, 34°52′55.29″S, 58°02′26.47″W; Gonnet, 34°53′20.26″S, 58°01′17.15″W). Refs.: Ronderos & Spinelli, 2000: 134; Ronderos & Spinelli, 2002: 93; Ronderos & Díaz, 2002: 44; Campos et al., 2011: 115; Marino et al., 2013a: 778 View Cited Treatment .
bahiensis Barbosa, 1947: 11 View in CoL . Type locality: Brazil, Bahia. ST F, M (USNM).
New record: Misiones: Posadas, Arroyo Zaimán , 27°25′46.64″S, 55°54′08.56″W GoogleMaps .
caridei (Brèthes), 1912: 451 View in CoL ( Cotocripus ; female). Type locality: Argentina, Buenos Aires, Berazategui, 34°45′26.67″S, 58°11′24.11″W. Type material (apparently lost). Distr.: Southeastern Brazil (Rio Grande do Sul), Argentina (Jujuy: Departamento Ledesma, Yuto, 23°38′S, 64°28′W; Libertador General San Martín, 23°48′S, 64°48′W. Buenos Aires: type locality; Isla Martín García, urban area, 34°11′08.00″S, 58°15′2.23″W; Isla Martín García, garbage place, 34°10′56.94″S, 58°15′27.91″W; Ensenada, Reserva Natural Integral Punta Lara, marginal forest, 34º47′21.2′′S, 57º59′56.8′′W; same data except grassland, 34°48′01.36″S, 58°00′26.26″W; Berisso, Los Talas, 34°52′16.47′′S, 57°51′08.76′′W; Magdalena, Arroyo Zapata, 35°08′29.55′′S, 57°26′32.17′′W, 32 m; Gonnet, 34°53′20.26″S, 58°01′17.15″W; Coronel Pringles, 38°00′11.12″S, 61°21′47.93″W), Uruguay. Refs.: Brèthes, 1912: 451; Cavalieri & Chiossone, 1966b: 147; Spinelli & Wirth, 1986: 59; Spinelli et al., 1989: 735; Ronderos & Spinelli, 1990: 83; Ronderos et al., 2011: 1187; Veggiani Aybar et al., 2015: 634; Cazorla et al., 2018: 6.
setifer (Lutz), 1913: 64 View in CoL ( Centrorhynchus ). Type locality: Brazil, Rio Grande do Sul, Santa Victoria do Palmar. LT H (FIOC)
New records: BuenosAires: San Nicolás, 33°19′56.49″S, 60°12′16.64″W GoogleMaps ; Zárate , 34°05′02.49″S, 59°01′6.88″W GoogleMaps ; Villa Elisa , 34°51′34.84″S, 58°05′11.19″W GoogleMaps ; La Plata, 34°55′13.78″S, 57°57′12.84″W GoogleMaps ; Berisso, Los Talas, bathing resort Bagliardi , 34°52′25.93″S, 57°48′39.46″W GoogleMaps ; Dolores , 36°18′11.56″S, 57°38′47.43″W GoogleMaps ; General Madariaga, Pinamar , 37°05′55.94″S, 56°58′05.99″W GoogleMaps ; Tandil , 37°20′13.00″S, 59°07′58″W GoogleMaps .
irwini Spinelli & Wirth, 1984b: 180 View in CoL (female). Type locality: Chile, Coquimbo Prov., Fray Jorge Pachingo. HT F (CASC). Distr.: Chile, Argentina (Neuquén: PN Lanín, Lago Rucachoroy, 39°13′59.07″S, 71°10′38.66″W; Lago Huechulafquen, 39°45′00.70″S, 71°27′21.48″W. Río Negro: El Bolsón, 41°57′53.72″S, 71°32′07.22″W). Refs.: Spinelli & Ronderos, 1994: 60; Spinelli & Ronderos, 2001a: 514.
New records: Neuquén : PN Nahuel Huapi, Península Quetrihué, Laguna Patagua, 40°51′05.7′′S, 71°37′18.3′′W, 817 m GoogleMaps ; PN Nahuel Huapi, Isla Victoria , unnamed stream, 40º53′26.3′′S, 71º33′14.1′′W, 780 m GoogleMaps ; PN Nahuel Huapi, Villa La Angostura , 40°45′14.51″S, 71°39′04.31″W GoogleMaps . Río Negro : Laguna Los Juncos, 41º03′33.8′′S, 71º00′10′′W, 908 m GoogleMaps ; PN Nahuel Huapi, Bariloche, Av. Bustillo km 4.7, 41°07′26.50″S, 71°21′44.32″W; PN Nahuel Huapi, Lago Gutiérrez, 41°10′49.26′′S, 71°23′15.08′′W GoogleMaps ; PN Nahuel Huapi, Arroyo Vertiente , 41°36′05.8′′S, 71°36′24.8′′W, 468 m. GoogleMaps
patagoniensis Ronderos & Spinelli, 1997: 34 (male, female). Type locality: Argentina, Chubut, Lago Fontana, 44°52′46.69″S, 71°32′45.32″W. HT F (MLPA). Distr.: Chile, Argentina (Neuquén: San Martín de los Andes, 40°09′39.65″S, 71°21′13.15″W; Río Limay, ″anfiteatro″, 41°00′49.85″S, 71°06′48.99″W. Río Negro: PN Nahuel Huapi, Cerro Challhuaco, 41°15′30′′S, 71°17′06′′W, 1326 m; Meseta de Somuncura, Estancia La Rinconada, springs Arroyo Comi-Có, 41°08′35.1′′S, 67°27′34.6′′W, 1000 m. Chubut: type locality; Arroyo Apeleg, 44°55′00.73″S, 70°09′01.74″W;northern shore Lago La Plata, 44°52′S, 71°39′W;Meseta de la Sierra Cuadrada,Estancia Don Eduardo, 44º39′25.9′′S, 68º00′09.9′′W. Santa Cruz: Lago Buenos Aires, Paso Pallavicini, 46°16′38.33″S, 71°43′37.30″W, 600 m; Río Pinturas, 47º09′21.8′′S, 70º39′31′′W, 570 m. Tierra del Fuego: Lago Yehuin, 54º21′39.6′′S, 67º46′44.4′′ W, 45 m). Refs.: Ronderos & Spinelli, 1997: 34; Ronderos & Spinelli, 2002: 94; Muzón et al., 2010: 113; Spinelli et al., 2010: 130.
New records: Neuquén: Lago Aluminé , 38°53′44.52″S, 71°05′7.15″W GoogleMaps ; PN Nahuel Huapi, Isla Victoria, unnamed stream, 40º53′26.3′′S, 71º33′14.1′′W, 780 m. GoogleMaps Río Negro : PN Nahuel Huapi, Cerro Challhuaco, 41°15′30′′S, 71°17′06′′W, 1326 m GoogleMaps . Santa Cruz: Meseta del Lago Buenos Aires, Estancia La Frontera, 46°46′10.41″S, 71°45′26.96″W GoogleMaps ; PN Perito Moreno, Lago Burmeister, 47°57′13.55″S, 72°07′12.58″W GoogleMaps .
Subgenus DRYMODESMYIA Vargas
DRYMODESMYIA Vargas, 1960: 40 (as subgenus of Culicoides View in CoL ). Type species: Culicoides copiosus Root & Hoffman View in CoL , by original designation.
chacoensis Spinelli & Wirth, 1984b: 174 View in CoL (male, female). Type locality: Argentina, Salta, Pocitos (Salvador Mazza), 22°03′19.64″S, 63°41′18.90″W.HT F(MLPA).Distr.: Bolivia, Central Chile, Paraguay, Argentina (Salta: type locality; Río Juramento at NRt. 9, 25°07′52.45″S, 65°00′58.95″W. Córdoba: La Serranita, 31°44′04.07″S, 64°27′38.15″W. Misiones: Departamento Cainguás, Municipio Campo Grande, 27°12′28′′S, 54°58′48′′W. Corrientes: Destacamento de Prefectura Paniagua, 10 km S Monte Caseros, 30°20′23.42″S, 57°39′21.48″W. Río Negro: Chipauquil, spring Arroyo Valcheta, 40°58′25′′S, 66°39′14.7′′W, 630 m; Meseta de Somuncura, Estancia El Rincón, 40°59′24.1′′S, 66°40′35.7′′W, 620 m. Chubut: Meseta de la Sierra Cuadrada, Estancia Don Eduardo, 44º39′25.9′′S, 68º00′09.9′′W), Uruguay. Refs.: Spinelli & Wirth, 1984b: 174; Spinelli & Ronderos, 1991: 89; Ronderos & Spinelli, 2002: 93; Muzón et al., 2005: 62; Ayala & Marino, 2018: 36.
