Megarthrus affinis, Miller, 1853: 28

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

affinis Miller, 1853: 28 ( Megarthrus ; [preoccupied]; Type locality: Wien).

— Redtenbacher, 1857: 258 ( Megarthrus ; characters; Austria).

— Kraatz, 1857d: 1029 ( Megarthrus ; [Note: Misidentification: Bernhauer and Schubert, 1910: 35 cited sinuatocollis Kraatz as a synonym of affinis ; Kraatz did not describe the species as new, he attributed it to Lacordaire]; characters; Germany).

— Kraatz, 1857d: 1029 ( Megarthrus ; synonym of sinuatocollis ).

— Kraatz, 1868c: 350 ( Megarthrus ; synonym of sinuatocollis ).

— Fauvel, 1871a: 53 [= 1872: 27] ( Megarthrus ; characters; France; Britain; Germany; Austria; Italy; Corsica; Spain; Turkey; Algeria).

— Fauvel, 1873b: 110 [= 1873c: 3] ( Megarthrus ; Sicily).

— Redtenbacher, 1874: 283 ( Megarthrus ; characters; Austria).

— Fauvel, 1878: 86 [= 1878a: 6] ( Megarthrus ; Algeria).

— Mulsant and Rey, 1878b: 240 ( Megarthrus ; characters; France).

— Fauvel, 1886: 12 [= 1886a: 4] ( Megarthrus ; Algeria; Tunisia).

— Fowler, 1888: 430 ( Megarthrus ; characters; Britain).

— Ganglbauer, 1895: 762 ( Megarthrus ; characters; middle Europe; Mediterranean region).

— Fauvel, 1902b: 49 ( Megarthrus ; Algeria; Tunisia).

— Sainte-Claire Deville, 1906: 67 ( Megarthrus ; Corsica).

— Montandon, 1908: 77 ( Megarthrus ; Romania).

— Reitter, 1909: 198 ( Megarthrus ; characters; Germany).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; catalog).

— Portevin, 1929: 457 ( Megarthrus ; characters; France).

— Gridelli, 1930: 60 ( Megarthrus ; Libya).

— Scheerpeltz, 1931a: 188 ( Megarthrus ; characters in key).

— Scheerpeltz, 1933: 1029 ( Megarthrus ; catalog).

— Paulian, 1941: 145 ( Megarthrus ; larval characters).

— Scheerpeltz, 1947: 259 ( Megarthrus ; characters).

— Horion, 1963: 23 ( Megarthrus ; Algeria; eastern Russia; Britain; Holland; Belgium; France; Spain; Italy; Dalmatia; Greece; Crete; Turkey; Romania; Hungary; Czechoslovakia; Poland; Russia; Germamy; Austria).

— Lohse, 1964: 23 ( Megarthrus ; characters; central Europe).

— Pototskaia, 1967: 19 ( Megarthrus ; larval characters).

— Pope, 1977: 22 ( Megarthrus ; Britain).

— Hammond, 1980: 135 ( Megarthrus ; Ireland).

— Tóth, 1982: 29 ( Megarthrus ; characters; Hungary).

— Lucht, 1987: 85 ( Megarthrus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Boháč, 1993: 40 ( Megarthrus ; Czech Republic; Slovakia).

— Sparacio, 1995: 137 ( Megarthrus ; characters; notes; Sicily).

— Zanetti, 1995: 5 ( Megarthrus ; Italy).

— Krasutskii, 1997: 304 ( Megarthrus ; found on Pleurotus calyptratus ; western Siberia).

— Cuccodoro and Löbl, 1997: 1354 ( Megarthrus ; synonym of bellevoyei ).

— Assing, Frisch, Kahlen, et al., 1998: 120 ( Megarthrus ; synonym of bellevoyei ).

bimaculatus Fauvel, 1904b: 87 ( Megarthrus ; Type locality: Ceylan).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; catalog).

— Cameron, 1924b: 165 ( Megarthrus ; characters; India).

— Cameron, 1930: 128 ( Megarthrus ; characters; Ceylon; India).

— Scheerpeltz, 1933: 1029 ( Megarthrus ; catalog).

- DISTRIBUTION: India, Sri Lanka.

birmanus Fauvel, 1895: 194 ( Megarthrus ; Type locality: Bírmanie, Carin Asciuii Ghécu , 1400–1500 m).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; catalog).

— Cameron, 1924b: 163, 165 ( Megarthrus ; characters; India).

— Cameron, 1930: 128 ( Megarthrus ; characters; Burma; India).

borealis Cuccodoro and Löbl, 1996: 163 ( Megarthrus ; Type locality: USA: Alaska: Prudhoe

Bay Rd., Fish Cr., 150°50′W, 66°32′N, 250 m).

- DISTRIBUTION: USA. calcaratus Coiffait, 1977: 256 ( Megarthrus ; Type locality: Khumba Pari, Népal, 3550 m).

— Coiffait, 1982: 40 ( Megarthrus ; Nepal).

- DISTRIBUTION: Nepal. chatterjeei Cameron, 1924b: 162 , 165 ( Megarthrus ; Type locality: Chakrata District, 5000 to

6500 feet).

— Cameron, 1930: 130 ( Megarthrus ; characters; India).

— Scheerpeltz, 1933: 1029 ( Megarthrus ; catalog).

- DISTRIBUTION: India. chinese J. Li, 1993: 53 ( Megarthrus ; Type locality: Mount Laoye).

- DISTRIBUTION: China. chobauti Fauvel, 1902b: 49 , 179 ( Megarthrus ; Type locality: Yakouren, Kabilie; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]. Originally cited from Kabylie:

sommet du Djebel Bou-Berak près Dellys; forêt d’Yakouren; Bougie).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; catalog).

— Cuccodoro and Löbl, 1997: 1357 ( Megarthrus ; lectotype designation: Yakouren , Kabilie; characters; Algeria) .

- DISTRIBUTION: Algeria. clarkei Cuccodoro and Löbl, 1995: 668 ( Megarthrus ; Type locality: Ethiopia: Shewa prov.,

Addis Abeba).

— Cuccodoro, 1999a: 376 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopa. conformis Sawada, 1962: 11 , 15 ( Megarthrus ; Type locality: Japan: Honshu: Kitashirakawa,

Kyoto City).

— Cuccodoro, 1996: 480 ( Megarthrus ; characters; Japan).

- DISTRIBUTION: Japan. congoensis Cameron, 1950 g: 4 ( Megarthrus ; Type locality: Belgian Congo: volc. Visoke,

alt. 2800–3300 m).

— Cuccodoro and Löbl, 1995: 669 ( Megarthrus ; characters; Rwanda).

- DISTRIBUTION: Congo, Rwanda. conspirator Cuccodoro, 1996: 481 ( Megarthrus ; Type locality: Japan: Honshu: Gunma Pref.:

Nikko N. P., below Konsei Pass, 1500–1600 m. Also cited from Sakhalin).

— Cuccodoro and Löbl, 1997: 1360 ( Megarthrus ; characters; Japan; Far East Russia).

- DISTRIBUTION: Japan, Sakhalin, Russia. constrictus Cuccodoro, 1996: 482 ( Megarthrus ; Type locality: Japan: Honshu: Nagano Pref.:

Minami Alps, Kitazawa Toge, 2000 m).

- DISTRIBUTION: Japan. convexus Sharp, 1874: 100 ( Megarthrus ; Type locality: Simabara; [Note: See lectotype designation by Cuccodoro, 1996]).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; catalog).

— Sawada, 1962: 11 ( Megarthrus ; cited as convex; characters; Japan).

— Nakane, 1963a: 82 ( Megarthrus ; characters; habitus photograph; Japan).

— Watanabe and Shibata, 1972: 60 ( Megarthrus ; Yaku-shima Island).

— Cuccodoro, 1996: 482 ( Megarthrus ; lectotype designation; characters; Japan).

