Eupines (Byraxis) impar ( Sharp, 1874 )

Shen, Jia-Wei & Leschen, Richard A. B., 2020, Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae), Zootaxa 4777 (1), pp. 1-84 : 41-43

Eupines (Byraxis) impar ( Sharp, 1874)

Fig. 26 ; map Fig. 52

Bryaxis impar Sharp, 1874: 500 .

Bryaxis impar : reprint from Sharp 1874 by Sharp 1876: 278.

Bryaxis impar : reprinted excerpt from Sharp 1874 by Broun 1880: 130.

Eupines (Byraxis) impar : combination and subgeneric assignment by Raffray 1904: 207. Raffray 1911: 82, 1924: 157; Hudson 1923: 364, 1934: 182; Kuschel 1990: 49; Nomura and Leschen 2006: 255; Maddison 2010: 417; Shen and Leschen 2019: 267 .

Bryaxis sanguineus Broun, 1880: 132 . Bryaxis vae Schaufuss, 1888: 34 . New name for Bryaxis sanguinea Broun, 1880: 132 . Unnecessary replacement name for Bryaxis sanginea Leach, 1817 . Bryaxis fulvitarsis Broun, 1893a: 1418 . New name for Bryaxis sanguinea Broun, 1880: 132 . Unnecessary replacement name for Bryaxis sanginea Leach, 1817 . Hudson 1923: 364, 1934: 183. Eupines (Byraxis) sanguinea : combination and subgeneric assignment by Raffray 1904: 207. Raffray 1911: 82, 1924: 157; Hudson 1923: 364, 1934: 182; Nomura and Leschen 2006: 255; Maddison 2010: 417; Shen and Leschen 2019: 273. Type locality: [CL] Tairua. Type depository: NZAC LT ♂, 2 PLT. NEW SYNONYMY.

Bryaxis foveatissima Broun, 1890: 233 . Bryaxis foveatissima : reprint from Broun 1890 by Broun 1893a: 1047. Eupines (Byraxis) foveatissima : combination and subgeneric assignment by Raffray 1904: 207. Raffray 1911: 82, 1924: 157; Hudson 1923: 364, 1934: 183; Nomura and Leschen 2006: 254; Maddison 2010: 417; Shen and Leschen 2019: 265. Type locality: [AK] Clevedon. Type depository: BMNH HT ♂. NEW SYNONYMY.

Type locality: [AK] Auckland .

Type depository: BMNH LT ♂, 3 PLT .

Diagnosis: Head with vertexal foveae punctiform; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7 and A8 unmodified, A6 slightly asymmetrical, A9 largest, about 1.5 × longer than A10, with apical impression at ventral side, impression about two-fifths length of A9, inserted with a barely visible protuberance at middle, A10 conical, with small basal impression at ventral side. Abdominal ventrite 2 with pair of barely visible discal protuberances, inserted with small apical setae; ventrite 6 with pair of long median struts, about half of ventral length. Protrochanters with a medium-sized ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified.

Redescription: Body length 1.38–1.49 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes well developed and prominent, with about 35 facets; vertexal foveae punctiform; antennal tubercles moderately developed; postantennal notches weak; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7 and A8 unmodified, A6 slightly asymmetrical, A9 largest, about 1.5 × longer than A10, with apical impression at ventral side, impression about twofifths length of A9, inserted with a barely visible protuberance at middle, A10 conical, with small basal impression at ventral side. Pronotum slightly wider than long, as wide as head at eye level; lateral antebasal foveae punctiform. Elytra slightly wider than long, widest at apical two-thirds; lacking humeral calli; sutural stria barely visible. Abdomen with tergite 1 slightly more than twice length of tergite 2; ventrite 2 with pair of barely visible discal protuberances, inserted with small apical setae; ventrite 6 with pair of long median struts, about half of ventral length. Legs with protrochanters modified with a medium-sized ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.28 mm long, with slender parameres, bent near base and sinuate towards apex, curved downwards, with two pairs of apical short setae and pair of subapical long setae; median lobe bent downwards in lateral view, with two long sclerites, about as long as parameres, both sclerites curved inwards at apex in dorsal view, left one slightly shorter than right one.