New records: Catamarca: Departamento Londres, El Shincal, 27°40′59.99″S, 67°10′00.02″W GoogleMaps . Santiago del Estero: Añatuya, 28°27′38.08″S, 62°50′03.71″W GoogleMaps . Córdoba: 4 km NW Cura Brochero , 31°38′16.68″S, 65°02′10.15″W GoogleMaps .
saltaensis Spinelli & Wirth, 1984b: 183 View in CoL (female). HT F (MLPA). Type locality: Argentina, Salta Cafayate , 26°03′54.14′′S, 65°58′21.99″W. Distr.: Argentina (Salta: type locality; 7 km W Cafayate, 26°05′44.37″S, 66°00′46.91′′W. La Rioja: Pozo de Piedra, 4 mi E Chepes, 31°20′03.81″S, 66°31′55.51″W. Tucumán: Alpachiri, 27°20′S, 65°46′W; Santa Ana, 27°28′S, 65°39′W; Villa Batiruana, 27°38′S, 65°44′W. Mendoza: Mendoza city, Zoo, 32°53′03.75″S, 68°53′20.74″W). Refs.: Spinelli & Wirth, 1984b: 183; Ronderos & Spinelli, 1990: 85; Ronderos & Spinelli, 2002: 94; Veggiani Aybar et al., 2015: 634.
New record: La Rioja: Chilecito , 29°10′27.22″S, 67°28′45.61″W GoogleMaps .
Subgenus HAEMATOMYIDIUM Goeldi
HAEMATOMYIDIUM Goeldi,1905:137 . Type species: Haematomyidium paraense Goeldi ,by original designation.
REFERENCES: Vitale et al., 1981 (key to females of debilipalpis View in CoL group); Wirth & Felippe-Bauer, 1989 (key to paraensis View in CoL group).
austroparaensis Spinelli View in CoL , in Spinelli et al., 2005a: 141 (male, female). Type locality: Argentina, Corrientes, Santa Tecla, 27°36′57.67″S, 56°23′14.66″W. HT M (MLPA). Distr.: Argentina (Misiones: Puerto Iguazú, 25°40′S, 54°33′W. Corrientes: type locality). Refs.: Spinelli et al., 2005a: 141; Veggiani Aybar et al., 2012b: 798.
crucifer Clastrier, 1968: 85 View in CoL (male, female). Type locality: French Guiana, Cayenne. HT M (MNHN). Distr.: Trinidad, Guyana, French Guiana, Brazil (Amazonas, Pará), Argentina (Misiones: Posadas, club San Francisco, 27°23′31.20″S, 55°55′59.37″W). Refs.: Spinelli et al., 2021: 406 View Cited Treatment .
debilipalpis Lutz, 1913: 60 View in CoL (female). Type locality: Brazil, S„o Paulo, Sierra da Bocaina. Type material (?). Distr.: Widespread from USA (Maryland, Kentucky, Nebraska south to Lousiana and Florida), Guatemala and Belize to Argentina ( Salta: Aguas Blancas , 22°34′S, 64°22′W; El Oculto, 23°06′S, 64°30′W; San Ramón de la Nueva Orán, 23°08′S, 64°20′W. Tucumán: Departamento Famaillá, 27°03′51.18″S, 65°23′30.10″W; La Montañita, 26°57′34.38″S, 65°23′20.84″W; Sargento Moya, 27°11′S, 65°38′W, 458 m; La Florida, 27°13′S, 65°37′W, 452 m; Capitán Cáceres, 27°11′S, 65°36′W, 414 m. Formosa: Estanislao del Campo, 25°03′12.50″S, 60°05′38.10″W; El Coatí, 25°43′37.96″S, 59°37′49.22″W; Los Matacos, 25°29′53.59″S, 59°26′26.16″W; Palo Santo, 25°33′48.14″S, 59°20′04.22″W; Formosa city, 26°11′24.06″S, 58°09′21.77″W. Chaco: Ciervo Petiso, 26°34′51.96″S, 59°37′53.94″W; Charadai, 27°39′16.64″S, 59°51′44.48″W; Colonia Elisa, 26°55′54.34″S, 59°31′05.34″W; Colonias Unidas, 26°41′53.95″S, 59°37′48.32″W; Laguna Limpia, 26°29′43.50″S, 59°40′52.36″W; Las Palmas, 27°03′02.58″S, 58°40′39.76″W; Quitilipi, 26°52′19.21″S, 60°13′16.21″W; Río de Oro, 26°54′45.57″S, 58°46′18.86″W. Misiones: Cataratas del Iguazú, 25°41′4.29″S, 54°26′32.29″W; Barra del Yacuí, 25°50′00′′S, 54°09′00′′W; Río Paranaí, 26°40′33.82″S, 54°49′13.44″W; Corpus, 27°06′22.43′′S, 55°31′24.71″W; Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. Corrientes: Ituzaingó, 27°35′22.76″S, 56°41′25.22″W. Entre Ríos: Ibicuy, 33°44′23.51″S, 59°10′06.53″W. Buenos Aires: Isla Martín García, garbage place, 34°10′56.94″S, 58°15′27.91″W; Ensenada, Reserva Natural Integral Punta Lara, marginal forest, 34º47′21.2′′S, 57º59′56.8′′W; Berisso, Los Talas, 34°52′16.47′′S, 57°51′08.76′′W). Refs.: Forattini, 1957: 383; Cavalieri & Chiossone, 1966b: 148; Spinelli et al., 1989: 735 (as lahillei ); Spinelli & Ronderos, 1997: 304; Ronderos et al., 2003b: 22; Veggiani Aybar et al., 2010: 97; Ronderos et al., 2010: 43 View Cited Treatment ; Veggiani Aybar et al., 2011: 1021; Ronderos et al., 2011: 1188; Veggiani Aybar et al., 2012a: 4; Cazorla et al., 2018: 6.
ichesi Ronderos & Spinelli, 1995b: 77 View in CoL . Type locality: Argentina, Misiones GoogleMaps , Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. HT F (MLPA).
Other localities for C. ichesi View in CoL from Argentina cited by Ronderos & Spinelli (1995): Jujuy: PN Calilegua, 23°40′17.58″S, 64°52′00.59″W GoogleMaps . Misiones: Montecarlo , 26°34′10.62″S, 54°47′50.59″W GoogleMaps ; Salto Encantado , 27°05′06″S, 54°49′58″W GoogleMaps ; Posadas, Arroyo Mártires , 27°24′26.66″S, 55°57′29.29″W GoogleMaps . Corrientes: Bella Vista View in CoL , 28°30′39.91″S, 59°03′04.46″W GoogleMaps ; Santa Tecla, 27°36′57.67″S, 56°23′14.66″W GoogleMaps ; Ituzaingó , 27°35′22.76″S, 56°41′25.22″W GoogleMaps ; Monte Caseros , 30°15′23.77″S, 57°37′21.10″W GoogleMaps ; Mocoretá , 30°37′06.63″S, 57°58′31.17″W GoogleMaps . Entre Ríos: Gualeguaychú , 33°00′26.40″S, 58°30′02.69″W GoogleMaps . Ciudad Autónoma de Buenos Aires, 34°32′18.65″S, 58°27′03.69″W GoogleMaps . Buenos Aires: Gonnet , 34°53′20.26″S, 58°01′17.15″W GoogleMaps ; General Belgrano , 35°45′56.68″S, 58°29′49.90″W GoogleMaps .