- DISTRIBUTION: Japan. corticale Lacordaire, 1835 , see: Metopsia . corticalis Sharp, 1889: 468 ( Megarthrus ; Type locality: Oyayama; [Note: See lectotype designation by Cuccodoro, 1996]).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; catalog).

— Nakane, 1963a: 82 ( Megarthrus ; characters; habitus photograph; Japan).

— Cuccodoro, 1996: 483 ( Megarthrus ; lectotype designation; characters; Japan).

- DISTRIBUTION: Japan.

danieli Cuccordoro, 1999 a: 380 ( Megarthrus ; Type locality: Ethiopia: Bale region , Bale Mountains National Park , Harenna Forest, Katcha area, 2400 m).

- DISTRIBUTION: Ethiopia.

dentatus Coiffait, 1977: 258 ( Megarthrus ; Type locality: Khumba Lager , Népal, 2200 m).

- DISTRIBUTION: Nepal.

denticollis Beck, 1817: 26 ( Omalium ; Type locality: München Umg., Engl. Garten Heu; [Note: See neotype designation by Cuccodoro and Löbl, 1997]. Originally cited from Baiern).

— Illiger, 1798: 355 ( Silpha ; variety α of hemipterus ; [Note: Cited as a synonym of denticollis by Mulsant and Rey, 1878b: 246 and Bernhauer and Schubert, 1910: 35; there is no available name]; characters).

— Erichson, 1840: 906 ( Megarthrus ; characters; Europe).

— Redtenbacher, 1849: 756 ( Megarthrus ; characters; Austria).

— Hardy, 1851: 64 ( Megarthrus ; notes; England).

— Fairmaire and Laboulbène, 1856: 655 ( Megarthrus ; characters; France).

— Redtenbacher, 1857: 257 ( Megarthrus ; characters; Austria).

— Kraatz, 1857d: 1030 ( Megarthrus ; characters; Germany).

— Peyron, 1858: 434 ( Megarthrus ; Turkey).

— Thomson, 1861: 219 ( Megarthrus ; characters; Scandinavia).

— Saulcy, 1862: 70 ( Megarthrus ; key characters).

— Fauvel, 1871a: 54 [= 1872: 28] ( Megarthrus ; characters; France; Scandinavia; Britain; Germany).

— Fauvel, 1873b: 110 [= 1873c: 3] ( Megarthrus ; Austria; Russia).

— Redtenbacher, 1874: 282 ( Megarthrus ; characters; Austria).

— Seidlitz, 1875: 232 ( Megarthrus ; characters; Baltic region).

— J. Sahlberg, 1876: 225 ( Megarthrus ; Finland).

— Mulsant and Rey, 1878b: 246 ( Megarthrus ; characters; France).

— Fowler, 1888: 429 ( Megarthrus ; characters; Britain).

— Ganglbauer, 1895: 762 ( Megarthrus ; characters; north and middle Europe; Asia Minor).

— Reitter, 1909: 198 ( Megarthrus ; characters; Germany).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; catalog).

— Johansen, 1914: 630 ( Megarthrus ; characters; Denmark).

— Portevin, 1929: 457 ( Megarthrus ; characters; France).

— Scheerpeltz, 1931a: 191 ( Megarthrus ; characters in key).

— Scheerpeltz, 1933: 1029 ( Megarthrus ; catalog).

— Kuusinen, 1933: 69 ( Megarthrus ; characters; illustrations; Finland).

— Scheerpeltz, 1947: 262 ( Megarthrus ; characters).

— Palm, 1948: 42 ( Megarthrus ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 24 ( Megarthrus ; eastern Russia; Caucasus; Ireland; Scotland; Lapland; Russia; France; Italy; Balkans; Germany; Austria).

— Lohse, 1964: 23 ( Megarthrus ; characters; central Europe).

— Raianu, 1964: 106 ( Megarthrus ; Romania).

— Kasule, 1966: 279 ( Megarthrus ; larval characters).

— Bacchus and Hammond, 1972: 62 ( Megarthrus ; donkey dung; Britain).

— Pope, 1977: 22 ( Megarthrus ; Britain).

— Hammond, 1980: 135 ( Megarthrus ; Ireland).

— Tóth, 1982: 30 ( Megarthrus ; characters; Hungary).

— Lucht, 1987: 85 ( Megarthrus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Dettner and Reissenweber, 1991 ( Megarthrus ; defensive secretions: chemistry and systematics).

— Welch, 1993: 227 ( Megarthrus ; ovariole number and ovary structure). — Zanetti, 1995: 5 ( Megarthrus ; Italy). — Muona and Viramo, 1995: 9 ( Megarthrus ; collected from fungus and dung of Ursus arctos ; Finland). — Cuccodoro, 1996: 484 ( Megarthrus ; characters; Japan). — Cuccodoro and Löbl, 1997: 1361 ( Megarthrus ; neotype designation: München Umg., Engl. Garten Heu; characters; collecting notes; Palaearctic region; Syrian record needs confirmation). — Assing, Frisch, Kahlen, et al., 1998: 120 ( Megarthrus ; note). — Kashcheev, 1999a: 57 ( Megarthrus ; Kazakhstan). - DISTRIBUTION: Europe, Russia, Turkey, Japan.

rufescens Stephens, 1834: 331 ( Megarthrus ; Type locality: Hertford; Ripley; London; Somersetshire; Salop; New Forest; Suffolk; Norfolk; Devon; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]). — Stephens, 1839: 423 ( Megarthrus ; characters; England). — Erichson, 1840: 906 ( Megarthrus ; synonym of hemipterus ). — Kraatz, 1857d: 1031 ( Megarthrus ; synonym of hemipterus ). — G. Waterhouse, 1858: 32 ( Megarthrus ; synonym of denticollis ). — Ganglbauer, 1895: 762 ( Megarthrus ; synonym of denticollis ). — Bernhauer and Schubert, 1910: 35 ( Megarthrus ; synonym of denticollis ). — Cuccodoro and Löbl, 1997: 1361, 1363 ( Megarthrus ; lectotype designation: [Note: Locality not cited]; synonym of denticollis ).

affinis Stephens, 1834: 333 ( Megarthrus ; [Note: Nomen dubium; see Cuccodoro and Löbl, 1997]; Type locality: Devonshire). — Stephens, 1839: 424 ( Megarthrus ; characters; England). — G. Waterhouse, 1858: 32 ( Megarthrus ; synonym of denticollis ). — Ganglbauer, 1895: 762 ( Megarthrus ; synonym of denticollis ). — Bernhauer and Schubert, 1910: 35 ( Megarthrus ; synonym of denticollis ). — Pope, 1977: 22 ( Megarthrus ; synonym of denticollis ). — Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; nomen dubium).

marginatus Stephens, 1834: 333 ( Megarthrus ; [Note: Nomen dubium; see Cuccodoro and Löbl, 1997]; Type locality: Essex near Southend). — Stephens, 1839: 424 ( Megarthrus ; characters; England). — Hardy, 1851: 64 ( Megarthrus ; synonym of denticollis ). — G. Waterhouse, 1858: 32 ( Megarthrus ; synonym of depressus and denticollis ). — Bernhauer and Schubert, 1910: 35 ( Megarthrus ; synonym of denticollis ). — Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; nomen dubium).

marginicollis Lacordaire, 1835: 492 ( Phloeobium ; [Note: Nomen dubium; see Cuccodoro and Löbl, 1997]; Type locality: Paris). — Erichson, 1839a: 644 ( Megarthrus ; characters; Germany). — Heer, 1839: 171 ( Megarthrus ; characters; Switzerland). — Erichson, 1840: 906 ( Megarthrus ; synonym of denticollis ). — Redtenbacher, 1849: 756 ( Megarthrus ; synonym of denticollis ). — Hardy, 1851: 64 ( Megarthrus ; synonym of denticollis ). — Kraatz, 1857d: 1030 ( Megarthrus ; synonym of denticollis ). — Mulsant and Rey, 1878b: 246 ( Megarthrus ; synonym of denticollis ). — Ganglbauer, 1895: 762 ( Megarthrus ; synonym of denticollis ). — Bernhauer and Schubert, 1910: 35 ( Megarthrus ; synonym of denticollis ). — Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; nomen dubium).

minimus Bruce, 1931: 194 ( Megarthrus ; variety of denticollis ; [Note: Nomen dubium; see Cuccodoro and Löbl, 1997]; Type locality: Tullstuguputten). — Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; nomen dubium).

dentipes Bernhauer, 1938: 17 ( Megarthrus ; Type locality: Nordwestl. China: Chinkiang; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]). — Cuccodoro and Löbl, 1997: 1364 ( Megarthrus ; lectotype designation; characters; China).

depressus Paykull, 1789: 70 ( Staphylinus ; Type locality: Suecia; [ Note : See lectotype designation by Cuccodoro and Löbl, 1997]).