Comments: The holotype of E. (B.) foveatissima and the lectotype of E. (B.) sanguinea were examined and were found having the identical modifications of A9 as the lectotype of E. (B.) impar . These names are formally synonymized here, with the fomer two names treated as junior synonyms. This species is similar to E. (B.) complector by having similar form of A9 but can be distinguished by its A9 with a barely visible protuberance ( Fig. 26B ), which is a conspicuous apical spine in E. (B.) complector ( Fig. 10B ).

Distribution: North Island. ND, AK, WO, CL, TO, BP, GB.

Additional material examined: 67 ♂♂, AK: 1, 239, Clevedon; 1, Coll. A.E. Brookes, 01/Jan/1916, Hunua Rg., Drury, Auckland, 239; 31, Lynfield, 10/May/1975, G. Kuschel, litter; 4, Bethells Matuku Res., 19/Sep–31/ Oct/1983, M.F. Tocker & B.G. Bennett; 1, AMNZ, Auckland Domain, 40 m, 12/Aug/2001, S.E. Thorpe, under rotten log; CL: 1, Little Barrier I., Pohutukawa Flat, foreshore, 03/Oct/1975, B.M. May, litter; 1, Little Bay, Kennedy Block, 22/Nov/1993, A. Larochelle, litter; WO: 1, Waharoa, 04/Jan/1992, P.A. Maddison, litter, 92/1; TO: 1, Pureora SF. Park, 610 m, 20/Oct/1991, B.M. May, litter; GB: 3, East Cape, Taikawakawa, 15/Apr/1992, G. Hall & R.C. Henderson, sifted litter, 92/58; 1, L. Waikaremoana Whareama Peninsula, Dec/2004, pit traps, Station 5; 1, Gray’s Bush, Gisborne Flats, 20/Aug/1976, A.R. Ferguson, litter; ND: 7, Kaikohe, 1289, Apr/1954, R. Hornabrook; BP: 5, Lottin Pt Rd., Waenga Bush, 16/Sep/1992, R.C. Henderson, litter; 2, Mt. Te Aroha smt., 03/Nov/1993, J.S. Dugdale, litter and liverworts; 2, Te Koau, Bush Walk, 243 m, 15/Mar–29/Apr/1993, J.S. Dugdale, Malaise trap; 1, Upper Waipiata Karakatuwhero V., 03/Feb/1993, J.S. Dugdale; 1, 243, Unknown locality: T. Broun Collection, A.E. Brookes Collection; 1, AMNZ, T. Broun Collection. Associated females (74 ♀♀): ND: 9, Kaikohe, 1289, Apr/1954, R. Hornabrook; AK: 12, Coll. A.E. Brookes, 01/Jan/1916, Hunua Rg., Drury, Auckland, 239; 50, Lynfield, 10/May/1975, G. Kuschel, litter; NN: 3, Dun Mt., 2000 ft, 6–10/20, A. Philpott.

Figures

FIGURE 10. Diagnostic characters of E. (B.) complector sp. n. A) Habitus. B) Antenna, in dorsal view. C) Same, in ventral view. D) Protrochanter. E) Ventrite 2. F) Ventrite 6. G) Aedeagus, in dorsal view. H) Same, in lateral view. I) Same, in ventral view. Scale bars: A = 1 mm, B–E = 0.2 mm, F–I = 0.1 mm.

FIGURE 26. Diagnostic characters of E. (B.) impar (Sharp, 1874). A) Habitus. B) Antenna, in lateral view. C) Same, in ventral view. D) Protrochanter. E) Ventrite 2. F) Ventrite 6. G) Aedeagus, in dorsal view. H) Same, in lateral view. I) Same, in ventral view. Scale bars: A = 1 mm, B–E = 0.2 mm, F–I = 0.1 mm.

FIGURE 52. Geographic distributions of E. (B.) impar (Sharp, 1874), E. (B.) impressifrons (Broun, 1880), E. (B.) insolita sp. n., E. (B.) lewisi Broun, 1910, E. (B.) longiceps Raffray, 1904, E. (B.) mayae sp. n., and E. (B.) micans (Sharp, 1874) in New Zealand.