New record: Buenos Aires: San Nicolás , 33°19′56.49″S, 60°12′16.64″W. dellapei Spinelli, Ronderos & Díaz, 2021: 404 (female). Type locality: Argentina GoogleMaps , Buenos Aires, Tandil , 37°20′13.00″S, 59°07′58″W. HT F ( MLPA). Distr.: Argentina (Buenos Aires: type locality) GoogleMaps . Refs.: Spinelli et al., 2021: 404.
dureti Ronderos & Spinelli, 1995a: 59 View in CoL (female). Type locality: Paraguay, Encarnación, Quiteria. HT F (MLPA). Distr.: Argentina (Misiones: Puerto Iguazú, 25°40′S, 54°33′W; Candelaria, 27°27′33.49″S, 55°45′28.76″W; Posadas, 27°24′26.66″S, 55°57′29.29″W), Paraguay. Refs.: Ronderos & Spinelli, 1995a: 59; Spinelli & Ronderos, 2007: 77; Veggiani Aybar et al., 2012b: 799.
New records: Misiones: Corpus , 27°06′22.43′′S, 55°31′24.71″W GoogleMaps ; San Ignacio, 27°15′37.88″S, 55°31′52.96″W GoogleMaps ; Arroyo Anyico at km 813 NRt. 14, aprox. 30 km N San José, 27°39′52.76″S, 55°36′31.35″W GoogleMaps .
flinti Wirth, 1982b: 251 View in CoL (female). Type locality: Argentina, Entre Ríos, Arroyo Perucho Verna at NRt. 14, 32°09′59.63′′S, 58°14′11.74″W. HT F (USNM). Distr.: Paraguay, sotheastern Brazil (Santa Catarina), Argentina (Misiones: Campo Viera, 27°21′57.62″S, 55°02′51.86″W; Corpus, 27°06′22.43′′S, 55°31′24.71″W; Candelaria, 27°27′33.49″S, 55°45′28.76″W; Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W; Colonia Liebig, Arroyo Chimiray, 27°55′05.87″S, 55°48′51.47″W; Mocoretá, 30°37′06.63″S, 57°58′31.17″W. Entre Ríos: type locality; Chajarí, 30°43′42.11″S, 57°52′39.32″W; Salto Grande, Puerto Luis, 31°23′36.01″S, 57°59′41.09″W. Buenos Aires: Berisso, Los Talas, 34°52′16.47′′S, 57°51′08.76′′W), Uruguay. Refs.: Wirth, 1982b: 251; Spinelli & Wirth, 1984b: 176; Spinelli & Ronderos, 1991: 91; Ronderos et al., 2003b: 22.
New records: Misiones: Eldorado , 26°23′55.73″S, 54°41′36.05″W GoogleMaps ; Aristóbulo del Valle, Arroyo Cuña Pirú , 27°05′16.30″S, 54°57′08.64″W GoogleMaps ; Arroyo Anyico at km 813 NRt. 14, aprox. 30 km N San José, 27°39′52.76″S, 55°36′31.35″W GoogleMaps ; Posadas, Refugio Don Lorenzo , 27°25′10.74″S, 55°52′38.88″W GoogleMaps . Corrientes: Arroyo Pay Ubre , 29º01′41.2′′S, 58º10′26.6′′W, 66 m GoogleMaps ; Entre Ríos: Gualeguaychú , 33°00′26.40″S, 58°30′02.69″W GoogleMaps . Buenos Aires: Estación Otamendi, INTA Delta , 34°10′30.44″S, 58°51′39.75″W GoogleMaps .
germanus Macfie, 1940b: 27 View in CoL (female). Type locality: Guyana. New River. HT F (BMNH). Distr.: Costa Rica, Colombia, Guyana, Argentina ( Formosa: Misión Laishi , 26°14′22.62″S, 58°37′36.61″). Refs.: Forattini, 1957: 381; Cavalieri & Chiossone, 1966b: 148.
ginesi Ortíz, 1951: 586 View in CoL (female). Type locality: Venezuela, Yaracuy, San Felipe. HT F (INHR). Distr.: El Salvador to Panama, Colombia, Venezuela, Trinidad, Brazil (Pará), Argentina (Misiones: Arroyo Anyico at km 813 NRt. 14, aprox. 30 km N San José, 27°39′52.76″S, 55°36′31.35″W). Refs.: Spinelli et al., 2005a: 149.
New record: Misiones : San Pedro, Reserva Biósfera Yaboti, 26°53′36.5″S, 53°52′42.5″W GoogleMaps .
hoffmani Fox, 1946: 251 View in CoL (female). Type locality: Trinidad, Camuto Village. HT F (UPRM). Distr.: Belize, Costa Rica, Cayman Islands, Jamaica to Trinidad, Argentina (Córdoba: 4 km NW Cura Brochero, 31°38′16.68″S, 65°02′10.15″W). Refs.: Spinelli et al., 2021: 406 View Cited Treatment .
horticola Lutz, 1913: 61 View in CoL (female). Type locality: Brazil, S„o Paulo, Tatuí. ST F (FIOC). Distr.: Perú, Paraguay, southeastern Brazil (Espirito Santo, Rio de Janeiro, S„o Paulo), Argentina (Misiones: Corpus, 27°06′22.43′′S, 55°31′24.71″W; Posadas, 9 de Julio, 27°25′27.46″S, 55°54′36.40″W; Posadas, Arroyo Mártires, 27°24′26.66″S, 55°57′29.29″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W. Buenos Aires: Ensenada, Reserva Natural Integral Punta Lara, marginal forest, 34º47′21.2′′S, 57º59′56.8′′W). Refs.: Spinelli et al., 1989: 735; Ronderos & Spinelli, 1998: 81; Ronderos et al., 2003b: 22; Felippe-Bauer & Spinelli, 2009: 28.
bachmanni Spinelli View in CoL , in Spinelli et al., 2005a: 146. Type locality: Argentina, Misiones GoogleMaps , Corpus, 27°06′22.43′′S, 55°31′24.71″W. HT F (MLPA).
lahillei (Iches), 1906: 264 ( Ceratopogon View in CoL , female). Type locality: Argentina, Chaco Austral. Type material (apparently lost). Distr.: Perú, southeastern Brazil (S„o Paulo), Paraguay, Argentina (Jujuy: 50 km W Fraile Pintado, 23°54′S, 65°02′W, 900 m. Salta: Aguas Blancas , 22°34′S, 64°22′W; El Oculto, 23°06′S, 64°30′W; San Ramón de la Nueva Orán, 23°08′S, 64°20′W. Tucumán: Sargento Moya, 27°11′S, 65°38′W, 458 m; La Florida, 27°13′S, 65°37′W, 452 m; Capitán Cáceres, 27°11′S, 65°36′W, 414 m; Chaco: Pampa del Indio, 26°08′S, 59º58′W; PN El Chaco, 26º48′25″S, 56º26′36.5″W, 76 m. Misiones: Iguazú Cué, 25°40′56.18″S, 54°34′36.11″W; Corpus, 27°06′22.43′′S, 55°31′24.71″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W; Destacamento de Prefectura Paniagua, 10 km S Monte Caseros, 30°20′23.42″S, 57°39′21.48″W; Mocoretá, 30°37′06.63″S, 57°58′31.17″W. Entre Ríos: Gualeguaychú, 33°00′26.40″S, 58°30′02.69″W). Refs.: Iches, 1906: 264; Spinelli & Wirth, 1986: 62; Shelley & Coscarón, 2001: 454; Spinelli & Ronderos, 1991: 91; Ronderos et al., 2003b: 22; Veggiani Aybar et al., 2010: 97; Veggiani Aybar et al., 2011: 1021; Veggiani Aybar et al., 2012a: 4; Refs.: Veggiani Aybar et al., 2012b: 799.