— Erichson, 1840: 905 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Switzerland; France).

— Heer, 1841: 566 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Switzerland).

— Hochhuth, 1849: 206 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Imeretia; Iberia).

— Hardy, 1851: 64 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; notes; England).

— Fairmaire and Laboulbène, 1856: 655 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; France).

— Redtenbacher, 1857: 258 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Austria).

— Kraatz, 1858b: 129 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Greece).

— Thomson, 1861: 218 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Scandinavia).

— Saulcy, 1862: 70 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; key characters).

— Brisout, 1862: xlviii ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters).

— Hochhuth, 1862: 112 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Russia).

— Kraatz, 1868c: 349 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; notes).

— Fauvel, 1871a: 54 [= 1872: 28] ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; France; Scandinavia; Britain; Germany; Switzerland; Austria; Greece; Russia; Alaska [Note: According to Cuccodoro and Löbl, 1996: 151, North American records cited as sinuatocollis are misidentifications of angulicollis ]).

— Fauvel, 1873b: 110 [= 1873c: 3] ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Caucasus).

— Fauvel, 1874: 320 [= 1874b: 26] ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Siberia).

— Redtenbacher, 1874: 283 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Austria).

— Seidlitz, 1875: 231 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Baltic region).

— J. Sahlberg, 1876: 224 ( Megarthrus ; cited as sinuatocollis , spelled as sinuaticollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Finland).

— Mulsant and Rey, 1878b: 243 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; France).

— Fowler, 1888: 430 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Britain).

— Ganglbauer, 1895: 762 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Palaearctic region; North America [Note: According to Cuccodoro and Löbl, 1996: 151, North American records cited as sinuatocollis are misidentifications of angulicollis ]).

— Montandon, 1908: 77 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Romania).

— Poppius, 1909: 3 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Lena Valley, Siberia ).

— Reitter, 1909: 198 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Germany).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; catalog).

— Johansen, 1914: 629 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Denmark).

— Kemner, 1925: 71 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; larval characters).

— Portevin, 1929: 457 ( Megarthrus ; cited as sinuatocollis , spelled as sinuaticollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; France).

— Scheerpeltz, 1931a: 190 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters in key).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; catalog).

— Kuusinen, 1933: 69 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; illustrations; Finland).

— Scheerpeltz, 1947: 260 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters).

— Palm, 1948: 42 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 24 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Transbaikal; Scotland; Lapland; France; Italy; Dalmatia; Balkans; Romania; Hungary; Czechoslovakia; Germany; Austria).

— Lohse, 1964: 23 ( Megarthrus ; cited as sinuatocollis , spelled as sinuaticollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; central Europe).

— Pope, 1977: 22 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Britain).

— Hammond, 1980: 135 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Ireland).

— Tóth, 1982: 29 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; characters; Hungary).

— Lucht, 1987: 85 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Dettner and Reissenweber, 1991 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; defensive secretions: chemistry and systematics).

— Welch, 1993: 227 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; ovariole number and ovary structure).

— Boháč, 1993: 40 ( Megarthrus ; Czech Republic; Slovakia).

— Boháč, 1993: 40 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Czech Republic; Slovakia).

— Øland, 1995: 23 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; host mushroom; Norway).

— Muona and Viramo, 1995: 9 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; collected from dung of Ursus arctos ; Finland).

— Zanetti, 1995: 5 ( Megarthrus ; cited as sinuatocollis ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1369]; Italy).

— Cuccodoro and Löbl, 1997: 1364 ( Megarthrus ; lectotype designation; specimens identified as sinuatocollis Lacordaire are depressus ; characters; collecting notes; Palaearctic region, absent from southern Mediterranean region and Japan).

— Ryabukhin, 1999: 12 ( Megarthrus ; cited as sinuatocollis ; [Note: Presumably misidentified,

see Cuccodoro and Löbl, 1997: 1369]; catalog; habitat; distribution; northeastern Russia).

— Kashcheev, 1999a: 57 ( Megarthrus ; Kazakhstan).

- DISTRIBUTION: Europe, Russia.

emarginatus Stephens, 1834: 332 ( Megarthrus ; [Note: Nomen dubium; see Cuccodoro and Löbl, 1997]; Type locality: London; Suffolk; Woodland, Devon.; Raehills ).

— Stephens, 1839: 423 ( Megarthrus ; characters; England). — Ganglbauer, 1895: 762 ( Megarthrus ; synonym of depressus ).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; synonym of depressus ).

— Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; nomen dubium).

pusillus Stephens, 1834: 333 ( Megarthrus ; [Note: Nomen dubium; see Cuccodoro and Löbl, 1997]; Type locality: New Forest, Hants).

— Stephens, 1839: 424 ( Megarthrus ; characters; England).

— Hardy, 1851: 64 ( Megarthrus ; synonym of depressus ).

— G. Waterhouse, 1858: 32 ( Megarthrus ; synonym of depressus ).

— Ganglbauer, 1895: 762 ( Megarthrus ; synonym of depressus ).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; synonym of depressus ).

— Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; nomen dubium).

depressus : sensu Fauvel and others, see: prosseni Schatzmayr, 1904 .

dissymetricus Coiffait, 1977: 259 ( Megarthrus ; Type locality: Phulchoki près Katmandu).

- DISTRIBUTION: Nepal.

dominicae Cuccodoro and Löbl, 1995: 670 ( Megarthrus ; Type locality: Uganda: Toro prov. , Ruwenzori N. P., John Mate Camp, 3350 m).

- DISTRIBUTION: Uganda.

elevatus Coiffait, 1977: 254 ( Megarthrus ; Type locality: Vallée Dzunda Khola près Talpi, région de Jumla, Népal occidental, 3000–3500 m).

- DISTRIBUTION: Nepal.

emarginatus Stephens, 1834 , see: depressus Paykull, 1789 .

excisus LeConte, 1863 a: 58 ( Megarthrus ; Type locality: Lake Superior).

— Fauvel, 1878d: 193 [= 1878e: 29] ( Megarthrus ; characters; Lake Superior; New York).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Scheerpeltz, 1933: 1029 ( Megarthrus ; catalog).

— Campbell and Davies, 1991: 87 ( Megarthrus ; checklist; Canada).

— Cuccodoro and Löbl, 1996: 165 ( Megarthrus ; characters; collecting notes; Alberta; British Columbia ; Manitoba; New Brunswick; Newfoundland; Nova Scotia ; Ontario; Prince Edward Island; Quebec; Yukon Territory; Alaska; Colorado; Illinois; Massachusetts; Minnesota; New Hampshire; New Jersey; New Mexico; New York; North Carolina; Oregon; Pennsylvania; South Dakota; Vermont; West Virginia; Wisconsin).

— Downie and Arnett, 1996: 429 ( Megarthrus ; characters; USA).

- DISTRIBUTION: Canada, USA.

falasha Cuccodoro and Löbl, 1995: 671 ( Megarthrus ; Type locality: Ethiopia: Gojam prov. , 8 km W Falega, Birham, 10 ° 46 ̍ N, 38 ° 03 ̍ E, 2820 m).