New records: Jujuy: PN Calilegua, 23°40′17.58″S, 64°52′00.59″W GoogleMaps . Formosa: Las Lomitas, 24°42′14.41″S, 60°35′11.08″W. Chaco: Laguna Limpia, 26°29′43.50″S, 59°40′52.36″W GoogleMaps ; Colonia Elisa, 26°55′54.34″S, 59°31′5.34″W GoogleMaps ; Colonias Unidas, 26°41′53.95″S, 59°37′48.32″W. Misiones: Puerto Iguazú, 25°40′S, 54°33′W GoogleMaps ; PN Iguazú, Garganta del Diablo , 25°41′33.28″S, 54°26′21.41″W GoogleMaps ; Montecarlo, 26°34′10.62″S, 54°47′50.59″W; Paranay, 26°40′33.82″S, 54°49′13.44″W; Tobuna, 26°28′01.24″S, 53°53′24.77″W; Posadas, Arroyo Mártires. Santa Fé: Reserva de la Defensa Campo Garabato, El Ombucito, charcas, 28°53′31.15″S, 60°12′36.91″W GoogleMaps . Corrientes: Garapé, 27°36′05.30″S, 56°14′44.44″W GoogleMaps .
pampaensis Spinelli & Wirth, 1984b: 182 View in CoL (female). Type locality: Argentina, La Pampa View in CoL , Colonia 25 de Mayo, 37°46′29.8′′S, 67°42′48.5′′W. HT F (MLPA). Distr.: Argentina (Jujuy: Libertador General San Martín, 23°48′S, 64°48′W. Salta: Cafayate, 7 km W Cafayate, Río Colorado, 26°05′44.37″S, 66°00′46.91W; San José de Metán, 25°30′S, 64°58′W. Catamarca: Departamento Londres, El Shincal, 27°40′59.99′′S, 67°10′00.02′′W. Chubut: Meseta de Somuncura, Arroyo Telsen, 42°28′35′′S, 66°52′18′′W, 276 m). Refs.: Spinelli & Wirth, 1984b: 182; Ronderos & Spinelli, 2002: 94; Spinelli & Ronderos, 2005: 43; Muzón et al., 2010: 113; Veggiani Aybar et al., 2015: 634.
paraensis (Goeldi), 1905: 137 View in CoL ( Haematomyidium ; female). Type locality: Brazil, Pará, Belém. NT F (FIOC). Distr.: USA (Colorado, Nebraska, Pennsylvania, Wisconsin to Louisiana and Florida) to Argentina (Jujuy: 50 km W Fraile Pintado, 23°54′S, 65°02′W, 900 m. Salta: Río Caraparí, 8 km S Pocitos (Salvador Mazza), 22°04′03.19′′S, 63°40′20.26′′W, 757 m; Aguas Blancas, 22°34′S, 64°22′W; El Oculto, 23°06′S, 64°30′W; San Ramón de la Nueva Orán, 23°08′S, 64°20′W; Cornejo, 24°48′21.85″S, 65°25′24.12″W; 44 km E Aguaray, 22°13′55.25″S, 63°19′47.06″W. Tucumán: Famaillá, 27°03′51.18″S, 65°23′30.10″W; Quebrada Del Loro, 26°37′02.76″S, 65°10′33.90″W; Sargento Moya, 27°11′S, 65°38′W, 458 m; La Florida, 27°13′S, 65°37′W, 452 m; Capitán Cáceres, 27°11′S, 65°36′W, 414 m. Mendoza: Villa Atuel, 34°49′51.31″S, 67°55′26.56″W. Chaco: Las Palmas, 27°03′02.58″S, 58°40′39.76″W. Misiones: Barra del Yacuí, 25°50′00′′S, 54°09′00′′W; Iguazú Cué, 25°40′56.18″S, 54°34′36.11″W; PN Iguazú, 25°39′S, 54°18′W; Corpus, 27°06′22.43′′S, 55°31′24.71″W; Candelaria, 27°27′33.49″S, 55°45′28.76″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W; Ituzaingó, 27°35′22.76″S, 56°41′25.22″W; Destacamento de Prefectura Paniagua, 10 km S Monte Caseros, 30°20′23.42″S, 57°39′21.48″W; Mocoretá, 30°37′06.63″S, 57°58′31.17″W). Refs.: Forattini, 1957: 426; Cavalieri & Chiossone, 1966b: 149; Wirth & Felippe Bauer: 1989: 553; Spinelli & Ronderos, 1991: 90; Shelley & Coscarón, 2001: 453; Ronderos & Spinelli, 2002: 94; Ronderos et al., 2003b: 22; Campos et al., 2011: 115; Veggiani Aybar et al., 2010: 97; Veggiani Aybar et al., 2011: 1021; Veggiani Aybar et at., 2012a: 4.
undecimpunctatus Kieffer, 1917b: 307 . Type locality: Argentina, Tucumán, San Pablo GoogleMaps , 26°52′19.75″S, 65°18′45.39″W. Type material (HNHM, lost).
New records: Jujuy: PN Calilegua, 23°40′17.58″S, 64°52′00.59″W GoogleMaps . Santiago del Estero: Añatuya, 28°27′38.08″S, 62°50′03.71″W GoogleMaps . Córdoba: Río San Antonio, paraje Las Jarillas , 31º32′02.8′′S, 64º33′01.7′′W, 820 m. GoogleMaps Misiones: Puerto Iguazú, 25°40′S, 54°33′W GoogleMaps ; Arroyo Anyico at km 813 NRt. 14, aprox. 30 km N San José, 27°39′52.76″S, 55°36′31.35″W GoogleMaps ; Posadas, 27°24′26.66″S, 55°57′29.29″W. Corrientes: Garapé, 27°36′05.30″S, 56°14′44.44″W GoogleMaps .
Subgenus HOFFMANIA Fox
HOFFMANIA Fox, 1948: 21 (as subgenus of Culicoides View in CoL ). Type species: Culicoides inamollae Fox & Hoffman View in CoL (= Culicoides insignis Lutz View in CoL ), by original designation.
REFERENCES: Spinelli et al., 1993 (key to species groups; key to males and females of guttatus View in CoL group); Wirth & Blanton, 1968 (key to males and females in hylas View in CoL group); Felippe-Bauer et al., 2009 (synopsis of hylas View in CoL group).
biestroi Spinelli & Ronderos, 1991: 86 (male, female) Type locality: Argentina, Corrientes, Destacamento de Prefectura Paniagua, 10 km S Monte Caseros, 30°20′23.42″S, 57°39′21.48″W. HT H (MLPA). Distr.: Argentina (Misiones: Puerto Iguazú, 25°40′S, 54°33′W. Corrientes: type locality. Buenos Aires: Magdalena, Arroyo Zapata, 35°08′29.55′′S, 57°26′32.17′′W, 32 m), Uruguay. Refs.: Spinelli & Ronderos, 1991: 86; Spinelli et al., 1993: 20; Veggiani Aybar et al., 2012b: 798.
New record: Misiones: Posadas , 27°24′26.66″S, 55°57′29.29″W GoogleMaps .
brasilianum Forattini, 1956: 81 (female). Type locality: Brazil: S„o Paulo, S„o Vicente. HT F (DEFS). Distr.: Brazil (Amazonas, Pará, Ceará, Pernambuco, Rio de Janeiro, S„o Paulo, Santa Catarina), Argentina (Misiones: Puerto Iguazú, 25°40′S, 54°33′W; Posadas, 27°24′26.66″S, 55°57′29.29″W). Refs.: Ronderos & Spinelli, 1990: 82; Veggiani Aybar et al., 2012b: 799.
calchaqui Spinelli & Veggiani Aybar View in CoL , in Spinelli et al., 2013c: 586 (female). Type locality: Argentina, Tucumán, Lules, Potrero de Las Tablas, 26°54′1.00″S, 65°26′59.99″W. HT F (IFML). Distr.: Argentina (Tucumán: type locality). Refs.: Spinelli et al., 2013c: 586.
charruus Spinelli & Martinez, 1992: 176 (as charrua View in CoL ; pupa, male, female). Type locality: Uruguay, Departamento San José, Playa Pascual, in Eryngium serra View in CoL . HTF (MLPA). Distr.: Paraguay, Argentina (Misiones: Corpus, 27°06′22.43″S, 55°31′24.71″W; Posadas, Arroyo Mártires, 27°24′26.66″S, 55°57′29.29″W. Buenos Aires: Mar de Ajó, 36°43′27.91″S, 56°42′23.86″W), Uruguay. Refs.: Ronderos & Spinelli, 1998: 81; Ronderos et al., 2003b: 22; Ronderos et al., 2008: 116.