— Cuccordoro, 1999a: 374, 376, 380 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

fennicus Lahtinen, 1938: 254 ( Megarthrus ; Type locality: Alajärvi; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]. Also cited from Turku, Ruissalo).

— Palm, 1948: 43 ( Megarthrus ; characters; Sweden; Norway; Finland).

— Cuccodoro and Löbl, 1997: 1369 ( Megarthrus ; lectotype designation: Alajärvi; characters; collecting notes; North Europe; Siberia).

- DISTRIBUTION: Europe, Russia.

fernandezi Jarrige, 1952 , see: Metopsia .

fijianus Cuccodoro, 1998: 107 ( Megarthrus ; Type locality: Fiji: Viti Levu, Navai , base of Mt. Tamanivi, 1000 m).

- DISTRIBUTION: Fiji.

flavolimbatus Cameron, 1924b: 164 , 165 ( Megarthrus ; Type locality: Simla Hills: Matiana, Narkanda, Kotgarh, 7000 to 9200 feet).

— Cameron, 1930: 131 ( Megarthrus ; characters; India).

— Scheerpeltz, 1933: 1029 ( Megarthrus ; catalog).

flavosignatus Bierig, 1940: 378 ( Megarthrus ; Type locality: Costa Rica: Carpintera ).

- DISTRIBUTION: Costa Rica.

flavus Stephens, 1834 , see: Metopsia .

franzi Scheerpeltz, 1947 , see: stercorarius Mulsant and Rey, 1878 .

gallicus C. Koch, 1938 , see: Metopsia .

gigas Fagel, 1957 a: 29 ( Megarthrus ; Type locality: Urundi: territoire de Ngozi, Samutuku, 2600 m. Also cited from Belgian Congo).

— Cuccodoro and Löbl, 1995: 672 ( Megarthrus ; characters; Burundi, Zaire).

- DISTRIBUTION: Burundi, Congo.

gomerense H. Franz, 1986 , see: Metopsia .

gressitti Cuccodoro, 1998: 107 ( Megarthrus ; Type locality: New Caledonia: Col des Roussettes , 450–550 m).

- DISTRIBUTION: New Caledonia.

harennaensis Cuccordoro, 1999 a: 380 ( Megarthrus ; Type locality: Ethiopia: Bale region ,

Bale Mountains National Park, Harenna Forest, 2 km below Rira, 2850 m).

- DISTRIBUTION: Ethiopia.

hemipterus Illiger, 1794: 597 ( Silpha ; Type locality: Amt Campen, einige Stunden von Braunschweig).

— Olivier, 1795:(42): 38 ( Staphylinus ; [Note: Misidentified as melanocephalus Fabricius but cited by Bernhauer and Schubert, 1910: 36 as synonym of hemipterus ; Olivier attributed melanocephalus to Paykull who attributed it to Fabricius. Cuccodoro and Löbl, 1997: 1372 cited melanocephalus Olivier as a synonym of hemipterus ]; characters).

— Illiger, 1798: 355 ( Silpha ; also variety β; characters).

— Gyllenhal, 1813: 699 ( Omalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 36 cited depressus variety c Gyllenhal as a synonym of hemipterus Illiger ; Gyllenhal’s variety is not an available name]; characters; Sweden).

— Gyllenhal, 1813: 699 ( Omalium ; synonym of depressus ).

— Erichson, 1839a: 645 ( Megarthrus ; characters; Germany).

— Heer, 1839: 172 ( Megarthrus ; characters; Switzerland).

— Erichson, 1840: 906 ( Megarthrus ; characters; Europe).

— Redtenbacher, 1849: 757 ( Megarthrus ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 655 ( Megarthrus ; characters; France).

— Redtenbacher, 1857: 258 ( Megarthrus ; characters; Austria).

— Kraatz, 1857d: 1031 ( Megarthrus ; characters; Germany).

— Thomson, 1861: 219 ( Megarthrus ; characters; Scandinavia).

— Saulcy, 1862: 70 ( Megarthrus ; key characters).

— Fauvel, 1871a: 54 [= 1872: 28] ( Megarthrus ; characters; France; Europe).

— Redtenbacher, 1874: 283 ( Megarthrus ; characters; Austria).

— Seidlitz, 1875: 232 ( Megarthrus ; characters; Baltic region).

— Mulsant and Rey, 1878b: 249 ( Megarthrus ; characters; France).

— Sharp, 1889: 468 ( Megarthrus ; Japan).

— Fowler, 1888: 430 ( Megarthrus ; characters; Britain).

— Ganglbauer, 1895: 763 ( Megarthrus ; characters; Europe).

— Reitter, 1909: 198 ( Megarthrus ; characters; Germany).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Johansen, 1914: 631 ( Megarthrus ; characters; Denmark).

— Portevin, 1929: 457 ( Megarthrus ; characters; France).

— Scheerpeltz, 1931a: 191 ( Megarthrus ; characters in key).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog).

— Scheerpeltz, 1947: 262 ( Megarthrus ; characters).

— Sawada, 1962: 14 ( Megarthrus ; characters; Japan).

— Raianu, 1963: 85 ( Megarthrus ; Romania).

— Lohse, 1964: 23 ( Megarthrus ; characters; central Europe).

— Szujecki, 1969a: 247 ( Megarthrus ; notes; Poland).

— Pope, 1977: 22 ( Megarthrus ; Britain).

— Hammond, 1980: 135 ( Megarthrus ; may not be in Ireland).

— Tóth, 1982: 30 ( Megarthrus ; characters; Hungary).

— Lucht, 1987: 85 ( Megarthrus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Welch, 1993: 227 ( Megarthrus ; ovariole number and ovary structure).

— Boháč, 1993: 40 ( Megarthrus ; Czech Republic; Slovakia).

— Zanetti, 1995: 5 ( Megarthrus ; Italy).

— Cuccodoro, 1996: 485 ( Megarthrus ; characters; Japan).

— Cuccodoro and Löbl, 1997: 1372 ( Megarthrus ; characters; collecting notes; Palaearctic region including Far East Russia and Japan).

— Fowles, Alexander, and Key, 1999: 129 ( Megarthrus ; saproxylic species; status as endangered or threatened species; Britain).

— Kashcheev, 1999a: 57 ( Megarthrus ; Kazakhstan).

- DISTRIBUTION: Europe, Russia, China, Japan.

nitiduloides Lacordaire, 1835: 493 ( Phloeobium ; Type locality: Paris; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]).

— Erichson, 1839a: 645 ( Megarthrus ; synonym of hemipterus ).

— Erichson, 1840: 906 ( Megarthrus ; synonym of denticollis ).

— Redtenbacher, 1849: 757 ( Megarthrus ; synonym of hemipterus ).

— Kraatz, 1857d: 1031 ( Megarthrus ; synonym of hemipterus ).

— Mulsant and Rey, 1878b: 250 ( Megarthrus ; synonym of hemipterus ).

— Ganglbauer, 1895: 763 ( Megarthrus ; synonym of hemipterus ).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; synonym of hemipterus ).

— Cuccodoro and Löbl, 1997: 1372 ( Megarthrus ; lectotype designation; synonym of hemipterus ).

heteropus Sawada, 1962: 14 , 15 ( Megarthrus ; Type locality: Japan: Honshu: Aomori Pref.: Tsuta ).

— J. Li, 1993: 18 ( Megarthrus ; cited as heterops; China).

— Cuccodoro, 1996: 495 ( Megarthrus ; synonym of hemipterus ).

— Cuccodoro and Löbl, 1997: 1372 ( Megarthrus ; synonym of hemipterus ).

heteropus Sawada, 1962 , see: hemipterus Illiger, 1794 .

hispanicum Scheerpeltz, 1960 , see: Metopsia .

horticola Cuccodoro and Löbl, 1995: 673 ( Megarthrus ; Type locality: Uganda: Bugisu prov. , Mt. Elgon , Sipi, 1750 m. Also cited from Kenya and Zaire).