New record: Misiones: Arroyo Anyico at km 813 NRt. 14, aprox. 30 km N San José, 27°39′52.76″S, 55°36′31.35″W GoogleMaps .
fernandoi Tavares & Souza, 1979: 611 View in CoL (male, female). Type locality: Brazil: Rio de Janeiro, Sumidouro. HT M (MNRJ). Distr.: Colombia, Brazil (Espirito Santo, Bahia, Maranh„o, Minas Gerais, Rio de Janeiro, Santa Catarina), Argentina (Chaco: PN El Chaco, 26º48′25″S, 56º26′36.5″W, 76 m. Misiones: Puerto Iguazú, 25°40′S, 54°33′W; Corpus, 27°06′22.43″S, 55°31′24.71″W; Candelaria, 27°27′33.49″S, 55°45′28.76″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W; Colonia Liebig, Arroyo Chimiray, 27°55′05.87″S, 55°48′51.47″W; Cuay Chico, 28°42′56.40″S, 56°24′03.00″W; Destacamento de Prefectura Paniagua, 10 km S Monte Caseros, 30°20′23.42″S, 57°39′21.48″W), Uruguay. Refs.: Spinelli & Wirth, 1986: 60; Spinelli & Ronderos, 1991: 89; Ronderos et al., 2003b: 22; Veggiani Aybar et al., 2012b: 798.
New records: Corrientes: Esteros del Iberá , Galarza , Arroyo Yacaré , 28°03′37.21″S, 56°42′04.00″W GoogleMaps ; Esteros del Iberá, Galarza , 28°05′40.29′′S, 56°41′11.25″W GoogleMaps .
ferreyrai Ronderos & Spinelli, 1995a: 61 View in CoL (female). Type locality: Argentina, Misiones, Posadas, Arroyo Mártires, 27°24′26.66″S, 55°57′29.29″W. HT F (MLPA). Distr.: Argentina (Misiones: type locality; Corpus, 27°06′22.43″S, 55°31′24.71″W; Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. Refs.: Ronderos et al., 2003b: 22.
New record: Misiones: Arroyo Garupá , 27°29′28.39″S, 55°50′24.23″W GoogleMaps .
foxi Ortíz, 1950: 461 View in CoL (male). Type locality: Puerto Rico, Campo Tortuguero ( Fox, 1949: 31, as C. guttatus View in CoL ). HT M (UPRM). Distr.: Mexico (Chiapas) to Bolivia, Puerto Rico to Argentina (Misiones: Puerto Iguazú, 25°40′S, 54°33′W. Corrientes: Cuay Chico, 28°42′56.40″S, 56°24′03.00″W; Mocoretá, 30°37′06.63″S, 57°58′31.17″W). Refs.: Forattini, 1957: 205; Cavalieri & Chiossone, 1966b: 148; Spinelli & Ronderos, 1991: 88; Veggiani Aybar et al., 2012b: 798.
New records: Tucumán: La Florida, 27°13′S, 65°37′W, 452m. GoogleMaps Misiones: Paranay , 26°40′33.82″S, 54°49′13.44″W GoogleMaps .
guttatus (Coquillett), 1904: 35 View in CoL ( Ceratopogon View in CoL ; female). Type locality: Brazil, S„o Paulo, Serra da Cantareira. LT F (USNM) View in CoL . Distr.: Southeastern Brazil (Minas Gerais, Rio de Janeiro, S„o Paulo, Santa Catarina), Paraguay, Argentina (Jujuy: Yuto, 23°38′S, 64°28′W. Salta: San Ramón de la Nueva Orán, 23°08′S, 64°20′W. Tucumán: Yánima, 27°39′S, 65°40′W. Misiones: Corpus, 27°06′22.43′′S, 55°31′24.71″W; Departamento Cainguás, Municipio Campo Grande, 27°12′28′′S, 54°58′48′′W; Candelaria; Posadas, Arroyo Mártires, 27°24′26.66″S, 55°57′29.29″W; Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. Buenos Aires: Berisso, Los Talas, 34°52′16.47′′S, 57°51′08.76′′W). Refs.: Ronderos & Spinelli, 1998: 81; Ronderos & Spinelli, 2002: 93; Ronderos et al., 2003b: 22; Veggiani Aybar et al., 2012b: 798; Veggiani Aybar et al., 2015: 634.
New record: Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W GoogleMaps .
ignacioi Forattini, 1957: 215 (male, female). Type locality: Brazil, S„o Paulo, Boracéa, M. Salesópolis HT M (DEFS). Distr.: Brazil (Acre, Amazonas, Pará, Rondônia, Roraima, Maranh „o, Minas Gerais, Rio de Janeiro, S„o Paulo GoogleMaps ), Paraguay, Argentina (Corrientes: Santa Tecla GoogleMaps , 27°36′57.67″S, 56°23′14.66″W). Refs.: Ronderos et al., 2003b: 22.
saintjusti Tavares & Ruiz, 1980: 27 View in CoL . Type locality: Brazil, Rio de Janeiro, Jacarepaguá ( Pau da Fome ). HT M (MNRJ).
New records: Buenos Aires: Berisso, Los Talas , 34°52′16.47′′S, 57°51′08.76′′W GoogleMaps ; Berisso, Los Talas, bathing resort Bagliardi , 34°52′25.93″S, 57°48′39.46″W GoogleMaps .
insignis Lutz, 1913: 51 View in CoL (pupa, male, female). Type locality: Brazil, Rio de Janeiro and Bahia. LT M (FIOC). Distr.: USA (Alabama, Georgia, Florida), Mexico (Yucatán, Chiapas), Central America and Caribbean to Argentina (Jujuy: Libertador General San Martín, 23°48′S, 64°48′W; Yuto, 23°38′S, 64°28′W. Salta:Aguas Blancas, 22°34′S, 64°22′W; El Oculto, 23°06′S, 64°30′W; San Ramón de la Nueva Orán, 23°08′S, 64°20′W. Tucumán: Sargento Moya, 27°11′S, 65°08′W, 458 m; La Florida, 27°13′S, 65°37′W, 452m; Capitán Cáceres, 27°11′S, 65°36′W, 414 m. Formosa: PN Río Pilcomayo, 25°04′20.09″S, 58°08′27.55″W. Chaco: General Obligado, 27°24′39.28″S, 59°24′57.71″W; Resistencia, 27°28′55.72″S, 58°57′59.93″W. Misiones: Corpus, 27°06′22.43′′S, 55°31′24.71″W; Candelaria, 27°27′33.49″S, 55°45′28.76″W; Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W; Ituzaingó, 27°35′22.76″S, 56°41′25.22″W; San Roque, 28°34′14.02″S, 58°43′4.65″W; Arroyo Ambrosio, 28º15′16.4′′S, 58º50′33.6′′W, 44 m; Destacamento de Prefectura Paniagua, 10 km S Monte Caseros, 30°20′23.42″S, 57°39′21.48″W; Mocoretá, 30°37′06.63″S, 57°58′31.17″W. Entre Ríos: Federación, Puerto San Rafael, 30°59′42.66″S, 57°54′43.02″W; Santa Ana, 30°54′18.67′′S, 57°56′32.17′′W, 6 m; Arroyo Perucho Verna at NRt. 14, 32°09′59.63″S, 58°14′11.74″W; Gualeguaychú, 33°00′26.40″S, 58°30′02.69″W. Buenos Aires: Isla Martín García, ″cantera tanque″, 34°11′0.60″S, 58°15′25.69″W; Ensenada, Reserva Natural Integral Punta Lara, marginal forest, 34º47′21.2′′S, 57º59′56.8′′W; same data except grassland, 34°48′01.36″S, 58°00′26.26″W; Berisso, Los Talas, 34°52′16.47′′S, 57°51′08.76′′W; Magdalena, Arroyo Zapata, 35°08′29.55′′S, 57°26′32.17′′W, 32 m). Refs.: Forattini, 1957: 223; Cavalieri & Chiossone, 1966b: 148; Spinelli & Balseiro, 1982: 226; Ronderos & Spinelli, 1989: 80; Spinelli et al., 1989: 735; Spinelli & Ronderos, 1991; Ronderos et al., 2003b: 22; Spinelli et al., 1993: 53; Veggiani Aybar et al., 2010: 97; Veggiani Aybar et al., 2011: 1021; Ronderos et al., 2011: 1188; Veggiani Ayber et al., 2012a: 4; Veggiani Aybar et al., 2012b: 798; Marino et al., 2013: 784; Veggiani Aybar et al., 2015: 634; Cazorla et al., 2018: 6.
inamollae Fox & Hoffman, 1944: 110 View in CoL . Type locality: Puerto Rico, Mayaguez. HT F (UPRM).
painteri Fox, 1946: 257 . Honduras, Puerto Castilla. HT F (UPRM).