- DISTRIBUTION: Congo, Kenya, Uganda.

hutu Cuccodoro and Löbl, 1995: 673 ( Megarthrus ; Type locality: Rwanda: Albert N. P., V. Visoke, Bishoke, 2800–3300 m. Also cited from Zaire).

- DISTRIBUTION: Congo, Rwanda.

inaequalis Bierig, 1940: 379 ( Megarthrus ; Type locality: Costa Rica: Vara Blanca , 1800 m, entre volcanes Barba y Poás).

- DISTRIBUTION: Costa Rica.

incubifer Cuccodoro, 1996: 486 ( Megarthrus ; Type locality: Japan).

- DISTRIBUTION: Japan.

integricollis Coiffait, 1977: 253 ( Megarthrus ; Type locality: Phulchoki près Katmandu).

- DISTRIBUTION: Nepal.

impressicollis Eppelsheim, 1893: 67 ( Megarthrus ; Type locality: Ost-Sibirien, in der Nähe von Irkutsk, von Tunka bis Munku-Sardik im Sajan’schen Gebirge ).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Cuccodoro, 1996: 485 ( Megarthrus ; characters; Japan).

— Cuccodoro and Löbl, 1997: 1375 ( Megarthrus ; characters; Russia; Japan).

japonicus Sharp, 1874: 99 ( Megarthrus ; Type locality: Japan; [ Note : See lectotype designation by Cuccodoro, 1996]).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Cameron, 1949b: 460 ( Megarthrus ; Japan).

— Sawada, 1962: 12 ( Megarthrus ; characters; Japan).

— Nakane, 1963a: 82 ( Megarthrus ; characters; habitus photograph; Japan).

— Watanabe and Shibata, 1972: 60 ( Megarthrus ; Yaku-shima Island).

— Cuccodoro, 1996: 488 ( Megarthrus ; lectotype designation; characters; Japan; China; Russia). — Cuccodoro and Löbl, 1997: 1375 ( Megarthrus ; characters; China; Asian Russia).

— Watanabe, 1998: 313 ( Megarthrus ; Japan).

- DISTRIBUTION: Japan, China, Russia.

kamerunensis Bernhauer, 1942c: 348 ( Megarthrus ; Type locality: Mont Cameroun: 1800–2000 m, versant Sud-Est).

— Cuccodoro and Löbl, 1995: 674 ( Megarthrus ; notes; Cameroon).

- DISTRIBUTION: Cameroon.

kuscheli Cuccodoro, 1998: 108 ( Megarthrus ; Type locality: New Caledonia: Table Unio , near Col d’Amieu, 900 m).

- DISTRIBUTION: New Caledonia.

longicornis Wollaston, 1854: 615 ( Megarthrus ; Type locality: Madeira: Funchal ).

— Wollaston, 1857: 206 ( Megarthrus ; habitat; Madeira Islands).

— Wollaston, 1865: 525 ( Megarthrus ; notes; Madeira Islands; [Note: Also listed from the Canary Islands but these specimens were misidentified according to Cuccodoro and Löbl, 1997: 1376 and are now identified as wollastoni ]).

— Wollaston, 1871: 313 ( Megarthrus ; notes).

— Fauvel, 1902b: 49 ( Megarthrus ; Madeira).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Smetana, 1963b: 31 ( Megarthrus ; Madeira).

— Cuccodoro and Löbl, 1997: 1376 ( Megarthrus ; characters; collecting notes; Madeira).

- DISTRIBUTION: Madeira.

magnicaudatus Cuccodoro and Löbl, 1995: 674 ( Megarthrus ; Type locality: Ethiopia: Balê prov. , Adelay Forest, 3100 m).

— Cuccodoro, 1999a: 376 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

mahnerti Cuccodoro and Löbl, 1995: 675 ( Megarthrus ; Type locality: Kenya: Central prov. , Mt. Aberdares, near N. P. entrance, 2300 m. Also cited from Tanzania).

- DISTRIBUTION: Kenya, Tanzania.

major Cuccodoro and Löbl, 1995: 675 ( Megarthrus ; Type locality: Zaire: Kivu, Albert N. P., Tshiaberimu sect., Kirungu, 2720 m).

- DISTRIBUTION: Congo.

malaisei Scheerpeltz, 1965: 144 ( Megarthrus ; Type locality: N.E. Burma, Kambaiti, 7000 ft.). - DISTRIBUTION: Myanmar.

mammiger Bierig, 1940: 375 ( Megarthrus ; Type locality: Costa Rica: Vara Blanca , 1800 m, entre volcanes Barba y Poás).

- DISTRIBUTION: Costa Rica.

maniwaata Cuccodoro and Löbl, 1995: 676 ( Megarthrus ; Type locality: Zaire: Ruwenzori Range, Albert N. P., Kalonge , river Nyamwamba, tributary of Butahu, 2010 m.

Also cited from Rwanda).

- DISTRIBUTION: Congo, Rwanda.

marginatus Stephens, 1834 , see: denticollis Beck, 1817 .

marginicollis Lacordaire, 1835 , see: denticollis Beck, 1817 .

maronitus Fagel, 1968 a: 190 ( Megarthrus ; Type locality: Liban: Nabeh Safa, 1000 m). martensi Coiffait, 1982: 131 ( Megarthrus ; Type locality: Nepal: nördl. Dhaulagiri, Gompa/Tarakot, 3300–3400 m).

- DISTRIBUTION: Nepal.

mastiger Bierig, 1940: 376 ( Megarthrus ; Type locality: Costa Rica: Vara Blanca , 1800 m, entre volcanes Barba y Poás).

- DISTRIBUTION: Costa Rica.

merabet Cuccodoro and Löbl, 1995: 676 ( Megarthrus ; Type locality: Rwanda: Cyangugu Pref., around Nyakabuye. Also cited from Zaire).

- DISTRIBUTION: Congo, Rwanda.

minimus Bruce, 1932 , see: denticollis Beck, 1817 .

minor Coiffait, 1977: 251 ( Megarthrus ; Type locality: Région de Taksang près Tukche, Takala, Népal central).

- DISTRIBUTION: Nepal.

montanus Sawada, 1962: 14 , 15 ( Megarthrus ; Type locality: Japan: Honshu: Nagano Pref.: Inago-yu , 1500–2000 m).

— Cuccodoro, 1996: 489 ( Megarthrus ; characters; China; Japan; Russia).

— Cuccodoro and Löbl, 1997: 1381 ( Megarthrus ; characters; Far East Russia; China; Japan).

- DISTRIBUTION: Russia, China, Japan.

subangulatus Sawada, 1962: 14 , 15 ( Megarthrus ; subspecies of montanus ; Type locality: Japan: Honshu: Nara Pref.: Mt. Oodaigahara).

— Cuccodoro, 1996: 489 ( Megarthrus ; synonym of montanus ).

— Cuccodoro and Löbl, 1997: 1381 ( Megarthrus ; synonym of montanus ).

monticola Cameron, 1942b: 322 ( Megarthrus ; Type locality: Chyulu Hills, altitude 5600 feet). — Cuccodoro and Löbl, 1995: 677 ( Megarthrus ; characters; Kenya).

- DISTRIBUTION: Kenya.

mukankundiyeorum Cuccodoro and Löbl, 1995: 678 ( Megarthrus ; Type locality: Rwanda: 25 km, N. Kibuye, Kayove, 2100 m).

- DISTRIBUTION: Rwanda.

mwami Cuccodoro and Löbl, 1995: 679 ( Megarthrus ; Type locality: Burundi, Bururi terr., Nyamurenbe, 900 m. Also cited from Rwanda).

- DISTRIBUTION: Burundi, Rwanda.

nanus Cuccodoro and Löbl, 1995: 680 ( Megarthrus ; Type locality: Africa or., Katona

[= Tanzania, Moshi prov.], Mt. Kilimanjaro).

- DISTRIBUTION: Tanzania.

negus Cuccodoro and Löbl, 1995: 680 ( Megarthrus ; Type locality: Ethiopia: Balê prov. , Dinshu, 3200 m).

— Cuccodoro, 1999a: 376, 382 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

newtoni Cuccodoro and Löbl, 1996: 168 ( Megarthrus ; Type locality: USA: Arizona: Apache Co., Big L. , 22 mi SW Eagar, 2700 m. Also cited from New Mexico).

- DISTRIBUTION: USA.

nigerrimus Cameron, 1941: 56 ( Megarthrus ; Type locality: Kashmir: Gulmarg).

- DISTRIBUTION: India.

nigrinus J. Sahlberg, 1876: 225 ( Megarthrus ; variety of sinuatocollis ; Type locality: Ryska Lappmarken, Porjeguba, Finland; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]. Also cited from Pekkala i Ruovesi; Gorki).

— Ganglbauer, 1895: 762 ( Megarthrus ; variety of sinuatocollis ).

— Münster, 1911: 277 ( Megarthrus ; synonym of sahlbergi ).

— Scheerpeltz, 1931a: 190 ( Megarthrus ; characters in key).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog).

— Palm, 1948: 42 ( Megarthrus ; characters; Sweden; Norway; Finland).

— Muona and Viramo, 1995: 9 ( Megarthrus ; collected from fungus; Finland).

— Cuccodoro and Löbl, 1997: 1381 ( Megarthrus ; lectotype designation: Ryska Lappmarken , Porjeguba , Finland; characters; collecting notes; North Europe; Asian Russia) .

- DISTRIBUTION: Europe, Russia.

sahlbergi Münster, 1911: 277 ( Megarthrus ; Type locality: Lakselv in Porsanger ; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]. Also cited from Norwegen: Südvaranger; Karasjok; Målselvtal; Kongsberg und Kristiana).

— Scheerpeltz, 1931a: 190 ( Megarthrus ; synonym of nigrinus ).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; synonym of nigrinus ).

— Scheerpeltz, 1947: 261 ( Megarthrus ; synonym of nigrinus ).

— Cuccodoro and Löbl, 1997: 1381 ( Megarthrus ; lectotype designation: Lakselv, Pors.; synonym of nigrinus ).

niloticus Cuccodoro and Löbl, 1995: 681 ( Megarthrus ; Type locality: Uganda: Bugisu prov. , Mt. Elgon , Sipi, 1750 m. Also cited from Rwanda and Zaire).

- DISTRIBUTION: Congo, Rwanda, Uganda.

nitiduloides Lacordaire, 1835 see: hemipterus Illiger, 1794 .

nitidulus Kraatz, 1857d: 1028 ( Megarthrus ; Type locality: Mollenfalde; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]. Originally cited from Grafschaft Glatz; Cassel).

— Redtenbacher, 1857: 995 ( Megarthrus ; characters; Austria).

— Saulcy, 1862: 70 ( Megarthrus ; key characters).

— Fauvel, 1871a: 53 [= 1872: 27] ( Megarthrus ; characters; France; Poland; Germany).

— Fauvel, 1873b: 109 [= 1873c: 2] ( Megarthrus ; Siberia).

— Redtenbacher, 1874: 283 ( Megarthrus ; characters).

— Mulsant and Rey, 1878b: 248 ( Megarthrus ; characters; Switzerland; Germany).

— Ganglbauer, 1895: 763 ( Megarthrus ; characters; Germany; Austria; Poland).

— Reitter, 1909: 198 ( Megarthrus ; characters; Germany).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Portevin, 1929: 457 ( Megarthrus ; characters; France).

— Scheerpeltz, 1931a: 191 ( Megarthrus ; characters in key).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog).

— Kuusinen, 1933: 70 ( Megarthrus ; characters; illustrations; Finland).

— Scheerpeltz, 1947: 262 ( Megarthrus ; characters).

— Horion, 1963: 25 ( Megarthrus ; Transbaikal; Britain; Norway; Sweden; Finland; Russia; Poland; Czech Republic; Slovakia; Romania; Balkans; Switzerland; Italy; France; Holland; Germany; Austria).

— Lohse, 1964: 23 ( Megarthrus ; characters; central Europe).

— Smetana, 1964d: 51 ( Megarthrus ; Czechoslovakia).

— Smetana, 1968: 227 ( Megarthrus ; Mongolia).

— Smetana 1975c: 154 ( Megarthrus ; Mongolia).

— Tóth, 1982: 30 ( Megarthrus ; characters).

— Lucht, 1987: 85 ( Megarthrus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Sweden).

— Boháč, 1993: 40 ( Megarthrus ; Czech Republic; Slovakia).

— Muona and Viramo, 1995: 9 ( Megarthrus ; collected from dung of Ursus arctos ; Finland).

— Zanetti, 1995: 5 ( Megarthrus ; Italy).

— Cuccodoro and Löbl, 1997: 1384 ( Megarthrus ; lectotype designation: Mollenfalde ; characters; Europe, Far East Russia; Mongolia) .

- DISTRIBUTION: Europe, Russia, Mongolia.

notabilis Cameron, 1941: 56 ( Megarthrus ; Type locality: Darjeeling: Ghum district).

- DISTRIBUTION: India.

occidentalis Cuccodoro and Löbl, 1996: 171 ( Megarthrus ; Type locality: Canada: British ogloblini Bruch, 1940: 111 ( Megarthrus ; Type locality: Argentina: Misiones, Loreto). - DISTRIBUTION: Argentina.

oromo Cuccordoro, 1999 a: 382 ( Megarthrus ; Type locality: Ethiopia: Bale region , Bale Mountains National Park, around Finchaya Habera, 3400 m). - DISTRIBUTION: Ethiopia.

ovalis Cameron, 1950 g: 5 ( Megarthrus ; Type locality: Belgian Congo: volc. Visoke, alt. 2800–3300 m). — Cuccodoro and Löbl, 1995: 681 ( Megarthrus ; characters; Rwanda; Zaire). — Assing and Wunderle, 2001: 36 ( Proteinus ; Cyprus). - DISTRIBUTION: Congo, Rwanda.

palmense H. Franz, 1986 , see: Metopsia .

panga Cuccodoro and Löbl, 1995: 682 ( Megarthrus ; Type locality: Zaire: Kivu prov., Albert N. P., V. Mikeno , near Rweru, 2400 m). - DISTRIBUTION: Congo.

parallelus Sharp, 1874: 99 ( Megarthrus ; Type locality: Nagasaki; [Note: See lectotype designation by Cuccodoro, 1996]). — Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog). — Sawada, 1962: 12 ( Megarthrus ; characters; Japan). — Cuccodoro, 1996: 491 ( Megarthrus ; lectotype designation; characters; Japan). - DISTRIBUTION: Japan.

subparallelus Sawada, 1962: 12 , 15 ( Megarthrus ; Type locality: Japan: Honshu: Osaka Pref.: Katsuoji, Minoo). — Watanabe and Shibata, 1972: 60 ( Megarthrus ; Yaku-shima Island). — Cuccodoro, 1996: 491 ( Megarthrus ; synonym of parallelus ).

pecki Cuccodoro and Löbl, 1996: 173 ( Megarthrus ; Type locality: Canada: British Columbia : Metlakatla. Also cited from Alberta, Ontario, Illinois, Iowa, Michigan, Minnesota, New Hampshire, New York, North Carolina, Pennsylvania, Tennessee, Washington, and West Virginia). - DISTRIBUTION: Canada, USA.

peckorum Cuccodoro, 1998: 109 ( Megarthrus ; Type locality: New Caledonia: Col d’Amieu , north of La Foa, 500 m). - DISTRIBUTION: New Caledonia.