New records: Jujuy: Posta de Lozano , 24°04′39.17″S, 65°24′36.80″W GoogleMaps . Salta: 44 km E Aguaray , 22°13′55.25″S, 63°19′47.06″W GoogleMaps . Chaco: PN El Chaco, 26º48′25″S, 56º26′36.5″W, 76 m. GoogleMaps Misiones: Campo Viera , 27°21′57.62″S, 55°02′51.86′′W GoogleMaps ; Posadas , Arroyo Mártires , 27°24′26.66″S, 55°57′29.29″W GoogleMaps . Corrientes: Garapé , 27°36′05.30″S, 56°14′44.44″W GoogleMaps ; Villa Olivari , 27°36′10.36″S, 56°53′09.48″W GoogleMaps ; Colonia Liebig , Arroyo Chimiray , 27°55′05.87″S, 55°48′51.47″W GoogleMaps ; Esteros del Iberá , Colonia Pellegrini , 28°32′17.58″S, 57°10′56.86″W GoogleMaps ; Esteros del Iberá, Estancia San Nicolás, casco, 28º07′41.0′′S, 57º26′03.5′′W GoogleMaps ; Esteros del Iberá , Estancia Rincón del Socorro , casa de Biólogos, 28°41′17.0′′S, 57°26′03.2′′W, 80 m. GoogleMaps Entre Ríos: Paraná, 31°44′28.63″S, 60°30′41.57″W GoogleMaps . Buenos Aires: La Plata, 34°55′13.78″S, 57°57′12.84″W GoogleMaps .
lutzi Costa Lima, 1937: 419 (female). Type locality: Brazil, Pará, Abaeté. ST F (FIOC). Distr.: Colombia to French Guiana, Argentina (Misiones: Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W), Brazil (Roraima, Amazonas, Pará, Mato Grosso, Goiás, São Paulo, Rio de Janeiro, Santa Catarina). Refs.: Ronderos & Spinelli, 1998: 82; Ronderos et al., 2003b: 22.
plaumanni Spinelli , in Spinelli et al., 1993: 69 (female). Type locality: Argentina, Chaco, PN El Chaco, 26º48′25″S, 56º26′36.5″W, 76 m. HT F (MLPA). Distr.: Colombia, Peru, Bolivia, Brazil (Amazonas, Pará, Rondônia, Minas Gerais), Argentina (Misiones: Departamento Cainguás, Municipio Campo Grande, 27°12′28′′S, 54°58′48′′W; Posadas, 27°26′11′′S, 55°53′47.4′′W. Chaco: type locality). Refs.: Spinelli et al., 1993: 69; Ayala & Marino, 2018: 37.
pseudoheliconiae Felippe-Bauer View in CoL , in Felippe-Bauer et al., 2008: 260 (female). Type locality: Peru, Departamento San Martín, Provincia San Martín, Distrito de Juan Guerra, El Porvenir. HT F (FIOC). Distr.: Peru, Brazil (Amazonas, Roraima), Argentina (Tucumán: Monteros, Reserva Provincial La Florida, 27°13′S, 65°37′W, 452 m. Misiones: Puerto Iguazú, Arroyo Mbocay, 25°37′51.16″S, 54°33′41.46″W; Puerto Iguazú, 25°40′S, 54°33′W). Refs.: FelippeBauer et al., 2009: 856; Veggiani Aybar et al., 2012b: 799; Spinelli et al., 2013c: 589.
Subgenus MATAEMYIA Vargas
MATAEMYIA Vargas, 1960: 43 (as subgenus of Culicoides View in CoL ). Type species: Culicoides mojingaensis Wirth & Blanton View in CoL , by original designation.
REFERENCE: Wirth & Soria, 1981 (key males and females to some species, as discrepans View in CoL group).
cuiabai Wirth, 1982b: 250 View in CoL (female). Type locality: Brazil, Mato Grosso, Cuiaba. HT F (USNM). Distr.: Brazil (Mato Grosso), Argentina (Corrientes: Ituzaingó, 27°35′22.76″S, 56°41′25.22″W). Refs.: Ronderos & Spinelli, 1998: 81; Ronderos et al., 2003b: 22.
Subgenus PSYCHOPHAENA Philippi
PSYCHOPHAENA Philippi, 1865: 628 . Type species: Psychophaena pictipennis Philippi (= Culicoides venezuelensis Ortíz & Mirsa ), by monotypy.
lacustris Ronderos, 1990a: 116 View in CoL (male, female). Type locality: Argentina, Río Negro, PN Nahuel Huapi, 6 km W southern shore Lago Mascardi, 41°21′01.70″S, 71°35′58.48″W. HT H (MLPA). Distr.: Argentina (Neuquén: PN Lanín, Lago Rucachoroy, 39°13′59.07″S, 71°10′38.66″W. Río Negro: type locality; Meseta de Somuncura, Estancia La Rinconada, springs Arroyo Comi-Có, 41°08′35.1′′S, 67°27′34.6′′W, 1000 m; Meseta de Somuncura, Estancia Jaguar Musso, paraje Chasicó, spring, 41°08′28′′S, 67°35′43.9′′W, 961 m; Meseta de Somuncura, Estancia El Rincón, spring Arroyo Valcheta, 40°59′24.1′′S, 66°40′35.7′′W, 620 m; Meseta de Somuncura, Chipauquil, spring Arroyo Valcheta, 40°58′25′′S, 66°39′14.7′′W, 630 m. Chubut: Meseta de Somuncura, Arroyo Telsen, 42°28′35.2′′S, 66°52′08′′W, 276 m; PRt. 12 at El Tropezón, 42°47′44.2′′S; 70°51′07.7′′W, 781 m). Refs.: Spinelli & Ronderos, 1994: 60; Muzón et al., 2005: 62; Muzón et al., 2010: 113; Anjos-Santos et al., 2014: 584 View Cited Treatment .
New records: Neuquén: PN Nahuel Huapi, Lago Correntoso, 40°43′58.90″S, 71°40′15.97″W GoogleMaps ; PN Nahuel Huapi, Península Quetrihué , Laguna Patagua, 40°51′05.7′′S, 71°37′18.3′′W, 817 m GoogleMaps ; PN Nahuel Huapi, Isla Victoria , unnamed stream, 40º53′26.3′′S, 71º33′14.1′′W, 780 m GoogleMaps ; PN Nahuel Huapi, Villa La Angostura , 40°45′14.51″S, 71°39′04.31″W GoogleMaps . Río Negro : Laguna Los Juncos, 41º03′33.8′′S, 71º00′10′′W, 908 m GoogleMaps ; PN Nahuel Huapi, Río Manso Superior , 41º14′28.4′′S, 71º44′12.6′′W, 837 m. GoogleMaps Chubut: 16 km SE Paso del Sapo , 42°50′33.49″S, 69°18′34.44″W GoogleMaps ; Meseta de la Sierra Cuadrada, Estancia Don Eduardo, 44º39′25.9′′S, 68º00′09.9′′W. Santa Cruz: Río Pinturas, 47º09′21.8′′S, 70º39′31′′W, 570 m GoogleMaps ; Gobernador Gregores, 48°45′20.09″S, 70°14′25.89″W.