pictus Motschulsky, 1845 a: 39 ( Megarthrus ; Type locality: Sitka;; [Note: See lectotype designation by Cuccodoro and Löbl, 1996]. Originally cited from Nord de la Californie). — Mäklin, 1852: 324 ( Megarthrus ; characters; notes; Sitka). — Fauvel, 1878d: 193 [= 1878e: 29] ( Megarthrus ; characters; Sitka Island). — Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog). — Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog). — Hatch, 1957: 108 ( Megarthrus ; characters; British Columbia ; Washington; Oregon). — Campbell and Davies, 1991: 87 ( Megarthrus ; checklist; USA; Canada). — Cuccodoro and Löbl, 1996: 177 ( Megarthrus ; lectotype designation: Sitka; characters; British Columbia ; California; Oregon; Washington). - DISTRIBUTION: Canada, USA.

primus Cuccodoro, 1995: 259 ( Megarthrus ; Type locality: Papua New Guinea, Morobe district, Biaru Road, Mt. Kolorong, 7 ° 28 ̍ 40 ̎ S, 146 ° 49 ̍ 20 ̎ E, 2200 m). — Cuccodoro, 1998: 112 ( Megarthrus ; characters; Papua New Guinea; Irian Jaya). - DISTRIBUTION: New Guinea.

prosseni Schatzmayr, 1904: 212 ( Megarthrus ; Type locality: Teufelsgraben, Villach, Dobratschgebiet, Carinthia, Austria; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]). — Olivier, 1795:(42): 36 ( Staphylinus ; cited as depressus ; [Note: Misidentified according to

Cuccodoro and Löbl, 1997: 1387]; characters). — Paykull, 1800: 412 ( Staphylinus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Sweden).

— Gyllenhal, 1810: 210 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters).

— Gyllenhal, 1813: 699 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters for two varieties; Sweden).

— Gyllenhal, 1827: 464 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]).

— C. Sahlberg, 1827: 280 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Finland).

— Curtis, 1829: 28 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; catalog; Britain).

— Curtis, 1829: 28 ( Megarthrus ; cited as macropterus Gravenhorst ; catalog; Britain).

— Mannerheim, 1830: 53 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Finland; Sweden).

— Mannerheim, 1831: 467 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Finland; Sweden).

— Stephens, 1834: 332 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; England).

— Stephens, 1834: 332 ( Megarthrus ; cited as macropterus Gravenhorst ; characters; England).

— Lacordaire, 1835: 494 ( Phloeobium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; France).

— Zetterstedt, 1838: 50 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; habitat; Lapland; Sweden).

— Westwood, 1838a: 18 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; type species).

— Erichson, 1839a: 644 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Germany).

— Heer, 1839: 171 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Switzerland).

— Stephens, 1839: 423 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; England).

— Erichson, 1839a: 644 ( Megarthrus ; cited as macropterus Gravenhorst ; synonym of depressus ).

— Stephens, 1839: 423 ( Megarthrus ; cited as macropterus Gravenhorst ; characters; England).

— Erichson, 1840: 905 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Europe).

— Erichson, 1840: 905 ( Megarthrus ; cited as macropterus Gravenhorst ; synonym of depressus ).

— Laporte, 1840: 191 ( Omalium ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; England; France; Sweden).

— Redtenbacher, 1849: 756 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Austria).

— Hardy, 1851: 64 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; notes; England).

— Hardy, 1851: 64 ( Megarthrus ; cited as macropterus Gravenhorst ; synonym of depressus ).

— Fairmaire and Laboulbène, 1856: 654 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; France).

— Fairmaire and Laboulbène, 1856: 654 ( Megarthrus ; cited as macropterus Gravenhorst ; characters; France).

— Redtenbacher, 1857: 258 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Austria).

— Kraatz, 1857d: 1027 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Germany).

— Thomson, 1859: 52 (type species of Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]).

— Thomson, 1861: 218 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Scandinavia).

— Saulcy, 1862: 70 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro

— Hochhuth, 1862: 112 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Russia).

— Fauvel, 1871a: 52 [= 1872: 26] ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; France; Europe).

— Redtenbacher, 1874: 283 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Austria).

— Seidlitz, 1875: 231 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Baltic region).

— J. Sahlberg, 1876: 224 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Finland).

— Mulsant and Rey, 1878b: 235 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; France).

— Mulsant and Rey, 1878b: 236 ( Megarthrus ; cited as macropterus Gravenhorst ; synonym of depressus ).

— Fowler, 1888: 430 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Britain).

— Ganglbauer, 1895: 762 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Europe).

— Ganglbauer, 1895: 762 ( Megarthrus ; cited as macropterus Gravenhorst ; synonym of depressus ).

— Poppius, 1905: 12 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Orkney Islands).

— Poppius, 1909: 3 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Lena Valley, Siberia ).

— Reitter, 1909: 198 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Germany).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Bernhauer and Schubert, 1910: 35 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; catalog).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; cited as macropterus Gravenhorst ; synonym of depressus ).

— Johansen, 1914: 629 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Denmark).

— Kemner, 1925: 64 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; larval characters).

— Portevin, 1929: 456 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; France).

— Scheerpeltz, 1931a: 189 ( Megarthrus ; characters in key).

— Scheerpeltz, 1931a: 188 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters in key).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog).

— Scheerpeltz, 1933: 1029 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; catalog).

— Kuusinen, 1933: 66 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; illustrations; Finland).

— Scheerpeltz, 1947: 258 ( Megarthrus ; characters).

— Scheerpeltz, 1947: 257 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters).

— Palm, 1948: 41 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 22 ( Megarthrus ; Alps).

— Horion, 1963: 21 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Germany; Austria).

— Lohse, 1964: 23 ( Megarthrus ; characters; central Europe).

— Lohse, 1964: 23 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro

— Smetana, 1964d: 51 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Czechoslovakia).

— Smetana, 1968: 226 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Mongolia).

— Szujecki, 1968a: 710 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Poland).

— Bacchus and Hammond, 1972: 62 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; donkey and elephant dung; Britain).

— Smetana 1975c: 154 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Mongolia).

— Pope, 1977: 22 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Britain).

— Hammond, 1980: 135 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; Ireland).

— Tóth, 1982: 29 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; Hungary).

— Lucht, 1987: 85 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Welch, 1993: 227 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; ovariole number and ovary structure).

— Zanetti, 1995: 5 ( Megarthrus ; “ depressus Auct. nec Paykull, 1789” cited as synonym; Italy).

— Cuccodoro, 1995: 262 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; water loading behavior).

— Muona and Viramo, 1995: 9 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; collected from dung of Ursus arctos ; Finland).

— Sparacio, 1995: 137 ( Megarthrus ; cited as depressus ; [Note: Misidentified according to Cuccodoro and Löbl, 1997: 1387]; characters; notes; Sicily).

— Cuccodoro and Löbl, 1997: 1387 ( Megarthrus ; lectotype designation: Teufelsgraben , Villach , Dobratschgebiet , Carinthia, Austria; characters; collecting notes; western Palaearctic region, evidently absent from Mediterranean region , records from Siberia and Far Eastern Russia need verification) .

— Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; cited as macropterus Gravenhorst ; nomen dubium).

— Ryabukhin, 1999: 13 ( Megarthrus ; cited as depressus ; [Note: Based on the subsequent literature cited by Ryabukhin, I assume this reference is based on misidentifications; see Cuccodoro and Löbl, 1997: 1387]; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Europe, Russia.

pusillus Stephens, 1834 , see: depressus Paykull, 1789 .

ras Cuccodoro and Löbl, 1995: 683 ( Megarthrus ; Type locality: Ethiopia: Gonder [= Simên] prov., Arghine, 3500 m or higher).

— Cuccodoro, 1999a: 374 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

retusus Stephens, 1834 , see: Metopsia .

riedeli Cuccodoro, 1998: 113 ( Megarthrus ; Type locality: Indonesia: Irian Jaya, Jayawijaya, Diuremna , 1900–2100 m).

- DISTRIBUTION: New Guinea.

rougemonti Cuccodoro and Löbl, 1995: 683 ( Megarthrus ; Type locality: Ethiopia: Balê prov. , Dinshu, 3200 m).