venezuelensis Ortíz & Mirsa, 1950: 137 (male, female). Type locality: Venezuela, Caracas, Los Chorros. HT F (INHR). Distr.: Costa Rica to Chile and Argentina (Jujuy: Yuto, 23°38′S, 64°28′W. Salta: Aguas Blancas, 22°34′S, 64°22′W; El Oculto, 23°06′S, 64°30′W; San Ramón de la Nueva Orán, 23°08′S, 64°20′W; Cafayate, 26°03′54.14″S, 65°58′21.99″W. Tucumán: Sargento Moya, 27°11′S, 65°38′W, 458 m; La Florida, 27°13′S, 65°37′W, 452 m; Capitán Cáceres, 27°11′S, 65°36′W, 414 m. Mendoza: San Rafael, 34°39′59.74″S, 68°14′31.71″W. Formosa: PN Río Pilcomayo, 25°04′20.09″S, 58°08′27.55″W. Chaco: PN El Chaco, 26º48′25″S, 56º26′36.5″W, 76 m. Misiones: Corpus, 27°06′22.43′′S, 55°31′24.71″W; Candelaria, 27°27′33.49″S, 55°45′28.76″W; Campo Viera, 27°21′57.62″S, 55°02′51.86″W; Posadas, Arroyo Zaimán, 27°25′46.64″S, 55°54′08.56″W. Corrientes: Santa Tecla, 27°36′57.67″S, 56°23′14.66″W; Ituzaingó, 27°35′22.76″S, 56°41′25.22″W; Destacamento de Prefectura Paniagua, 10 km S Monte Caseros, 30°20′23.42″S, 57°39′21.48″W; Mocoretá, 30°37′06.63″S, 57°58′31.17″W. Entre Ríos: Federación, Puerto San Rafael, 30°59′42.66″S, 57°54′43.02″W; Santa Ana, 30°54′18.67′′S, 57°56′32.17′′W, 6 m. Buenos Aires: Isla Martín García, ″arenal chico″, 34°10′49.16″S, 58°15′22.52″W; Ensenada, Reserva Natural Integral Punta Lara, marginal forest, 34º47′21.2′′S, 57º59′56.8′′W; same data except grassland, 34°48′01.36″S, 58°00′26.26″W; Berisso, Los Talas, 34°52′16.47′′S, 57°51′08.76′′W; Sierra de la Ventana, Arroyo Belisario, 38º05′18.7′′S, 61º55′46.9′′W, 376 m; Sierra de la Ventana, Reserva Parque Provincial Ernesto Tornquist, Arroyo de la Clausura, 38º02′46.1′′S, 61º59′10.1′′W; Sierra de la Ventana, Reserva Parque Provincial Ernesto Tornquist, unnamed stream, 38º01′09.4′′S, 62º00′56.6′′W). Refs.: Forattini, 1957: 463; Spinelli & Balseiro, 1982: 226; Cavalieri & Chiossone, 1966b: 149; Ronderos & Spinelli, 1989: 78; Spinelli et al., 1989: 735; Spinelli & Ronderos, 1991: 89; Ronderos et al., 2003b: 12; Spinelli, 2003: 22; Díaz et al., 2005: 376; Veggiani Aybar Aybar et al., 2010: 97; Marino et al., 2011a: 203; Veggiani Aybar et al., 2011: 1021; Ronderos et al., 2011: 1189; Veggiani Aybar et al., 2012a: 4; Veggiani Aybar et al., 2012b: 798; Veggiani Aybar et al., 2015: 636; Cazorla et al., 2018: 7.
pictipennis (Philippi), 1865: 628 ( Psychophaena ) (preoccupied by Culicoides pictipennis (Staeger), 1839 View in CoL ). Type locality: Chile, Santiago, San Cristobal LT H (MUNS).
ortizi Fox, 1952: 366. Type locality: Venezuela, Zulia, La Salina. HT F ( UPRM) .
New records: Jujuy: Posta de Lozano , 24°04′39.17′′S, 65°24′36.80″W GoogleMaps . Salta: Río Juramento at NRt. 9, 25°07′52.45″S, 65°00′58.95″W GoogleMaps . Mendoza: Río Malargue , 35°32′12.22″S, 69°38′42.74″W GoogleMaps . Buenos Aires: Gonnet, Arroyo Rodríguez , 34°53′03.08″S, 58°02′30.64″W GoogleMaps ; General Madariaga, Pinamar , 37°05′55.94″S, 56°58′05.99″W GoogleMaps .
| UPRM |
University of Puerto Rico at Mayagueez, Rhizobium Culture Collection |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Ceratopogon aeschnosuga de Meijere
| Spinelli, Gustavo R., Ronderos, María M., Ayala, Mahia M. & Díaz, Florentina 2023 |
bachmanni
| Spinelli, G. R. & Ronderos, M. M. & Diaz, F. & Marino, P. I. 2005: 146 |
soibelzoni
| Marino, P. I. & Spinelli, G. R. 2001: 14 |
calchaqui
| Spinelli, G. R. & Marino, P. I. 1997: 188 |
riojana
| Muzon, J. & Spinelli, G. R. & Von Ellenrieder, N. & Estevez, A. L. & Marino, P. I. & Perez Goodwyn, P. J. & Angrisano, E. B. & Diaz, F. & Fernandez, L. A. & Mazzucconi, S. & Rossi, G. & Salomon, O. D. 2005: 62 |
| Marino, P. I. & Spinelli, G. R. 2001: 16 |
| Spinelli, G. R. & Marino, P. I. 1997: 188 |
ichesi
| Ronderos, M. M. & Spinelli, G. R. 1995: 77 |
dureti
| Veggiani Aybar, C. A. & Dantur, M. J. & Stein, M. & Oria, G. & Ramirez, P. & Lizarralde de Grosso, M. & Spinelli, G. R. 2012: 799 |
| Ronderos, M. M. & Spinelli, G. R. & Diaz, F. 2007: 77 |
| Ronderos, M. M. & Spinelli, G. R. 1995: 59 |
| Ronderos, M. M. & Spinelli, G. R. 1995: 59 |
ferreyrai
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Ronderos, M. M. & Spinelli, G. R. 1995: 61 |
biestroi
| Veggiani Aybar, C. A. & Dantur, M. J. & Stein, M. & Oria, G. & Ramirez, P. & Lizarralde de Grosso, M. & Spinelli, G. R. 2012: 798 |
| Spinelli, G. R. & Ronderos, M. M. 1991: 86 |
| Spinelli, G. R. & Ronderos, M. M. 1991: 86 |
lacustris
| Anjos-Santos, D. & Funes, A. & Spinelli, G. R. & Ronderos, M. M. 2014: 584 |
| Muzon, J. & Spinelli, G. & Rossi, G. C. & Marino, P. I. & Diaz, F. & Melo, M. C. 2010: 113 |
| Muzon, J. & Spinelli, G. R. & Von Ellenrieder, N. & Estevez, A. L. & Marino, P. I. & Perez Goodwyn, P. J. & Angrisano, E. B. & Diaz, F. & Fernandez, L. A. & Mazzucconi, S. & Rossi, G. & Salomon, O. D. 2005: 62 |
| Ronderos, M. M. & Spinelli, G. R. 1994: 60 |
| Ronderos, M. M. 1990: 116 |
irwini
| Spinelli, G. R. & Ronderos, M. M. 2001: 514 |
| Ronderos, M. M. & Spinelli, G. R. 1994: 60 |
| Spinelli, G. R. & Wirth, W. W. 1984: 180 |
chacoensis
| Ayala, M. M. & Marino, P. I. 2018: 36 |
| Muzon, J. & Spinelli, G. R. & Von Ellenrieder, N. & Estevez, A. L. & Marino, P. I. & Perez Goodwyn, P. J. & Angrisano, E. B. & Diaz, F. & Fernandez, L. A. & Mazzucconi, S. & Rossi, G. & Salomon, O. D. 2005: 62 |
| Ronderos, M. M. & Spinelli, G. R. 2002: 93 |
| Spinelli, G. R. & Ronderos, M. M. 1991: 89 |
| Spinelli, G. R. & Wirth, W. W. 1984: 174 |
| Spinelli, G. R. & Wirth, W. W. 1984: 174 |
saltaensis
| Veggiani Aybar, C. A. & Dantur, M. J. & Claps, G. L. & Lizarralde de Grosso, M. & Spinelli, G. R. 2015: 634 |
| Ronderos, M. M. & Spinelli, G. R. 2002: 94 |
| Ronderos, M. M. & Spinelli, G. R. 1990: 85 |
| Spinelli, G. R. & Wirth, W. W. 1984: 183 |
| Spinelli, G. R. & Wirth, W. W. 1984: 183 |
pampaensis
| Veggiani Aybar, C. A. & Dantur, M. J. & Claps, G. L. & Lizarralde de Grosso, M. & Spinelli, G. R. 2015: 634 |
| Muzon, J. & Spinelli, G. & Rossi, G. C. & Marino, P. I. & Diaz, F. & Melo, M. C. 2010: 113 |
| Ronderos, M. M. & Spinelli, G. R. 2002: 94 |
| Spinelli, G. R. & Wirth, W. W. 1984: 182 |
| Spinelli, G. R. & Wirth, W. W. 1984: 182 |
flinti
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Spinelli, G. R. & Ronderos, M. M. 1991: 91 |
| Spinelli, G. R. & Wirth, W. W. 1984: 176 |
| Wirth, W. W. 1982: 251 |
| Wirth, W. W. 1982: 251 |
cuiabai
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Ronderos, M. M. & Spinelli, G. R. 1998: 81 |
| Wirth, W. W. 1982: 250 |
saintjusti
| Tavares, O. & Ruiz, R. A. 1980: 27 |
fernandoi
| Veggiani Aybar, C. A. & Dantur, M. J. & Stein, M. & Oria, G. & Ramirez, P. & Lizarralde de Grosso, M. & Spinelli, G. R. 2012: 798 |
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Spinelli, G. R. & Ronderos, M. M. 1991: 89 |
| Spinelli, G. R. & Wirth, W. W. 1986: 60 |
| Tavares, O. & Souza, M. A. de 1979: 611 |
SALIOHELEA
| Wirth, W. W. & Ratanaworabhan, N. C. 1978: 494 |
leei
| Cazorla, C. G. & Marino, P. I. & Diaz, F. & Campos, R. E. 2018: 6 |
| Marino, P. I. & Cazorla, C. G. & Diaz, F 2011: 202 |
| Wirth, W. W. & Ratanaworabhan, N. C. 1978: 498 |
goniognatha
| Wirth, W. W. & Messersmith, D. H. 1971: 20 |
leptognatha
| Wirth, W. W. & Messersmith, D. H. 1971: 21 |
paraensis (Goeldi), 1905: 137
| Campos, R. E. & Spinelli, G. R. & Mogi, M. 2011: 115 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2011: 1021 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2010: 97 |
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Ronderos, M. M. & Spinelli, G. R. 2002: 94 |
| Shelley, A. J. & Coscaron, S. 2001: 453 |
| Spinelli, G. R. & Ronderos, M. M. 1991: 90 |
| Cavalieri, F. & Chiossone, I. F. 1966: 149 |
ANILOMYIA
| Vargas, L. 1960: 37 |
DRYMODESMYIA
| Vargas, L. 1960: 40 |
MATAEMYIA
| Vargas, L. 1960: 43 |
WARMKEA
| Saunders, L. G. 1957: 671 |
aeria
| Wirth, W. W. & Soria, S. J. 1980: 143 |
| Saunders, L. G. 1957: 675 |
tuberculata
| Spinelli, G. R. & Balseiro, E. G. 1982: 226 |
| Saunders, L. G. 1957: 677 |
nigrigenus
| Ronderos, M. M. & Spinelli, G. R. 1990: 85 |
| Wirth, W. W. & Blanton, F. S. 1956: 222 |
AVARITIA
| Fox, I. 1955: 218 |
sanctaeclarae
| Muzon, J. & Spinelli, G. & Rossi, G. C. & Marino, P. I. & Diaz, F. & Melo, M. C. 2010: 115 |
| Marino, P. I. & Spinelli, G. R. 2001: 14 |
| Wirth, W. W. 1952: 90 |
ginesi Ortíz, 1951: 586
| Spinelli, G. R. & Ronderos, M. M. & Diaz, F. & Marino, P. I. 2005: 149 |
| Ortiz, I. 1951: 586 |
venezuelensis Ortíz & Mirsa, 1950: 137
| Cazorla, C. G. & Marino, P. I. & Diaz, F. & Campos, R. E. 2018: 7 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Claps, G. L. & Lizarralde de Grosso, M. & Spinelli, G. R. 2015: 636 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Santana, M. & Lizarralde de Grosso, M. & Spinelli, G. R. 2012: 4 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Stein, M. & Oria, G. & Ramirez, P. & Lizarralde de Grosso, M. & Spinelli, G. R. 2012: 798 |
| Marino, P. I. & Cazorla, C. G. & Diaz, F 2011: 203 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2011: 1021 |
| Ronderos, M. M. & Marino, P. I. & Diaz, F. & Estevez, A. L. 2011: 1189 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2010: 97 |
| Diaz, F. & Ronderos, M. M. & Spinelli, G. R. 2005: 376 |
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 12 |
| Spinelli, G. R. & Ronderos, M. M. 1991: 89 |
| Spinelli, G. R. & Balseiro, E. G. 1982: 226 |
| Cavalieri, F. & Chiossone, I. F. 1966: 149 |
| Ortiz, I. & Mirsa, M. 1950: 137 |
HOFFMANIA
| Fox, I. 1948: 21 |
bahiensis
| Barbosa, F. A. S. 1947: 11 |
painteri
| Fox, I. 1946: 257 |
inamollae
| Fox, I. & Hoffman, W. A. 1944: 110 |
APELMA
| Macfie, J. W. S. 1940: 16 |
germanus
| Cavalieri, F. & Chiossone, I. F. 1966: 148 |
| Macfie, J. W. S. 1940: 27 |
limnetis
| Marino, P. I. & Spinelli, G. R. 2004: 2254 |
| Ingram, A. & Macfie, J. W. S. 1931: 169 |
| Ingram, A. & Macfie, J. W. S. 1931: 169 |
THYRIDOMYIA
| Saunders, L. G. 1925: 268 |
TRICHOHELEA
| Goetghebuer, M. 1920: 39 |
undecimpunctatus
| Kieffer, J. J. 1917: 307 |
pusillus
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Ronderos, M. M. & Spinelli, G. R. 1998: 82 |
| Lutz, A. 1913: 52 |
bambusicola
| Marino, P. I. & Cazorla, C. G. & Ronderos, M. M. 2013: 778 |
| Campos, R. E. & Spinelli, G. R. & Mogi, M. 2011: 115 |
| Ronderos, M. M. & Spinelli, G. R. 2002: 93 |
| Ronderos, M. M. & Diaz, F. 2002: 44 |
| Ronderos, M. M. & Spinelli, G. R. 2000: 134 |
| Lutz, A. 1913: 62 |
debilipalpis
| Cazorla, C. G. & Marino, P. I. & Diaz, F. & Campos, R. E. 2018: 6 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Santana, M. & Lizarralde de Grosso, M. & Spinelli, G. R. 2012: 4 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2011: 1021 |
| Ronderos, M. M. & Marino, P. I. & Diaz, F. & Estevez, A. L. 2011: 1188 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2010: 97 |
| Ronderos, M. M. & Cazorla, C. G. & Spinelli, G. R. 2010: 43 |
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Ronderos, M. M. & Spinelli, G. R. 1997: 304 |
| Cavalieri, F. & Chiossone, I. F. 1966: 148 |
| Lutz, A. 1913: 60 |
horticola
| Felippe-Bauer, M. L. & Spinelli, G. R. 2009: 28 |
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Ronderos, M. M. & Spinelli, G. R. 1998: 81 |
| Lutz, A. 1913: 61 |
insignis
| Cazorla, C. G. & Marino, P. I. & Diaz, F. & Campos, R. E. 2018: 6 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Claps, G. L. & Lizarralde de Grosso, M. & Spinelli, G. R. 2015: 634 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Stein, M. & Oria, G. & Ramirez, P. & Lizarralde de Grosso, M. & Spinelli, G. R. 2012: 798 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2011: 1021 |
| Ronderos, M. M. & Marino, P. I. & Diaz, F. & Estevez, A. L. 2011: 1188 |
| Veggiani Aybar, C. A. & Dantur, M. J. & Lizarralde de Grosso, M. & Spinelli, G. R. 2010: 97 |
| Ronderos, M. M. & Greco, N. & Spinelli, G. R. 2003: 22 |
| Spinelli, G. R. & Balseiro, E. G. 1982: 226 |
| Cavalieri, F. & Chiossone, I. F. 1966: 148 |
| Lutz, A. 1913: 51 |
COTOCRIPUS Brèthes, 1912: 451
| Brethes, J. 1912: 451 |
CULICOIDINI
| Kieffer, J. J. 1911: 1 |
| Kieffer, J. J. 1911: 319 |
HAEMATOMYIDIUM
| Goeldi, E. 1905: 137 |
PSYCHOPHAENA
| Philippi, R. A. 1865: 628 |
CERATOPOGONINAE
| Newman, E. 1834: 388 |
CULICOIDES
| Latreille, P. A. 1809: 251 |