— Cuccodoro, 1999a: 376, 386 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

rufomarginatus Cameron, 1914: 525 ( Megarthrus ; Type locality: S. India: Nilgiri Hills ).

— Cameron, 1924b: 166 ( Megarthrus ; characters; India).

— Cameron, 1930: 129 ( Megarthrus ; characters; India; Sumatra).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog).

— Cameron, 1934: 17 ( Megarthrus ; India).

- DISTRIBUTION: India, Indonesia.

rufescens Stephens, 1834 , see: denticollis Beck, 1817 .

sahlbergi Munster, 1911 , see: nigrinus J. Sahlberg, 1876 .

sawadai Cuccodoro, 1996: 492 ( Megarthrus ; Type locality: Japan: Honshu: Gunma Pref.: below Usui Pass , 750 m).

- DISTRIBUTION: Japan.

scotti Cuccodoro and Löbl, 1995: 684 ( Megarthrus ; Type locality: Ethiopia: Gamo Gofa prov. , Gughé highlands, Dita, c. 3000 m).

— Cuccodoro, 1999a: 376 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

scriptus Sharp, 1889: 469 ( Megarthrus ; Type locality: Chuzenji , Gunma Pref., Honshu , Japan; [Note: See lectotype designation by Cuccodoro, 1996]. Also cited from Yuyama).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Sawada, 1962: 10 ( Megarthrus ; characters; Japan).

— Nakane, 1963a: 82 ( Megarthrus ; characters; habitus photograph; Japan).

— Cuccodoro, 1996: 492 ( Megarthrus ; lectotype designation: Chuzenji , Gunma Pref., Honshu, Japan; characters; Japan) .

- DISTRIBUTION: Japan.

sculpticollis Coiffait, 1982: 131 ( Megarthrus ; Type locality: Nepal: environs de Lughla, Khumbu).

- DISTRIBUTION: Nepal.

selenitus Cuccodoro and Löbl, 1995: 684 ( Megarthrus ; Type locality: Uganda: Ruwenzori Range, Toro prov., above Kilembe, 2000 m).

- DISTRIBUTION: Uganda.

septempunctatus Champion, 1925: 102 ( Megarthrus ; Type locality: W. Almora in Kumaon).

— Cameron, 1930: 132 ( Megarthrus ; characters; India).

— Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog).

- DISTRIBUTION: India.

serrula Wollaston, 1865: 525 , 76 appendix ( Megarthrus ; Type locality: Gomera: above Hermigua; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]).

— Fauvel, 1902b: 50 ( Megarthrus ; Canary Islands).

— Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog).

— Cuccodoro and Löbl, 1997: 1390 ( Megarthrus ; lectotype designation; characters; collecting notes; Canary Islands).

— Hernández, Outerelo, and Gamarra, 1994: 192 ( Megarthrus ; Canary Islands).

— Assing, 1999a: 345 ( Megarthrus ; Canary Islands).

- DISTRIBUTION: Canary Islands.

sexpunctatus Cameron, 1941: 57 ( Megarthrus ; Type locality: Darjeeling: Ghum district).

- DISTRIBUTION: India.

shibatai Sawada, 1962: 13 , 15 ( Megarthrus ; Type locality: Japan: Honshu: Nagano Pref.: Inago-yu , 1500–2000 m).

— Cuccodoro, 1996: 494 ( Megarthrus ; characters; Japan).

- DISTRIBUTION: Japan.

simienensis Fagel, 1957 a: 30 ( Megarthrus ; Type locality: Ethiopia: Simien, E. of Mindigabsa , over 10,000 ft.).

— Cuccodoro and Löbl, 1995: 685 ( Megarthrus ; characters; Ethiopia).

— Cuccodoro, 1999a: 376 ( Megarthrus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

sinuatocollis Lacordaire, 1835: 493 ( Phloeobium ; Type locality: Paris; [Note: Nomen dubium; see Cuccodoro and Löbl, 1997]). — Cuccodoro and Löbl, 1997: 1351 ( Megarthrus ; nomen dubium). - DISTRIBUTION: France.

smetanai Cuccodoro and Löbl, 1996: 181 ( Megarthrus ; Type locality: Canada: British Columbia : 4 mi W Midway. Also cited from Northwest Territories, Quebec, and Alaska). - DISTRIBUTION: Canada, USA.

solitarius Sharp, 1887: 743 ( Megarthrus ; Type locality: Guatemala, Purula). — Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog). - DISTRIBUTION: Costa Rica, Guatemala.

adelphus Bierig, 1940: 377 ( Megarthrus ; variety of solitarius ; Type locality: Costa Rica: Carpintera, 1800 m).

spathuliformis Assing and Wunderle, 1999: 438 ( Megarthrus ; Type locality: E. Islas Canarias, 11, La Palma, SW Laguna de Barlovento, Mte. Hueco, 1000 m). — Assing, 2000b: 115 ( Megarthrus ; collecting notes; Canary Islands). - DISTRIBUTION: Canary Islands.

spinosus Cuccodoro and Löbl, 1995: 686 ( Megarthrus ; Type locality: Uganda: Bugisu prov. , Mt. Elgon, Sipi, 1750 m). - DISTRIBUTION: Uganda.

stercorarius Mulsant and Rey, 1878b: 238 ( Megarthrus ; Type locality: Hautes-Pyrénées; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]). — Bernhauer and Schubert, 1910: 36 ( Megarthrus ; catalog). — Scheerpeltz, 1931a: 188 ( Megarthrus ; characters in key). — Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog). — Scheerpeltz, 1947: 257 ( Megarthrus ; characters). — Zanetti, 1995: 5 ( Megarthrus ; Italy). — Cuccodoro and Löbl, 1997: 1392 ( Megarthrus ; lectotype designation; characters; collecting notes; Pyrenees; Alps; Carpathians; Caucasus). — Assing, Frisch, Kahlen, et al., 1998: 120 ( Megarthrus ; note). - DISTRIBUTION: Pyrenees, Alps, Carpathians, Caucasus.

franzi Scheerpeltz, 1947: 252 ( Megarthrus ; Type locality: Österreich: Kärnten, Hohe Tauern, Sonnblick-Gruppe, Fleissgasthof; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]). — Horion, 1963: 22 ( Megarthrus ; Austria; Switzerland). — Lohse, 1964: 23 ( Megarthrus ; characters; central Europe). — Lucht, 1987: 85 ( Megarthrus ; checklist; Germany; Austria; Switzerland). — Zanetti, 1995: 5 ( Megarthrus ; synonym of stercorarius ). — Cuccodoro and Löbl, 1997: 1392 ( Megarthrus ; lectotype designation: Fleisssgasthof, Sonnblick-Gruppe, Hohe Tauern, Kärnten, Austria; synonym of stercorarius ). — Assing, Frisch, Kahlen, et al., 1998: 120 ( Megarthrus ; synonym of stercorarius ).

strandi Scheerpeltz, 1931 a: 188, 189 ( Megarthrus ; Type locality: Asker am Westufer des Oslo-Fjords, zwischen Drammen und Oslo; [Note: See lectotype designation by Cuccodoro and Löbl, 1997]). — Scheerpeltz, 1933: 1030 ( Megarthrus ; catalog). — Kuusinen, 1933: 66 ( Megarthrus ; characters; illustrations; Finland). — Scheerpeltz, 1947: 260 ( Megarthrus ; characters). — Palm, 1948: 41 ( Megarthrus ; characters; Sweden; Norway; Finland). — Cuccodoro and Löbl, 1997: 1395 ( Megarthrus ; lectotype designation: W Oslo Fjord, Asker, Vestfold, Norway; characters; collecting notes; North Europe). - DISTRIBUTION: Finland, Sweden, Norway.

stylifer Cuccodoro and Löbl, 1995: 686 ( Megarthrus ; Type locality: Zaire: Ruwenzori range, Albert N. P. , Kyandolire, river Mulaku, tributary of Kakalari, 1750 m).